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<document id="9DE80A2536C6610A5B450B3770261CA9" ID-DOI="10.5281/zenodo.197753" ID-GBIF-Dataset="779898d0-f0ad-4f5e-8e0d-adddba683374" ID-ISSN="1175-5326" ID-Zenodo-Dep="197753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460277580866" checkinUser="plazi" docAuthor="Zhang, Dong &amp; Qiao, Gexia" docDate="2010" docId="03F187F809543A6EFF09DE1BFB582382" docLanguage="en" docName="zt02608p024.pdf" docOrigin="Zootaxa 2608" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mollitrichosiphum (Metatrichosiphon) nigrum Zhang &amp; Qiao, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFC8FF80095F3A60FF9ED959FFAD2210" masterDocTitle="Mollitrichosiphum Suenaga from China (Hemiptera: Aphididae), with the description of one new species" masterLastPageNumber="24" masterPageNumber="1" pageNumber="12" updateTime="1698248838330" updateUser="plazi">
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<mods:title id="5D075854A9A8085E6A2963D2A8576E87">Mollitrichosiphum Suenaga from China (Hemiptera: Aphididae), with the description of one new species</mods:title>
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<mods:namePart id="50BEB50D689B3712742FB5B7DC96A464">Zhang, Dong</mods:namePart>
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<mods:namePart id="F7FD012E394276F2CE02E5E93B6B33E3">Qiao, Gexia</mods:namePart>
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<mods:date id="BABFE8F4C9C1BEB391CB411C760AC619">2010</mods:date>
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<treatment id="03F187F809543A6EFF09DE1BFB582382" ID-DOI="http://doi.org/10.5281/zenodo.5614791" ID-GBIF-Taxon="119396294" ID-Zenodo-Dep="5614791" LSID="urn:lsid:plazi:treatment:03F187F809543A6EFF09DE1BFB582382" httpUri="http://treatment.plazi.org/id/03F187F809543A6EFF09DE1BFB582382" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C342656509543A6BFF09DE1BFEC2256F" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BE736EE09543A6BFF09DE1BFC9D254C" blockId="11.[151,816,1858,1919]" box="[151,816,1858,1884]" pageId="11" pageNumber="12">
<heading id="D0AF818209543A6BFF09DE1BFC9D254C" bold="true" box="[151,816,1858,1884]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B92CEAFC09543A6BFF09DE1BFC9D254C" bold="true" box="[151,816,1858,1884]" pageId="11" pageNumber="12">
<taxonomicName id="4C584D6D09543A6BFF09DE1BFD66254C" box="[151,715,1858,1884]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov." subGenus="Metatrichosiphon">
<emphasis id="B92CEAFC09543A6BFF09DE1BFED4254C" bold="true" box="[151,377,1858,1884]" italics="true" pageId="11" pageNumber="12">Mollitrichosiphum</emphasis>
(
<emphasis id="B92CEAFC09543A6BFE14DE1BFDCD254C" bold="true" box="[394,608,1858,1884]" italics="true" pageId="11" pageNumber="12">Metatrichosiphon</emphasis>
)
<emphasis id="B92CEAFC09543A6BFDECDE1BFD66254C" bold="true" box="[626,715,1858,1884]" italics="true" pageId="11" pageNumber="12">nigrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F578709543A6BFD4DDE1BFC9D254C" box="[723,816,1858,1884]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE736EE09543A6BFF09DE3CFEC2256F" blockId="11.[151,816,1858,1919]" box="[151,367,1893,1919]" pageId="11" pageNumber="12">
(
<figureCitation id="13632A6B09543A6BFF01DE3CFEBC256F" box="[159,273,1893,1919]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="11" pageNumber="12">Figures 6</figureCitation>
,
<figureCitation id="13632A6B09543A6BFE83DE3CFE96256F" box="[285,315,1893,1919]" captionStart="FIGURE 13" captionStartId="21.[151,255,1363,1387]" captionTargetBox="[151,1436,190,1358]" captionTargetId="figure@21.[151,1436,190,1364]" captionTargetPageId="21" captionText="FIGURE 13. Dorsal view of body, a. Mollitrichosiphum (Metatrichosiphon) yamabiwae Suenaga (oviparous female), b. Mollitrichosiphum (Metatrichosiphon) yamabiwae Suenaga (alate viviparous female), c. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov. (apterous viviparous female), d. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov. (alate viviparous female)." httpUri="https://zenodo.org/record/197766/files/figure.png" pageId="11" pageNumber="12">13</figureCitation>
c, d)
</paragraph>
</subSubSection>
<subSubSection id="C342656509543A6BFF09DEF6FBF425FF" pageId="11" pageNumber="12" type="description">
<paragraph id="8BE736EE09543A6BFF09DEF6FBF425FF" blockId="11.[151,1436,1967,2031]" pageId="11" pageNumber="12">
<emphasis id="B92CEAFC09543A6BFF09DEF6FE9725D9" bold="true" box="[151,314,1967,1993]" pageId="11" pageNumber="12">Description.</emphasis>
Apterous viviparous female: Body black with thin white pruinosity in life. For general measurements see
<tableCitation id="C6DA035509543A6BFEECDE8CFE6725FF" box="[370,458,2005,2031]" captionStart="TABLE 1" captionStartId="2.[151,241,562,586]" captionTargetBox="[151,1436,688,1905]" captionTargetPageId="2" captionText="TABLE 1. Biometric data (mean, range and Standard Division) of Mollitrichosiphum (Metatrichosiphon) nigrum sp." httpUri="http://table.plazi.org/id/DF276666095D3A62FF09DB6BFA31205A" pageId="11" pageNumber="12" tableUuid="DF276666095D3A62FF09DB6BFA31205A">Table 1</tableCitation>
. Mounted specimens: body oblong oval (
<figureCitation id="13632A6B09543A6BFC31DE8CFBAF25FF" box="[943,1026,2005,2031]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="11" pageNumber="12">Figs. 6</figureCitation>
o, 13c).
</paragraph>
</subSubSection>
<subSubSection id="C342656509533A6DFF5BD9CEFA9125A9" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BE736EE09533A6CFF5BD9CEFB3F2037" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
<emphasis id="B92CEAFC09533A6CFF5BD9CEFE6422A1" bold="true" box="[197,457,151,177]" pageId="12" pageNumber="13">Mounted specimens:</emphasis>
Head: Dorsum of head, antennal segments III, and ultimate rostral segment dark brown, antennal segments IIIVI brownish. Head fused with prothorax. Antennal segments IIIVI with transverse imbrications, head with ventral spinulose imbrications on the lateral sides. Median portion of front flat, antennal tubercles slightly developed (
<figureCitation id="13632A6B09533A6CFD3AD851FD432332" box="[676,750,264,290]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
a). Dorsal setae thick and long, mostly acuminate, rarely furcate; ventral setae fine, long and pointed. Head with two pairs of frontal setae, three pairs of dorsal setae between antennae, and three or four pairs of dorsal setae between eyes. Eyes multi-faceted, with ocular tubercles. Antennae six-segmented (
<figureCitation id="13632A6B09533A6CFDD7D821FD302382" box="[585,669,376,402]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Figs. 6</figureCitation>
b, 6c), proportion of segments IVI: 20, 14, 100, 31, 38, 31+45, respectively. Antennal setae pointed at apex, setae directed outwards distinctly longer than setae pointing inwards, segments IVI each with 79, 57, 2735, 5 or 6, 4 or 5, 3+4 setae, respectively; apex of processus terminalis with three short setae. Rostrum reaching abdominal segment I; ultimate rostral segment long wedge-shaped (
<figureCitation id="13632A6B09533A6CFED0DB54FE382037" box="[334,405,525,551]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
d), with 3 pairs of primary setae and 6-7 pairs of secondary setae.
</paragraph>
<paragraph id="8BE736EE09533A6CFF5BDB6BFEDB2117" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
Thorax: dorsum of thorax brown. Venter with spinulose imbrications. Dorsal setae of thorax thick and long, pointed or furcate. Pronotum with 2224 setae; mesonotum with 2426 setae; metanotum with 2830 setae. Mesosternal furca blackish brown, sclerotized distinctly, with a short stem (
<figureCitation id="13632A6B09533A6CFBCDDB24FB362087" box="[1107,1179,637,663]" captionStart="FIGURE 4" captionStartId="10.[151,255,1305,1329]" captionTargetBox="[277,1301,218,1253]" captionTargetId="figure@10.[264,1323,197,1284]" captionTargetPageId="10" captionText="FIGURE 4. Mollitrichosiphum (Metatrichosiphon) nandii Busu, apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal hair on abdominal tergite VIII; h. siphunculus; i. cauda; j. anal plate; k. genital plate. Alate viviparous female: l. antennal segment III. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197757/files/figure.png" pageId="12" pageNumber="13">Fig. 4</figureCitation>
), length of single arm
<quantity id="4CA09B0B09533A6CFF09DBFBFEEB20AC" box="[151,326,674,700]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="12" pageNumber="13" unit="mm" value="0.13" valueMax="0.15" valueMin="0.11">0.110.15 mm</quantity>
, 2.693.68 basal diameter of antennal segment III. Femora and tibiae brownish, tarsi brown.
<collectingCountry id="F34F767E09533A6CFF09DB91FF7E20F2" box="[151,211,712,738]" name="India" pageId="12" pageNumber="13">Hind</collectingCountry>
tibiae with 5363 transverse ridges in basal 3/4, tarsi with weakly transverse imbrications. First tarsal chaetotaxy: 7, 7, 7.
</paragraph>
<paragraph id="8BE736EE09533A6CFF5BDA4BFBFA266C" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
Abdomen: brown, except siphunculi dark brown. Venter of abdominal segments IVI with coarsely spinules. Dorsal setae of abdomen thick and long, pointed or furcate, ventral setae fine and pointed, dorsal setae 1.893.19 times as long as ventral setae. Abdominal tergites IVI with 1824, 3238, 4044, 4448, 30 34, 1618 setae, respectively; tergite VII with two pairs of spinal setae; tergite VIII with one pair of spinal setae. Spiracles on prothorax and abdominal segments IVII round, and open. Siphunculi extremely long, slightly swollen in median (
<figureCitation id="13632A6B09533A6CFE43DA94FD8E21F7" box="[477,547,973,999]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
h), flange distinct, densely spinulose imbrications in apical, other parts with spinules; with 110130 long setae. Cauda and anal plate spinulose. Cauda semi-round (
<figureCitation id="13632A6B09533A6CFB26DAABFAAF261C" box="[1208,1282,1010,1036]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">
Fig.
<date id="FFE6102E09533A6CFB6FDAABFAAF261C" box="[1265,1282,1010,1036]" pageId="12" pageNumber="13">6</date>
</figureCitation>
i), with 810 thick and long setae. Anal plate transversely elliptical, with 1622 thick and long setae and a transverse band of cell-like markings in apical (
<figureCitation id="13632A6B09533A6CFD9ADD64FDE72647" box="[516,586,1085,1111]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
j). Genital plate transverse oval (
<figureCitation id="13632A6B09533A6CFC5BDD64FBA72647" box="[965,1034,1085,1111]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
k), with 1418 setae and spinulose imbrications. Gonapophyses three, each with two or three short and pointed setae.
</paragraph>
<paragraph id="8BE736EE09533A6CFF5BDDDEFF5926D7" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
<emphasis id="B92CEAFC09533A6CFF5BDDDEFDAF26B1" bold="true" box="[197,514,1159,1185]" pageId="12" pageNumber="13">Alate viviparous female:</emphasis>
Body
<quantity id="4CA09B0B09533A6CFDCBDDD1FCA826B2" box="[597,773,1160,1186]" metricMagnitude="-3" metricUnit="m" metricValue="2.195" metricValueMax="2.68" metricValueMin="1.71" pageId="12" pageNumber="13" unit="mm" value="2.195" valueMax="2.68" valueMin="1.71">1.712.68 mm</quantity>
long,
<quantity id="4CA09B0B09533A6CFCCADDD1FBA826B2" box="[852,1029,1160,1186]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.6" metricValueMin="7.4" pageId="12" pageNumber="13" unit="mm" value="0.9" valueMax="1.06" valueMin="0.74">0.741.06 mm</quantity>
wide. General measurements see
<tableCitation id="C6DA035509533A6CFF09DDF4FF5C26D7" box="[151,241,1197,1223]" captionStart="TABLE 2" captionStartId="5.[151,241,460,484]" captionTargetBox="[151,1436,544,1836]" captionTargetPageId="5" captionText="TABLE 2. Biometric data (mean, range and Standard Division) of Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov. (Alate viviparous females and first instar nymph in mm)." httpUri="http://table.plazi.org/id/DF276666095A3A65FF09D895FC802014" pageId="12" pageNumber="13" tableUuid="DF276666095A3A65FF09D895FC802014">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE736EE09533A6CFF58DD8BFCA827F8" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
<emphasis id="B92CEAFC09533A6CFF58DD8BFE6726FC" bold="true" box="[198,458,1234,1260]" pageId="12" pageNumber="13">Mounted specimens:</emphasis>
Body oblong oval (
<figureCitation id="13632A6B09533A6CFD24DD8BFCA126FC" box="[698,780,1234,1260]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Figs. 6</figureCitation>
p, 13d). Head: Dorsum of head, antennal segments I to III and rostral segment V dark brown, antennal segments IV to VI and rostral segment IV brownish. Head separated from prothorax. Antennal segments IV to VI with transverse imbrications. Head with 1517 dorsal setae. Antennae six-segmented. Length in proportion of antennal segments IVI: 16: 12: 100: 31: 39: 31+48, respectively. Antennal setae stout and long, pointed at apex, segments IVI each with 57, 4 or 5, 2125, 79, 5, 4+4 setae, respectively; apex of processus terminalis with four short setae. Antennal segment III with 1416 round secondary rhinaria; primary rhinaria round and weakly ciliated. Rostrum reaching hind coxa, with 3 pairs of primary setae and 7 pairs of secondary setae.
</paragraph>
<paragraph id="8BE736EE09533A6CFF5BDCABFBF4248C" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
Thorax: Dorsum of thorax dark brown. Dorsal setae of thorax long and pointed. Pronotum with 2024 dorsal setae. Femora, basal and apical tibiae, and tarsi dark brown, median part of tibiae brownish. Femora with sparse spinules in distal part on inner surface, tarsi with weakly transverse imbrications.
<collectingCountry id="F34F767E09533A6CFB77DF63FA892444" box="[1257,1316,1594,1620]" name="India" pageId="12" pageNumber="13">Hind</collectingCountry>
tibia with 4953 short transverse ridges in basal 3/5. Veins of wings and pterostigma dark brown. In fore wings media twice branched; pterostigma long wedge shaped; hind wings with 2 oblique veins.
</paragraph>
<paragraph id="8BE736EE09533A6CFF5BDFFFFB2C25F0" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="13">
Abdomen: From the median part of tergites III to VI fused each other, forming a large sclerotized brown patches; marginal part brownish; siphunculi, cauda, anal plate and genital plate dark brown. Venter of abdominal segments IIIV with coarsely spinules. Abdominal tergite I with 10 setae, tergite VII with 4 setae, tergite VIII with 2 setae. Spiracles on prothorax and abdominal segments IVII oval and open, spiracular sclerite oval and brown. Siphunculi extremely long (
<figureCitation id="13632A6B09533A6CFC95DE6FFCFE2540" box="[779,851,1846,1872]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
p), flange distinct, with spinulose imbrications in basal and distal part, other parts with imbrications; 21.2323.16 times as long as its distal width; with 189 210 long and short setae. Cauda sub-triangle with spinules, and 810 thick and long setae. Anal plate transversely elliptical with spinules, with 1517 thick and long setae. Genital plate transverse oval, with spinulose imbrications, with 1822 setae. Others same as apterous viviparous female.
</paragraph>
<paragraph id="8BE736EE09523A6DFF5BD9CEFD5B22A2" blockId="13.[151,1437,151,738]" box="[197,758,151,178]" pageId="13" pageNumber="14">
<emphasis id="B92CEAFC09523A6DFF5BD9CEFE1022A1" bold="true" box="[197,445,151,177]" pageId="13" pageNumber="14">First-instar nymph:</emphasis>
Body long oval (
<figureCitation id="13632A6B09523A6DFD17D9C1FD6222A2" box="[649,719,152,178]" captionStart="FIGURE 6" captionStartId="13.[151,255,1463,1487]" captionTargetBox="[284,1314,795,1435]" captionTargetId="figure@13.[264,1323,791,1441]" captionTargetPageId="13" captionText="FIGURE 6. Mollitrichosiphum (Metatrichosiphon) nigrum sp. nov., apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="13" pageNumber="14">
Fig.
<quantity id="4CA09B0B09523A6DFD5ED9C1FD6222A2" box="[704,719,152,178]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="m" value="6.0">6</quantity>
</figureCitation>
m).
</paragraph>
<paragraph id="8BE736EE09523A6DFF5BD9E4FBB620F2" blockId="13.[151,1437,151,738]" pageId="13" pageNumber="14">
<emphasis id="B92CEAFC09523A6DFF5BD9E4FE7822C7" bold="true" box="[197,469,189,215]" pageId="13" pageNumber="14">Mounted specimens:</emphasis>
Head fused with prothorax. Median front slightly convex, antennal tubercles indistinct. Eyes with eight facets and ocular tubercles. Antennae four-segmented, about 0.50 times as long as body length, segments III and IV with transverse imbrications; length in proportion of segments IIV: 37: 28: 100: 52+69, respectively; processus terminalis about 1.33 times as long as base of segment IV; segments IIV each with 0 or 1, 2, 1 or 2, 2+0 stout and long setae, respectively; apex of processus terminalis with two or three setae; length of setae on segment III about 2.33 times as long as widest diameter of the segment. Rostrum reaching abdominal segment VI; ultimate rostral segment slender and long wedge-shaped, 5.76 basal width, 2.14 second hind tarsal segment. Dorsal setae thick and long, pointed. Head with one pair of frontal setae, two pairs of spinal dorsal setae, and two pairs of lateral setae. Pronotum with one pair of spinal setae and one pair of long and stout lateral setae; mesonotum and metanotum each with one pair of spinal setae, and two pairs of pleural setae. Abdominal tergites IV each with one pair of spinal, one pair of pleural, and one pair of marginal setae; tergite VI with one pair of pleural setae; tergite VII with one pair of spinal setae and one pair of marginal tubercles, each with one stout hair at apex; tergite VIII with one pair of spinal tubercles, each with one stout hair at apex. First tarsal chaetotaxy: 2, 2, 2. Siphunculi bare, cone-shaped, about 0.14 times as long as body. Cauda and anal plate each with 4 thick and long setae.
</paragraph>
<caption id="DF27666609523A6DFF09DCEEFE67247E" httpUri="https://zenodo.org/record/197759/files/figure.png" pageId="13" pageNumber="14" targetBox="[284,1314,795,1435]" targetPageId="13">
<paragraph id="8BE736EE09523A6DFF09DCEEFE67247E" blockId="13.[151,1437,1463,1646]" pageId="13" pageNumber="14">
<emphasis id="B92CEAFC09523A6DFF09DCEEFEB727DF" bold="true" box="[151,282,1463,1487]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
<taxonomicName id="4C584D6D09523A6DFEBEDCEEFCBD27DE" box="[288,784,1463,1486]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nigrum" status="sp. nov." subGenus="Metatrichosiphon">
<emphasis id="B92CEAFC09523A6DFEBEDCEEFE4B27DE" box="[288,486,1463,1486]" italics="true" pageId="13" pageNumber="14">Mollitrichosiphum</emphasis>
(
<emphasis id="B92CEAFC09523A6DFE68DCEEFD1E27DE" box="[502,691,1463,1486]" italics="true" pageId="13" pageNumber="14">Metatrichosiphon</emphasis>
)
<emphasis id="B92CEAFC09523A6DFD5DDCEEFCBD27DE" box="[707,784,1463,1486]" italics="true" pageId="13" pageNumber="14">nigrum</emphasis>
</taxonomicName>
<emphasis id="B92CEAFC09523A6DFC89DCEEFCDF27DF" bold="true" box="[791,882,1463,1487]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A21F578709523A6DFC89DCEEFCC327DF" box="[791,878,1463,1487]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
apterous viviparous female: a. dorsal view of head; b. antennal segment III; c. antennal segment VI; d. ultimate rostral segment; e. mesosternal furca; f. hind tibia, tarsi and claws; g. dorsal setae on abdominal tergite I; h. siphunculus; i. cauda; j. anal plate; k. genital plate; o. dorsal view of body. Alate viviparous female: l. antennal segment III; p. dorsal view of body. First instar nymph: m. dorsal view of body. n. groups of the speceis colonized on undersides of leaves of host plant from Guangxi, China photoed by J. F. Wang. Scale bars = 0.10mm.
</paragraph>
</caption>
<paragraph id="8BE736EE09523A6DFF5BDFC0FD3924C9" blockId="13.[151,1436,1689,2015]" pageId="13" pageNumber="14">
<emphasis id="B92CEAFC09523A6DFF5BDFC0FE1424A3" bold="true" box="[197,441,1689,1715]" pageId="13" pageNumber="14">Material examined.</emphasis>
<typeStatus id="54E3884C09523A6DFE5EDFC3FD9D24A4" box="[448,560,1690,1716]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
, apterous viviparous female,
<emphasis id="B92CEAFC09523A6DFC16DFC0FBE324A3" bold="true" box="[904,1102,1689,1715]" pageId="13" pageNumber="14">
<collectingCountry id="F34F767E09523A6DFC16DFC0FC7B24A3" box="[904,982,1689,1715]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
, Guangxi
</emphasis>
: Longsheng,
<date id="FFE6102E09523A6DFB71DFC3FAC824A4" box="[1263,1381,1690,1716]" pageId="13" pageNumber="14" value="2006-05-21">21.v.2006</date>
, No. 18913, on
<taxonomicName id="4C584D6D09523A6DFE8DDF99FD3D24C9" authority="J.F Wang" authorityName="J.F Wang" box="[275,656,1727,1753]" class="Magnoliopsida" family="Sabiaceae" genus="Meliosma" kingdom="Plantae" order="Sabiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="B92CEAFC09523A6DFE8DDF99FDAE24C9" box="[275,515,1728,1753]" italics="true" pageId="13" pageNumber="14">Meliosma cuneifolia</emphasis>
(J.F Wang)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE736EE09523A6DFF5BDFBCFA9125A9" blockId="13.[151,1436,1689,2015]" pageId="13" pageNumber="14">
<typeStatus id="54E3884C09523A6DFF5BDFBCFE9624EF" box="[197,315,1765,1791]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
: 3 apterous viviparous females and 2 alate viviparous females, data same as
<typeStatus id="54E3884C09523A6DFB27DFBCFA8E24EF" box="[1209,1315,1765,1791]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
; Guangxi: Xingan: Yongancun,
<date id="FFE6102E09523A6DFE3DDE53FDB32534" box="[419,542,1802,1828]" pageId="13" pageNumber="14" value="2006-07-02">2.vii.2006</date>
, No. 19258, 5 apterous viviparous females and 1 alate viviparous female on
<taxonomicName id="4C584D6D09523A6DFF09DE69FD862559" authority="J.H. Huang" authorityName="J.H. Huang" box="[151,555,1839,1865]" class="Magnoliopsida" family="Simaroubaceae" genus="Ailanthus" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="altissima">
<emphasis id="B92CEAFC09523A6DFF09DE69FE2D2559" box="[151,384,1840,1865]" italics="true" pageId="13" pageNumber="14">Ailanthus altissima</emphasis>
(J.H. Huang)
</taxonomicName>
;
<emphasis id="B92CEAFC09523A6DFDA9DE76FD462559" bold="true" box="[567,747,1839,1865]" pageId="13" pageNumber="14">
<collectingCountry id="F34F767E09523A6DFDA9DE76FD2A2559" box="[567,647,1839,1865]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
, Fujian
</emphasis>
: Mt. Wuyi, alt.
<quantity id="4CA09B0B09523A6DFC31DE76FC5A2559" box="[943,1015,1839,1865]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="13" pageNumber="14" unit="m" value="820.0">820m</quantity>
,
<date id="FFE6102E09523A6DFB9ADE76FB2C2559" box="[1028,1153,1839,1865]" pageId="13" pageNumber="14" value="2003-07-06">6.vii.2003</date>
, No. 14417, 2 apterous viviparous females, 1 alate viviparous female and 2 apterous viviparous nymphs on
<taxonomicName id="4C584D6D09523A6DFBEEDE0CFF5F2584" authority="X.L. Huang" authorityName="X.L. Huang" class="Magnoliopsida" family="Elaeagnaceae" genus="Elaeagnus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="pungens">
<emphasis id="B92CEAFC09523A6DFBEEDE0CFAF8257E" box="[1136,1365,1877,1902]" italics="true" pageId="13" pageNumber="14">Elaeagnus pungens</emphasis>
(X.L. Huang)
</taxonomicName>
; Jianyang,
<date id="FFE6102E09523A6DFEEBDE23FE422584" box="[373,495,1914,1940]" pageId="13" pageNumber="14" value="1973-07-02">2.vii.1973</date>
, No. 5265, 2 apterous viviparous females, 4 alate viviparous females, 1 apterous viviparous nymph and 4 alate viviparous nymphs on
<taxonomicName id="4C584D6D09523A6DFC9ADEF9FA9525A9" authority="C.L. Fang &amp; B.L. Zhang" authorityName="C.L. Fang &amp; B.L. Zhang" box="[772,1336,1951,1977]" class="Magnoliopsida" family="Sabiaceae" genus="Meliosma" kingdom="Plantae" order="Sabiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="myriantha">
<emphasis id="B92CEAFC09523A6DFC9ADEF9FC5525A9" box="[772,1016,1952,1977]" italics="true" pageId="13" pageNumber="14">Meliosma myriantha</emphasis>
(C.L. Fang &amp; B.L. Zhang)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C342656509523A6EFF5BDE9DFDC222C7" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BE736EE09523A6DFF5BDE9DFD3925CF" blockId="13.[151,1436,1689,2015]" box="[197,660,1988,2015]" pageId="13" pageNumber="14">
<emphasis id="B92CEAFC09523A6DFF5BDE9DFEC825CE" bold="true" box="[197,357,1988,2014]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingCountry id="F34F767E09523A6DFEF2DE9CFE1F25CF" box="[364,434,1989,2015]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
(Fujian, Guangxi).
</paragraph>
<paragraph id="8BE736EE09513A6EFF5BD9CEFDC222C7" blockId="14.[151,1436,151,402]" pageId="14" pageNumber="15">
<emphasis id="B92CEAFC09513A6EFF5BD9CEFEF722A1" bold="true" box="[197,346,151,177]" pageId="14" pageNumber="15">Host plants.</emphasis>
<taxonomicName id="4C584D6D09513A6EFEFFD9C1FCBF22A2" box="[353,786,152,178]" class="Magnoliopsida" family="Simaroubaceae" genus="Ailanthus" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="altissima">
<emphasis id="B92CEAFC09513A6EFEFFD9C1FDEF22A1" box="[353,578,152,177]" italics="true" pageId="14" pageNumber="15">Ailanthus altissima</emphasis>
(Simaroubaceae)
</taxonomicName>
,
<taxonomicName id="4C584D6D09513A6EFC82D9C1FBA122A1" box="[796,1036,152,177]" class="Magnoliopsida" family="Sabiaceae" genus="Meliosma" kingdom="Plantae" order="Sabiales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="B92CEAFC09513A6EFC82D9C1FBA122A1" box="[796,1036,152,177]" italics="true" pageId="14" pageNumber="15">Meliosma cuneifolia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C584D6D09513A6EFB84D9C1FA3122A2" box="[1050,1436,152,178]" class="Magnoliopsida" family="Sabiaceae" genus="Meliosma" kingdom="Plantae" order="Sabiales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="myriantha">
<emphasis id="B92CEAFC09513A6EFB84D9C1FAA122A1" box="[1050,1292,152,177]" italics="true" pageId="14" pageNumber="15">Meliosma myriantha</emphasis>
(Sabiaceae)
</taxonomicName>
and
<taxonomicName id="4C584D6D09513A6EFF56D9E4FDC622C7" box="[200,619,189,215]" class="Magnoliopsida" family="Elaeagnaceae" genus="Elaeagnus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pungens">
<emphasis id="B92CEAFC09513A6EFF56D9E4FE0022C6" box="[200,429,189,214]" italics="true" pageId="14" pageNumber="15">Elaeagnus pungens</emphasis>
(Elaeagnaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C342656509513A6EFF5BD9BBFB582382" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BE736EE09513A6EFF5BD9BBFB582382" blockId="14.[151,1436,151,402]" pageId="14" pageNumber="15">
<emphasis id="B92CEAFC09513A6EFF5BD9BBFEFF22EC" bold="true" box="[197,338,226,252]" pageId="14" pageNumber="15">Etymology.</emphasis>
The new species is named for its body color in life. “
<taxonomicName id="4C584D6D09513A6EFC5BD9BBFBB722EB" authority="Latin" authorityName="Latin" box="[965,1050,226,251]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B92CEAFC09513A6EFC5BD9BBFBB722EB" box="[965,1050,226,251]" italics="true" pageId="14" pageNumber="15">nigrum</emphasis>
</taxonomicName>
” (Latin) means black, dark.
<emphasis id="B92CEAFC09513A6EFF5BD85EFEF92331" bold="true" box="[197,340,263,289]" pageId="14" pageNumber="15">Comments.</emphasis>
This new species can be easily separated from other congeneric species by black body with slightly wax powders in life. The new one is similar with
<taxonomicName id="4C584D6D09513A6EFCD4D874FBD32356" box="[842,1150,301,326]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rhusae">
<emphasis id="B92CEAFC09513A6EFCD4D874FBD32356" box="[842,1150,301,326]" italics="true" pageId="14" pageNumber="15">Mollitrichosiphum rhusae</emphasis>
</taxonomicName>
, but differed from it in: more transverse ridges on hind tibia, with
<quantity id="4CA09B0B09513A6EFD16D80BFD5D237C" box="[648,752,338,364]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" metricValueMax="1.6002" metricValueMin="1.3462" pageId="14" pageNumber="15" unit="in" value="58.0" valueMax="63.0" valueMin="53.0">5363 in</quantity>
apterae,
<quantity id="4CA09B0B09513A6EFCC5D80BFC6E237C" box="[859,963,338,364]" metricMagnitude="0" metricUnit="m" metricValue="1.2953999999999999" metricValueMax="1.3462" metricValueMin="1.2446" pageId="14" pageNumber="15" unit="in" value="51.0" valueMax="53.0" valueMin="49.0">4953 in</quantity>
alatae (
<taxonomicName id="4C584D6D09513A6EFBBED80BFBC3237B" box="[1056,1134,338,363]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rhusae">
<emphasis id="B92CEAFC09513A6EFBBED80BFBC3237B" box="[1056,1134,338,363]" italics="true" pageId="14" pageNumber="15">rhusae</emphasis>
</taxonomicName>
:
<quantity id="4CA09B0B09513A6EFB1ED80BFB45237C" box="[1152,1256,338,364]" metricMagnitude="-1" metricUnit="m" metricValue="9.271" metricValueMax="11.684" metricValueMin="6.858" pageId="14" pageNumber="15" unit="in" value="36.5" valueMax="46.0" valueMin="27.0">2746 in</quantity>
apterae,
<quantity id="4CA09B0B09513A6EFACCD80BFF032382" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.0922" metricValueMin="1.016" pageId="14" pageNumber="15" unit="in" value="41.5" valueMax="43.0" valueMin="40.0">4043 in</quantity>
alatae); in alatae: secondary rhinaria on antennal segment III fewer, with 1416 (
<taxonomicName id="4C584D6D09513A6EFBF6D821FB1A2381" box="[1128,1207,376,401]" class="Insecta" family="Aphididae" genus="Mollitrichosiphum" kingdom="Animalia" order="Aphidomorpha" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rhusae">
<emphasis id="B92CEAFC09513A6EFBF6D821FB1A2381" box="[1128,1207,376,401]" italics="true" pageId="14" pageNumber="15">rhusae</emphasis>
</taxonomicName>
: 20).
</paragraph>
</subSubSection>
</treatment>
</document>