301 lines
31 KiB
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301 lines
31 KiB
XML
<document id="3485CCDA853D449F6F6FC462633E78FB" ID-DOI="10.1649/0010-065X-72.2.331" ID-ISSN="1938-4394" ID-Zenodo-Dep="5382545" ID-ZooBank="urn:lsid:zoobank.org:pub:11BE44A2-4C73-4D36-A761-C9059C9C4222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630295096312" checkinUser="diego" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2018" docId="03F187ED6F1EFF8ADBDCFD4EFBCDFD45" docLanguage="en" docName="0010-065X-72.2.331.pdf" docOrigin="The Coleopterists Bulletin 72 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-72.2.331" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Tomarus maimon Erichson, Lectoallotype, B. C. Ratcliffe 1847" docType="treatment" docVersion="4" lastPageNumber="335" masterDocId="FFC8FF956F1BFF8FDB2BFF98FFFBFFF0" masterDocTitle="Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="338" masterPageNumber="331" pageNumber="335" updateTime="1699218372114" updateUser="plazi">
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<mods:title id="6082A2D914D1344E392D15D025B5D41D">Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<mods:namePart id="B4E9FCDBB7951DFE73319C49331AA56F">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="2B72D4A2E9DBC4F043D424CA4D1732C8">Deloya, Cuauhtémoc</mods:namePart>
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<mods:typeOfResource id="4FA0A60AAFEBB615188F83642D7CC3F6">text</mods:typeOfResource>
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<mods:title id="FF51D88CFCE152064FDB398942C0F701">The Coleopterists Bulletin</mods:title>
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<mods:part id="D0FDF7018E7480FF4E5F9A5B2D708BB3">
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<mods:date id="A9B103DE2B1E46692097B9A39C9D1F6E">2018</mods:date>
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<mods:number id="9C05315D96E59120935C9A2A442C18CE">2018-06-20</mods:number>
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<mods:number id="90A321D117F6F7E42662693A0BB88509">72</mods:number>
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<mods:identifier id="CF75BB4B8BE22F214C0FEF5309C51EA0" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="C34265706F1EFF8ADBDCFD4EFDB2FD1B" box="[247,585,726,747]" pageId="5" pageNumber="335" type="nomenclature">
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<paragraph id="8BE736FB6F1EFF8ADBDCFD4EFDB2FD1B" blockId="5.[247,585,726,747]" box="[247,585,726,747]" pageId="5" pageNumber="335">
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<heading id="D0AF81976F1EFF8ADBDCFD4EFDB2FD1B" box="[247,585,726,747]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="335" reason="2">
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<taxonomicName id="4C584D786F1EFF8ADBDCFD4EFDB2FD1B" authority="Erichson, 1847" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[247,585,726,747]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">
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<emphasis id="B92CEAE96F1EFF8ADBDCFD4EFDB2FD1B" bold="true" box="[247,585,726,747]" pageId="5" pageNumber="335">
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<emphasis id="B92CEAE96F1EFF8ADBDCFD4EFE53FD1B" bold="true" box="[247,424,726,747]" italics="true" pageId="5" pageNumber="335">Tomarus maimon</emphasis>
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Erichson, 1847
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34265706F1EFF8ADB86FD66FE63FCB8" pageId="5" pageNumber="335" type="reference_group">
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<paragraph id="8BE736FB6F1EFF8ADB86FD66FE63FCB8" blockId="5.[173,659,766,840]" pageId="5" pageNumber="335">
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<treatmentCitationGroup id="AB4811D56F1EFF8ADB86FD66FE63FCB8" pageId="5" pageNumber="335">
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<treatmentCitation id="0AF910EA6F1EFF8ADB86FD66FD68FCE3" author="Burmeister, H." box="[173,659,766,787]" page="101" pageId="5" pageNumber="335" year="1847">
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<taxonomicName id="4C584D786F1EFF8ADB86FD66FD68FCE3" authority="Burmeister 1847: 101" authorityName="Burmeister" authorityPageNumber="101" authorityYear="1847" box="[173,659,766,787]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="fossator">
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<emphasis id="B92CEAE96F1EFF8ADB86FD66FE61FCE3" box="[173,410,766,787]" italics="true" pageId="5" pageNumber="335">Heteronychus fossator</emphasis>
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<bibRefCitation id="EFC94B0A6F1EFF8ADA8CFD66FD68FCE3" author="Burmeister, H." box="[423,659,766,787]" pageId="5" pageNumber="335" refId="ref4587" refString="Burmeister, H. 1847. Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister 1847: 101</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(designated as a subspecies by
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<treatmentCitation id="0AF910EA6F1EFF8ADAC3FC80FD7CFCDD" author="Dupuis, F." box="[488,647,792,813]" page="31" pageId="5" pageNumber="335" year="2016">
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<bibRefCitation id="EFC94B0A6F1EFF8ADAC3FC80FD7CFCDD" author="Dupuis, F." box="[488,647,792,813]" pageId="5" pageNumber="335" pagination="30 - 37" refId="ref4870" refString="Dupuis, F. 2016. Tomarus maimon fossator Burmeister, 1847, nouvelle sous-espece du bouclier Guyanais (Coleoptera, Dynastidae). Coleopteres 22 (4): 30 - 37." type="journal article" year="2016">Dupuis 2016: 31</bibRefCitation>
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</treatmentCitation>
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).
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<emphasis id="B92CEAE96F1EFF8ADBECFCABFE63FCB8" bold="true" box="[199,408,819,840]" pageId="5" pageNumber="335">NEW SYNONYMY.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34265706F1EFF8ADBEEFCC3FE09FA73" pageId="5" pageNumber="335" type="distribution">
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<paragraph id="8BE736FB6F1EFF8ADBEEFCC3FE8DFC55" blockId="5.[173,660,859,1730]" pageId="5" pageNumber="335">
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<emphasis id="B92CEAE96F1EFF8ADBEEFCC3FEBDFC80" bold="true" box="[197,326,859,880]" pageId="5" pageNumber="335">Distribution.</emphasis>
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<collectingCountry id="F34F766B6F1EFF8ADA64FCC3FE5BFC80" box="[335,416,859,880]" name="Ecuador" pageId="5" pageNumber="335">Ecuador</collectingCountry>
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,
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<collectingCountry id="F34F766B6F1EFF8ADA87FCC3FE21FC80" box="[428,474,859,880]" name="Peru" pageId="5" pageNumber="335">Peru</collectingCountry>
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,
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<collectingCountry id="F34F766B6F1EFF8ADACDFCC3FDE7FC80" box="[486,540,859,880]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
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,
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<collectingCountry id="F34F766B6F1EFF8AD903FCC3FD9FFC80" box="[552,612,859,880]" name="Brazil" pageId="5" pageNumber="335">Brazil</collectingCountry>
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, and
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<collectingCountry id="F34F766B6F1EFF8ADB86FCEDFF0CFC7A" box="[173,247,885,906]" name="Guyana" pageId="5" pageNumber="335">Guyana</collectingCountry>
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(
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<bibRefCitation id="EFC94B0A6F1EFF8ADA2CFCEDFE6BFC7A" author="Endrodi, S." box="[263,400,885,906]" pageId="5" pageNumber="335" pagination="1 - 145" refId="ref4930" refString="Endrodi, S. 1969. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
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; Ratcliffe and Dechambre 1983;
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<bibRefCitation id="EFC94B0A6F1EFF8ADBC0FC08FE90FC55" author="Dupuis, F." box="[235,363,912,933]" pageId="5" pageNumber="335" pagination="30 - 37" refId="ref4870" refString="Dupuis, F. 2016. Tomarus maimon fossator Burmeister, 1847, nouvelle sous-espece du bouclier Guyanais (Coleoptera, Dynastidae). Coleopteres 22 (4): 30 - 37." type="journal article" year="2016">Dupuis 2016</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BE736FB6F1EFF8ADBEEFC32FE09FA73" blockId="5.[173,660,859,1730]" pageId="5" pageNumber="335">
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<emphasis id="B92CEAE96F1EFF8ADBEEFC32FDB1FC4F" bold="true" box="[197,586,938,959]" pageId="5" pageNumber="335">Specimens Examined. Type material:</emphasis>
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<materialsCitation id="3B303CA66F1EFF8AD97EFC32FD76FB8A" collectionCode="MNHUB" collectorName="B. C. Ratcliffe" pageId="5" pageNumber="335" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
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<typeStatus id="54E388596F1EFF8AD97EFC32FF2DFC2A" pageId="5" pageNumber="335" type="lectotype">Lectotype</typeStatus>
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deposited in
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<collectionCode id="ED49AE3E6F1EFF8ADA79FC5DFE50FC2A" box="[338,427,965,986]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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(16): “Peru, v. Tschidi //
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<typeStatus id="54E388596F1EFF8ADB86FC78FEFDFC05" box="[173,262,992,1013]" pageId="5" pageNumber="335" type="syntype">Syntypus</typeStatus>
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,
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<taxonomicName id="4C584D786F1EFF8ADA24FC78FDBCFC05" authority="Erichson, 1847" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[271,583,992,1013]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson, 1847</taxonomicName>
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, labeled by
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<collectionCode id="ED49AE3E6F1EFF8ADBF9FC62FED0FBFF" box="[210,299,1018,1039]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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, 2013//
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<taxonomicName id="4C584D786F1EFF8ADAAFFC62FD6BFBFF" authority="Erichson" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[388,656,1018,1039]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson</taxonomicName>
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,
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<typeStatus id="54E388596F1EFF8ADB86FB8DFEF8FBDA" box="[173,259,1045,1066]" pageId="5" pageNumber="335" type="lectotype">lectotype</typeStatus>
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,
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<collectorName id="26AD532D6F1EFF8ADA20FB8DFE75FBDA" box="[267,398,1045,1066]" pageId="5" pageNumber="335">B. C. Ratcliffe</collectorName>
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”. Lectoallotype deposited in
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<collectionCode id="ED49AE3E6F1EFF8ADB86FBB7FEFFFBB4" box="[173,260,1071,1092]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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(
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<specimenCount id="9D5EFD726F1EFF8ADA3AFBB7FED3FBB5" box="[273,296,1071,1093]" pageId="5" pageNumber="335" type="female">1♀</specimenCount>
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): “Peru, v. Tschidi //
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<typeStatus id="54E388596F1EFF8ADAC8FBB7FDC7FBB4" box="[483,572,1071,1092]" pageId="5" pageNumber="335" type="syntype">Syntypus</typeStatus>
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,
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<taxonomicName id="4C584D786F1EFF8AD969FBB7FE72FBAF" authority="Erichson, 1847" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson, 1847</taxonomicName>
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, labeled by
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<collectionCode id="ED49AE3E6F1EFF8ADAD9FBD2FDB7FBAF" box="[498,588,1098,1119]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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, 2013 //
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<taxonomicName id="4C584D786F1EFF8ADB86FBFCFD7FFB89" authority="Erichson, Lectoallotype, B. C. Ratcliffe" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[173,644,1124,1145]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson,Lectoallotype,B.C.Ratcliffe</taxonomicName>
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”
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</materialsCitation>
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.
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<materialsCitation id="3B303CA66F1EFF8ADB86FBE7FDD2FB14" collectionCode="MNHUB" pageId="5" pageNumber="335" specimenCount="3" typeStatus="paralectotype">
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<typeStatus id="54E388596F1EFF8ADB86FBE7FECCFB64" box="[173,311,1151,1172]" pageId="5" pageNumber="335" type="paralectotype">Paralectotypes</typeStatus>
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deposited in
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<collectionCode id="ED49AE3E6F1EFF8ADA96FBE7FDEDFB64" box="[445,534,1151,1172]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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(26): “Peru, v. Tschidi //
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<typeStatus id="54E388596F1EFF8ADA0EFB02FE7BFB5F" box="[293,384,1178,1199]" pageId="5" pageNumber="335" type="syntype">Syntypus</typeStatus>
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,
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<taxonomicName id="4C584D786F1EFF8ADAA1FB02FF25FB39" authority="Erichson, 1847" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson, 1847</taxonomicName>
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, labeled by
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<collectionCode id="ED49AE3E6F1EFF8ADA61FB2CFE5FFB39" box="[330,420,1204,1225]" pageId="5" pageNumber="335">MNHUB</collectionCode>
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, 2013 //
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<taxonomicName id="4C584D786F1EFF8ADADAFB2CFF00FB14" authority="Erichson" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">Tomarus maimon Erichson</taxonomicName>
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,
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<typeStatus id="54E388596F1EFF8ADA2BFB57FE8FFB14" box="[256,372,1231,1252]" pageId="5" pageNumber="335" type="paralectotype">Paralectotype</typeStatus>
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,B. C.
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<bibRefCitation id="EFC94B0A6F1EFF8ADA82FB57FDDBFB14" author="Ratcliffe, B. C." box="[425,544,1231,1252]" pageId="5" pageNumber="335" pagination="663 - 680" refId="ref5362" refString="Ratcliffe, B. C. 2014. A new genus and species of Dynastinae (Coleoptera: Scarabaeidae) from the Galapagos Islands, Ecuador, other new species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a revised key to the genera of New World Pentodontini. The Coleopterists Bulletin 68 (4): 663 - 680." type="journal article" year="2014">Ratcliffe 2014</bibRefCitation>
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”
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</materialsCitation>
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.
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<emphasis id="B92CEAE96F1EFF8AD919FB57FEF2FB0E" bold="true" pageId="5" pageNumber="335">Additional material:</emphasis>
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<materialsCitation id="3B303CA66F1EFF8ADA32FB71FD6BFB0E" box="[281,656,1257,1279]" collectingDate="1983-11" collectionCode="FSCA" country="Peru" location="Tacna" pageId="5" pageNumber="335" specimenCount="1" specimenCount-female="1" stateProvince="Tacna">
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<emphasis id="B92CEAE96F1EFF8ADA32FB71FE9AFB0E" bold="true" box="[281,353,1257,1278]" pageId="5" pageNumber="335">
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<collectingCountry id="F34F766B6F1EFF8ADA32FB71FEA6FB0E" box="[281,349,1257,1278]" name="Peru" pageId="5" pageNumber="335">PERU</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="499CF8196F1EFF8ADA5AFB71FE57FB0E" box="[369,428,1257,1278]" country="Peru" name="Tacna" pageId="5" pageNumber="335">Tacna</collectingRegion>
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,
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<date id="FFE6103B6F1EFF8ADA94FB71FDFCFB0E" box="[447,519,1257,1278]" pageId="5" pageNumber="335" value="1983-11">
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<collectingDate id="EFA2E9D36F1EFF8ADA94FB71FDFCFB0E" box="[447,519,1257,1278]" pageId="5" pageNumber="335" value="1983-11">xi.1983</collectingDate>
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</date>
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[
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<specimenCount id="9D5EFD726F1EFF8AD934FB71FDC3FB0F" box="[543,568,1257,1279]" pageId="5" pageNumber="335" type="female">1♀</specimenCount>
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,
|
|
<collectionCode id="ED49AE3E6F1EFF8AD965FB71FD70FB0E" box="[590,651,1257,1278]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="335" type="Museum">FSCA</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3B303CA66F1EFF8ADB86FA9CFEC0FA99" collectingDate="1919-11-20" collectingDateMax="1966-02-27" collectingDateMin="1919-11-20" collectionCode="UFRJ, MZUSP" collectorName="D. Mendes & Col. Malkin & Col. P. Santo" country="Brazil" county="Para" location="Para" municipality="Utinga" pageId="5" pageNumber="335" specimenCount="2" specimenCount-female="2">
|
|
<emphasis id="B92CEAE96F1EFF8ADB86FA9CFEF0FAE9" bold="true" box="[173,267,1284,1305]" pageId="5" pageNumber="335">
|
|
<collectingCountry id="F34F766B6F1EFF8ADB86FA9CFEFCFAE9" box="[173,263,1284,1305]" name="Brazil" pageId="5" pageNumber="335">BRAZIL</collectingCountry>
|
|
:
|
|
</emphasis>
|
|
<collectingCounty id="62864E776F1EFF8ADA3FFA9CFEC6FAE9" box="[276,317,1284,1305]" pageId="5" pageNumber="335">Pará</collectingCounty>
|
|
,
|
|
<collectingMunicipality id="6B83AC816F1EFF8ADA61FA9CFE73FAE9" box="[330,392,1284,1305]" pageId="5" pageNumber="335">Utinga</collectingMunicipality>
|
|
,
|
|
<date id="FFE6103B6F1EFF8ADABEFA9CFE0FFAE9" box="[405,500,1284,1305]" pageId="5" pageNumber="335" value="1919-11-20">
|
|
<collectingDate id="EFA2E9D36F1EFF8ADABEFA9CFE0FFAE9" box="[405,500,1284,1305]" pageId="5" pageNumber="335" value="1919-11-20">20.xi.1919</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="26AD532D6F1EFF8AD929FA9CFD93FAE9" box="[514,616,1284,1305]" pageId="5" pageNumber="335">D. Mendes</collectorName>
|
|
[16 MN/
|
|
<collectionCode id="ED49AE3E6F1EFF8ADBF2FA86FEEBFAC3" box="[217,272,1310,1331]" country="Brazil" name="Universidade Federal Rural do Rio de Janeiro" pageId="5" pageNumber="335" type="Herbarium">UFRJ</collectionCode>
|
|
].
|
|
<location id="8E8760206F1EFF8ADA0FFA86FEB6FAC3" LSID="urn:lsid:plazi:treatment:03F187ED6F1EFF8ADBDCFD4EFBCDFD45:8E8760206F1EFF8ADA0FFA86FEB6FAC3" box="[292,333,1310,1331]" country="Brazil" county="Para" municipality="Utinga" name="Para" pageId="5" pageNumber="335">Pará</location>
|
|
,
|
|
<location id="8E8760206F1EFF8ADA77FA86FE52FAC3" LSID="urn:lsid:plazi:treatment:03F187ED6F1EFF8ADBDCFD4EFBCDFD45:8E8760206F1EFF8ADA77FA86FE52FAC3" box="[348,425,1310,1331]" country="Brazil" county="Para" municipality="Utinga" name="Caninde" pageId="5" pageNumber="335">Canindé</location>
|
|
,
|
|
<location id="8E8760206F1EFF8ADA93FA86FDDDFAC3" LSID="urn:lsid:plazi:treatment:03F187ED6F1EFF8ADBDCFD4EFBCDFD45:8E8760206F1EFF8ADA93FA86FDDDFAC3" box="[440,550,1310,1331]" country="Brazil" county="Para" municipality="Utinga" name="Rio Gurupi" pageId="5" pageNumber="335">Rio Gurupi</location>
|
|
,
|
|
<date id="FFE6103B6F1EFF8AD91EFA86FD74FAC3" box="[565,655,1310,1331]" pageId="5" pageNumber="335" value="1966-02-27">
|
|
<collectingDate id="EFA2E9D36F1EFF8AD91EFA86FD74FAC3" box="[565,655,1310,1331]" pageId="5" pageNumber="335" value="1966-02-27">27.ii.1966</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="26AD532D6F1EFF8ADB86FAA1FEDAFABE" box="[173,289,1337,1358]" pageId="5" pageNumber="335">Col. Malkin</collectorName>
|
|
[
|
|
<specimenCount id="9D5EFD726F1EFF8ADA1AFAA1FEB1FABE" box="[305,330,1336,1358]" pageId="5" pageNumber="335" type="female">1♀</specimenCount>
|
|
,
|
|
<collectionCode id="ED49AE3E6F1EFF8ADA73FAA1FE56FABE" box="[344,429,1337,1358]" country="Brazil" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="335">MZUSP</collectionCode>
|
|
].
|
|
<location id="8E8760206F1EFF8ADA96FAA1FDF2FABE" LSID="urn:lsid:plazi:treatment:03F187ED6F1EFF8ADBDCFD4EFBCDFD45:8E8760206F1EFF8ADA96FAA1FDF2FABE" box="[445,521,1337,1358]" country="Brazil" county="Para" municipality="Utinga" name="Rio Jan" pageId="5" pageNumber="335">Rio Jan</location>
|
|
,
|
|
<collectorName id="26AD532D6F1EFF8AD93FFAA1FD68FABE" box="[532,659,1337,1358]" pageId="5" pageNumber="335">Col. P. Santo</collectorName>
|
|
[
|
|
<specimenCount id="9D5EFD726F1EFF8ADB9EFACCFF36FA99" box="[181,205,1363,1385]" pageId="5" pageNumber="335" type="female">1♀</specimenCount>
|
|
,
|
|
<collectionCode id="ED49AE3E6F1EFF8ADBC9FACCFECCFA99" box="[226,311,1364,1385]" country="Brazil" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="335">MZUSP</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3B303CA66F1EFF8ADA67FACCFE16FA73" collectingDate="1967-12-05" collectionCode="UFRJ" collectorName="Col. J. Neuhausen" country="Chile" location="Arica" pageId="5" pageNumber="335" specimenCount="1" specimenCount-female="1">
|
|
<emphasis id="B92CEAE96F1EFF8ADA67FACCFE5BFA99" bold="true" box="[332,416,1364,1385]" pageId="5" pageNumber="335">
|
|
<collectingCountry id="F34F766B6F1EFF8ADA67FACCFE67FA99" box="[332,412,1364,1385]" name="Chile" pageId="5" pageNumber="335">CHILE</collectingCountry>
|
|
:
|
|
</emphasis>
|
|
<location id="8E8760206F1EFF8ADA85FACCFE1EFA99" LSID="urn:lsid:plazi:treatment:03F187ED6F1EFF8ADBDCFD4EFBCDFD45:8E8760206F1EFF8ADA85FACCFE1EFA99" box="[430,485,1364,1385]" country="Chile" name="Arica" pageId="5" pageNumber="335">Arica</location>
|
|
,
|
|
<date id="FFE6103B6F1EFF8ADADDFACCFDA2FA99" box="[502,601,1364,1385]" pageId="5" pageNumber="335" value="1967-12-05">
|
|
<collectingDate id="EFA2E9D36F1EFF8ADADDFACCFDA2FA99" box="[502,601,1364,1385]" pageId="5" pageNumber="335" value="1967-12-05">5.xii.1967</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="26AD532D6F1EFF8AD941FACCFED6FA73" pageId="5" pageNumber="335">Col. J. Neuhausen</collectorName>
|
|
[16,
|
|
<specimenCount id="9D5EFD726F1EFF8ADA75FAF6FE8CFA74" box="[350,375,1390,1412]" pageId="5" pageNumber="335" type="female">1♀</specimenCount>
|
|
MN/
|
|
<collectionCode id="ED49AE3E6F1EFF8ADA85FAF6FE13FA73" box="[430,488,1390,1411]" country="Brazil" name="Universidade Federal Rural do Rio de Janeiro" pageId="5" pageNumber="335" type="Herbarium">UFRJ</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C34265706F1EFF8ADBEEFA11FBCDFD45" pageId="5" pageNumber="335" type="discussion">
|
|
<paragraph id="8BE736FB6F1EFF8ADBEEFA11FBCDFD45" blockId="5.[173,660,859,1730]" lastBlockId="5.[707,1194,194,693]" pageId="5" pageNumber="335">
|
|
<emphasis id="B92CEAE96F1EFF8ADBEEFA11FECCFA6E" bold="true" box="[197,311,1417,1438]" pageId="5" pageNumber="335">Comments.</emphasis>
|
|
<taxonomicName id="4C584D786F1EFF8ADA6CFA11FE0DFA6E" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[327,502,1417,1438]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">
|
|
<emphasis id="B92CEAE96F1EFF8ADA6CFA11FE0DFA6E" box="[327,502,1417,1438]" italics="true" pageId="5" pageNumber="335">Tomarus maimon</emphasis>
|
|
</taxonomicName>
|
|
was described from
|
|
<collectingCountry id="F34F766B6F1EFF8ADBCDFA3BFEEAFA48" box="[230,273,1443,1464]" name="Peru" pageId="5" pageNumber="335">Peru</collectingCountry>
|
|
and additional records from
|
|
<collectingCountry id="F34F766B6F1EFF8AD914FA3BFD6BFA48" box="[575,656,1443,1464]" name="Ecuador" pageId="5" pageNumber="335">Ecuador</collectingCountry>
|
|
,
|
|
<collectingCountry id="F34F766B6F1EFF8ADB86FA26FF1DFA23" box="[173,230,1470,1491]" name="Brazil" pageId="5" pageNumber="335">Brazil</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EFC94B0A6F1EFF8ADBD2FA26FE7DFA23" author="Endrodi, S." box="[249,390,1470,1491]" pageId="5" pageNumber="335" pagination="1 - 145" refId="ref4930" refString="Endrodi, S. 1969. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
|
|
), and
|
|
<collectingCountry id="F34F766B6F1EFF8ADAE0FA26FE05FA23" box="[459,510,1470,1491]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
|
|
(Ratcliffe and Dechambre 1983) were subsequently published. Its junior synonym,
|
|
<taxonomicName id="4C584D786F1EFF8ADA48FA6BFDBAF9F8" authorityName="Burmeister" authorityYear="1847" box="[355,577,1523,1544]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="fossator">
|
|
<emphasis id="B92CEAE96F1EFF8ADA48FA6BFDBAF9F8" box="[355,577,1523,1544]" italics="true" pageId="5" pageNumber="335">Heteronychus fossator</emphasis>
|
|
</taxonomicName>
|
|
known from
|
|
<collectingCountry id="F34F766B6F1EFF8ADBF4F996FED2F9D3" box="[223,297,1550,1571]" name="Guyana" pageId="5" pageNumber="335">Guyana</collectingCountry>
|
|
and French Guiana, was proposed as a subspecies by
|
|
<bibRefCitation id="EFC94B0A6F1EFF8ADA6EF9B0FE21F9CD" author="Dupuis, F." box="[325,474,1576,1597]" pageId="5" pageNumber="335" pagination="30 - 37" refId="ref4870" refString="Dupuis, F. 2016. Tomarus maimon fossator Burmeister, 1847, nouvelle sous-espece du bouclier Guyanais (Coleoptera, Dynastidae). Coleopteres 22 (4): 30 - 37." type="journal article" year="2016">Dupuis (2016)</bibRefCitation>
|
|
. He considered a disjunct distribution between
|
|
<taxonomicName id="4C584D786F1EFF8ADAF8F9DBFD68F9A8" authority="maimon" box="[467,659,1603,1624]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="subSpecies" species="maimon" subSpecies="maimon">
|
|
<emphasis id="B92CEAE96F1EFF8ADAF8F9DBFD68F9A8" box="[467,659,1603,1624]" italics="true" pageId="5" pageNumber="335">T. maimon maimon</emphasis>
|
|
</taxonomicName>
|
|
and its subspecies, the first from the Andes (
|
|
<collectingCountry id="F34F766B6F1EFF8ADB98F9E0FEFEF97D" box="[179,261,1656,1677]" name="Ecuador" pageId="5" pageNumber="335">Ecuador</collectingCountry>
|
|
,
|
|
<collectingCountry id="F34F766B6F1EFF8ADA25F9E0FEC7F97D" box="[270,316,1656,1677]" name="Peru" pageId="5" pageNumber="335">Peru</collectingCountry>
|
|
, and
|
|
<collectingCountry id="F34F766B6F1EFF8ADA40F9E0FE59F97D" box="[363,418,1656,1677]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
|
|
), and the subspecies from the Guyanese shield. However, there is an omission of the records by
|
|
<bibRefCitation id="EFC94B0A6F1EFF8ADA64F935FE1BF932" author="Endrodi, S." box="[335,480,1709,1730]" pageId="5" pageNumber="335" pagination="1 - 145" refId="ref4930" refString="Endrodi, S. 1969. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
|
|
from
|
|
<collectingRegion id="499CF8196F1EFF8AD933F935FD68F932" box="[536,659,1709,1730]" country="Brazil" name="Mato Grosso" pageId="5" pageNumber="335">Mato Grosso</collectingRegion>
|
|
and Pará (
|
|
<collectingCountry id="F34F766B6F1EFF8AD809FF5AFCA7FF27" box="[802,860,194,215]" name="Brazil" pageId="5" pageNumber="335">Brazil</collectingCountry>
|
|
). Here, additional records from Pará were found in Brazilian collections. The male from Pará has no evident lateral teeth in the parameres, which coincides with
|
|
<taxonomicName id="4C584D786F1EFF8AD860FE89FB53FED6" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[843,1192,273,294]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="subSpecies" species="maimon" subSpecies="fossator">
|
|
<emphasis id="B92CEAE96F1EFF8AD860FE89FC02FED6" box="[843,1017,273,294]" italics="true" pageId="5" pageNumber="335">T. maimon fossator</emphasis>
|
|
(
|
|
<bibRefCitation id="EFC94B0A6F1EFF8ADF2DFE89FB59FED6" author="Burmeister, H." box="[1030,1186,273,294]" pageId="5" pageNumber="335" refId="ref4587" refString="Burmeister, H. 1847. Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister, 1847</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
and differs from the specimens from
|
|
<collectingCountry id="F34F766B6F1EFF8ADF31FEB4FBB8FEB1" box="[1050,1091,300,321]" name="Peru" pageId="5" pageNumber="335">Peru</collectingCountry>
|
|
and
|
|
<collectingCountry id="F34F766B6F1EFF8ADF58FEB4FB5DFEB1" box="[1139,1190,300,321]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
|
|
. However, that difference could be the result of the geographical distance as only specimens from the extremes of the distribution of the species have been recently examined. In any case, a unique character should not differentiate subspecies in the genus, as a character can vary between populations and even more when they are in allopatry.In conclusion, the subspecies
|
|
<taxonomicName id="4C584D786F1EFF8AD9E8FD98FC94FDE5" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[707,879,512,533]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="subSpecies" species="maimon" subSpecies="fossator">
|
|
<emphasis id="B92CEAE96F1EFF8AD9E8FD98FC94FDE5" box="[707,879,512,533]" italics="true" pageId="5" pageNumber="335">T. maimon fossator</emphasis>
|
|
</taxonomicName>
|
|
is here placed into synonymy with
|
|
<taxonomicName id="4C584D786F1EFF8AD9E8FD83FCDEFDC0" authorityName="Erichson, Lectoallotype, B. C. Ratcliffe" authorityYear="1847" box="[707,805,539,560]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="maimon">
|
|
<emphasis id="B92CEAE96F1EFF8AD9E8FD83FCDEFDC0" box="[707,805,539,560]" italics="true" pageId="5" pageNumber="335">T. maimon</emphasis>
|
|
</taxonomicName>
|
|
as there is no clear disruption in the distribution of the taxa, which is likely an artefact of lack of sampling. The records from
|
|
<collectingCountry id="F34F766B6F1EFF8AD8EDFDC8FC0AFD95" box="[966,1009,592,613]" name="Peru" pageId="5" pageNumber="335">Peru</collectingCountry>
|
|
,
|
|
<collectingCountry id="F34F766B6F1EFF8AD8D0FDC8FBBCFD95" box="[1019,1095,592,613]" name="Ecuador" pageId="5" pageNumber="335">Ecuador</collectingCountry>
|
|
, and
|
|
<collectingCountry id="F34F766B6F1EFF8ADF53FDC8FB52FD95" box="[1144,1193,592,613]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
|
|
are mainly from arid and coastal areas and some from tropical forest, so the Amazonian forest should not be a strong geographic barrier for the species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |