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<document id="D61AF3B5A62DC7C37E77486F3F9B2BE0" ID-CLB-Dataset="160067" ID-DOI="10.11646/zootaxa.5194.4.1" ID-GBIF-Dataset="30119506-845f-4008-9c90-209a5369b7f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7156853" ID-ZooBank="EB6859A0-935C-44BF-9B2C-7FDF6FE76793" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665139293186" checkinUser="plazi" docAuthor="Kubicki, Brian, Reyes, Aura &amp; Arias, Erick" docDate="2022" docId="03C687EDFFFCFFC0A4C547A52B50FEBD" docLanguage="en" docName="zootaxa.5194.4.1.pdf" docOrigin="Zootaxa 5194 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nototriton (Taylorotriton) tapanti Good &amp; Wake 1993" docType="treatment" docVersion="7" lastPageNumber="476" masterDocId="FFFFFF95FFE4FFD9A452427C2F1FFFD8" masterDocTitle="Revised taxonomy and distributions of Costa Rican moss salamanders (Caudata: Plethodontidae: Nototriton), with descriptions of new taxa" masterLastPageNumber="496" masterPageNumber="451" pageNumber="475" updateTime="1699389481015" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D3E8087CB1EBA9A5B4F4B3AC69A2B45">Revised taxonomy and distributions of Costa Rican moss salamanders (Caudata: Plethodontidae: Nototriton), with descriptions of new taxa</mods:title>
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<mods:namePart id="F617810295A5819A3CE3BF3C7E956986">Kubicki, Brian</mods:namePart>
<mods:affiliation id="F6901AF9008C782D34C24410A4EC812B">Costa Rican Amphibian Research Center, Guayacán, Provincia de Limón, Costa Rica.</mods:affiliation>
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<mods:namePart id="6CE112B0AAC433778D44A88E56C5246A">Reyes, Aura</mods:namePart>
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<mods:affiliation id="E977001B1DF1D390878808DCEA3740C7">Costa Rican Amphibian Research Center, Guayacán, Provincia de Limón, Costa Rica. &amp; auralr 22 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5845 - 0528</mods:affiliation>
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<mods:namePart id="F9777C7424E1D5524035FD5A3AE9D1D1">Arias, Erick</mods:namePart>
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<mods:affiliation id="BC67F9747A273935D32A4400C096A066">Departamento de Zoología, Instituto de Biología, UNAM, AP 70 - 153 Ciudad Universitaria, CP 04510, México, D. F., México. eapiedra @ gmail. com; https: // orcid. org / 0000 - 0002 - 4449 - 1070 &amp; Escuela de Biología, Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, Costa Rica.</mods:affiliation>
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<mods:title id="0C44E27F6C7AA09C492EBD18BC05473A">Zootaxa</mods:title>
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<treatment id="03C687EDFFFCFFC0A4C547A52B50FEBD" ID-DOI="http://doi.org/10.5281/zenodo.7156825" ID-GBIF-Taxon="202157136" ID-Zenodo-Dep="7156825" LSID="urn:lsid:plazi:treatment:03C687EDFFFCFFC0A4C547A52B50FEBD" httpUri="http://treatment.plazi.org/id/03C687EDFFFCFFC0A4C547A52B50FEBD" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<subSubSection id="C3756570FFFCFFC1A4C547A52EADF9CF" pageId="24" pageNumber="475" type="nomenclature">
<paragraph id="8BD036FBFFFCFFC1A4C547A52C31FA2B" blockId="24.[151,814,1497,1560]" box="[151,814,1497,1524]" pageId="24" pageNumber="475">
<heading id="D0988197FFFCFFC1A4C547A52C31FA2B" bold="true" box="[151,814,1497,1524]" fontSize="11" level="1" pageId="24" pageNumber="475" reason="1">
<taxonomicName id="4C6F4D78FFFCFFC1A4C547A52C31FA2B" ID-CoL="6SDDP" authority="Good &amp; Wake, 1993" authorityName="Good &amp; Wake" authorityYear="1993" box="[151,814,1497,1524]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="tapanti" subGenus="Taylorotriton">
<emphasis id="B91BEAE9FFFCFFC1A4C547A52C31FA2B" bold="true" box="[151,814,1497,1524]" pageId="24" pageNumber="475">
<emphasis id="B91BEAE9FFFCFFC1A4C547A52E0AFA2B" bold="true" box="[151,277,1497,1523]" italics="true" pageId="24" pageNumber="475">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFFCFFC1A57447A52EDAFA2C" bold="true" box="[294,453,1497,1524]" italics="true" pageId="24" pageNumber="475">Taylorotriton</emphasis>
)
<emphasis id="B91BEAE9FFFCFFC1A58447A52D34FA2B" bold="true" box="[470,555,1497,1523]" italics="true" pageId="24" pageNumber="475">tapanti</emphasis>
<bibRefCitation id="EFFE4B0AFFFCFFC1A66047A52C31FA2B" author="Good, D. A. &amp; Wake, D. B." box="[562,814,1497,1523]" pageId="24" pageNumber="475" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD036FBFFFCFFC1A4C547812EADF9CF" blockId="24.[151,814,1497,1560]" box="[151,434,1533,1560]" pageId="24" pageNumber="475">Tapanti moss salamander</paragraph>
</subSubSection>
<subSubSection id="C3756570FFFCFFC1A4C5443E2C3CF9A1" pageId="24" pageNumber="475" type="reference_group">
<paragraph id="8BD036FBFFFCFFC1A4C5443E2D74F982" blockId="24.[151,803,1602,1657]" box="[151,619,1602,1626]" pageId="24" pageNumber="475">
<treatmentCitationGroup id="AB7F11D5FFFCFFC1A4C5443E2D74F982" box="[151,619,1602,1626]" pageId="24" pageNumber="475">
<treatmentCitation id="0ACE10EAFFFCFFC1A4C5443E2D78F982" author="Good, D. A. &amp; Wake, D. B." box="[151,615,1602,1626]" page=")" pageId="24" pageNumber="475" year="1993">
<taxonomicName id="4C6F4D78FFFCFFC1A4C5443E2D78F982" ID-CoL="6SDDP" authority="Good &amp; Wake (1993: 134)" authorityName="Good &amp; Wake" authorityYear="1993" box="[151,615,1602,1626]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="tapanti">
<emphasis id="B91BEAE9FFFCFFC1A4C5443E2E50F981" box="[151,335,1602,1625]" italics="true" pageId="24" pageNumber="475">Nototriton tapanti</emphasis>
<bibRefCitation id="EFFE4B0AFFFCFFC1A507443E2D78F982" author="Good, D. A. &amp; Wake, D. B." box="[341,615,1602,1626]" pageId="24" pageNumber="475" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake (1993: 134)</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BD036FBFFFCFFC1A4C5441D2C3CF9A1" blockId="24.[151,803,1602,1657]" box="[151,803,1632,1657]" pageId="24" pageNumber="475">
<treatmentCitationGroup id="AB7F11D5FFFCFFC1A4C5441D2C3CF9A1" box="[151,803,1632,1657]" pageId="24" pageNumber="475">
<treatmentCitation id="0ACE10EAFFFCFFC1A4C5441D2C00F9A1" author="Dubois, A. &amp; Raffaelli, J." box="[151,799,1632,1657]" page=")" pageId="24" pageNumber="475" year="2012">
<taxonomicName id="4C6F4D78FFFCFFC1A4C5441D2C00F9A1" ID-CoL="6SDDP" authority=": Dubois &amp; Raffaelli (2012: 141)" authorityName=": Dubois &amp; Raffaelli" authorityPageNumber="141" authorityYear="2012" box="[151,799,1632,1657]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="tapanti" subGenus="Nototriton">
<emphasis id="B91BEAE9FFFCFFC1A4C5441D2E1EF9A0" box="[151,257,1633,1656]" italics="true" pageId="24" pageNumber="475">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFFCFFC1A542441D2E65F9A0" box="[272,378,1633,1656]" italics="true" pageId="24" pageNumber="475">Nototriton</emphasis>
)
<emphasis id="B91BEAE9FFFCFFC1A5DA441D2ECFF9A0" box="[392,464,1633,1656]" italics="true" pageId="24" pageNumber="475">tapanti</emphasis>
:
<bibRefCitation id="EFFE4B0AFFFCFFC1A58F441C2C00F9A1" author="Dubois, A. &amp; Raffaelli, J." box="[477,799,1632,1657]" pageId="24" pageNumber="475" pagination="77 - 161" refId="ref26197" refString="Dubois, A. &amp; Raffaelli, J. (2012) A new ergotaxonomy of the order Urodela Dumeril, 1805 (Amphibia, Batrachia). Alytes, 28, 77 - 161." type="journal article" year="2012">Dubois &amp; Raffaëlli (2012: 141)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C3756570FFFCFFC1A4C544DB2EC0F8D1" pageId="24" pageNumber="475" type="materials_examined">
<paragraph id="8BD036FBFFFCFFC1A4C544DB2EC0F8D1" blockId="24.[151,1437,1703,2017]" pageId="24" pageNumber="475">
<materialsCitation id="3B073CA6FFFCFFC1A4C544DB2EC0F8D1" ID-GBIF-Occurrence="3921375308" collectingDate="1986-05-25" collectionCode="MVZ" collectorName="A. Collazo &amp; D. A. Good" country="Costa Rica" county="Canton de Paraiso" elevation="1300" location="Rio Quiri" municipality="Distrito de Orosi" pageId="24" pageNumber="475" specimenCode="MVZ 203746" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<emphasis id="B91BEAE9FFFCFFC1A4C544DB2E14F919" bold="true" box="[151,267,1703,1729]" pageId="24" pageNumber="475">
<typeStatus id="54D48859FFFCFFC1A4C544DB2E19F919" box="[151,262,1703,1729]" pageId="24" pageNumber="475" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBC99E80FFFCFFC1A54144DB2EAFF919" box="[275,432,1703,1729]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="24" pageNumber="475" type="Museum">MVZ 203746</specimenCode>
, by original designation, an adult female from
<collectingCountry id="F378766BFFFCFFC1A79144DB2B5CF919" box="[963,1091,1703,1729]" name="Costa Rica" pageId="24" pageNumber="475">Costa Rica</collectingCountry>
:
<collectingRegion id="49ABF819FFFCFFC1A01C44DB2A5FF919" box="[1102,1344,1703,1729]" country="Costa Rica" name="Cartago" pageId="24" pageNumber="475">Provincia de Cartago</collectingRegion>
:
<collectingCounty id="62B14E77FFFCFFC1A11944DB2E0FF93D" pageId="24" pageNumber="475">
Cantón
<collectorName id="269A532DFFFCFFC1A4C544B72E0FF93D" box="[151,272,1739,1765]" pageId="24" pageNumber="475">de Paraíso</collectorName>
</collectingCounty>
:
<collectingMunicipality id="6BB4AC81FFFCFFC1A54944B72EC2F93D" box="[283,477,1739,1765]" pageId="24" pageNumber="475">
Distrito
<collectorName id="269A532DFFFCFFC1A52A44B72EC2F93D" box="[376,477,1739,1765]" pageId="24" pageNumber="475">de Orosi</collectorName>
</collectingMunicipality>
:
<location id="8EB06020FFFCFFC1A5B544B72D4AF93D" LSID="urn:lsid:plazi:treatment:03C687EDFFFCFFC0A4C547A52B50FEBD:8EB06020FFFCFFC1A5B544B72D4AF93D" box="[487,597,1739,1765]" country="Costa Rica" county="Canton de Paraiso" municipality="Distrito de Orosi" name="Rio Quiri" pageId="24" pageNumber="475" stateProvince="Cartago">Río Quirí</location>
, near
<location id="8EB06020FFFCFFC1A6CA44B72C8EF93D" LSID="urn:lsid:plazi:treatment:03C687EDFFFCFFC0A4C547A52B50FEBD:8EB06020FFFCFFC1A6CA44B72C8EF93D" box="[664,913,1739,1765]" country="Costa Rica" county="Canton de Paraiso" municipality="Distrito de Orosi" name="Tapanti National Park" pageId="24" pageNumber="475" stateProvince="Cartago">Tapanti National Park</location>
, ca.
<elevation id="0042D1C8FFFCFFC1A79644B72B48F93D" box="[964,1111,1739,1765]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="475" unit="m" value="1300.0">
<quantity id="4C979B1EFFFCFFC1A79644B72B05F93D" box="[964,1050,1739,1765]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="475" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
, collected by
<collectorName id="269A532DFFFCFFC1A0A044B72A73F93D" box="[1266,1388,1739,1765]" pageId="24" pageNumber="475">A. Collazo</collectorName>
and
<collectorName id="269A532DFFFCFFC1A4C544932E07F8D1" box="[151,280,1775,1801]" pageId="24" pageNumber="475">D. A. Good</collectorName>
on
<date id="FFD1103BFFFCFFC1A51044922EC4F8D1" box="[322,475,1774,1801]" pageId="24" pageNumber="475" value="1986-05-25">
<collectingDate id="EF95E9D3FFFCFFC1A51044922EC4F8D1" box="[322,475,1774,1801]" pageId="24" pageNumber="475" value="1986-05-25">25 May 1986</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFFCFFC1A495456F2A6BF889" pageId="24" pageNumber="475" type="etymology">
<paragraph id="8BD036FBFFFCFFC1A495456F2A6BF889" blockId="24.[151,1437,1703,2017]" pageId="24" pageNumber="475">
<emphasis id="B91BEAE9FFFCFFC1A495456F2E51F8F5" bold="true" box="[199,334,1811,1837]" pageId="24" pageNumber="475">Etymology.</emphasis>
The specific epithet is a noun in apposition and was given in recognition of the site where the
<typeStatus id="54D48859FFFCFFC1A4C5454B2FE6F889" box="[151,249,1847,1873]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
of this taxon was collected, a small settlement near what is presently known as Tapantí National Park.
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFFCFFC0A49545262B50FEBD" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475" type="distribution">
<paragraph id="8BD036FBFFFCFFC0A49545262DE9FEC5" blockId="24.[151,1437,1703,2017]" lastBlockId="25.[151,1436,150,357]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<emphasis id="B91BEAE9FFFCFFC1A49545262E7EF8AC" bold="true" box="[199,353,1882,1908]" pageId="24" pageNumber="475">Distribution.</emphasis>
<taxonomicName id="4C6F4D78FFFCFFC1A53845272D29F8AC" authorityName=": Dubois &amp; Raffaelli" authorityYear="2012" box="[362,566,1883,1908]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="tapanti">
<emphasis id="B91BEAE9FFFCFFC1A53845272D29F8AC" box="[362,566,1883,1908]" italics="true" pageId="24" pageNumber="475">Nototriton tapanti</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F378766BFFFCFFC1A6B145272C40F8AD" box="[739,863,1883,1909]" name="Costa Rica" pageId="24" pageNumber="475">Costa Rica</collectingCountry>
and was only known from a single specimen, the
<typeStatus id="54D48859FFFCFFC1A4C545032FE3F841" box="[151,252,1919,1945]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
, collected near Río Quirí on the northern section of the Cordillera de Talamanca, but during this study BK discovered and collected additional specimens that resulted in extending the known range approximately four kilometers to the south of the type locality within Tapanti National Park (ca.
<quantity id="4C979B1EFFFCFFC1A7BD45BB2B5BF839" box="[1007,1092,1991,2017]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="475" unit="m" value="1400.0">1400 m</quantity>
a.s.l.) and 30 kilometers to the NNE along the eastern flank of the Turrialba Volcano (ca.
<quantity id="4C979B1EFFFDFFC0A77842EA2C9FFF69" box="[810,896,150,177]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="25" pageNumber="476" unit="m" value="2250.0">2250 m</quantity>
a.s.l.), which is part of the Cordillera Volcanica Central. With the additional individuals of this poorly known taxon discovered during this study,
<taxonomicName id="4C6F4D78FFFDFFC0A08042C72A83FF0C" authorityName=": Dubois &amp; Raffaelli" authorityYear="2012" box="[1234,1436,187,212]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="tapanti">
<emphasis id="B91BEAE9FFFDFFC0A08042C72A83FF0C" box="[1234,1436,187,212]" italics="true" pageId="25" pageNumber="476">Nototriton tapanti</emphasis>
</taxonomicName>
is now known to inhabit sites within the northern Cordillera de Talamanca and extreme eastern Cordillera Volcanica Central at elevations from roughly
<quantity id="4C979B1EFFFDFFC0A672437E2DA4FEC5" box="[544,699,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.775" metricValueMax="2.25" metricValueMin="1.3" pageId="25" pageNumber="476" unit="m" value="1775.0" valueMax="2250.0" valueMin="1300.0">1300–2250 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BD036FBFFFDFFC0A495435B2B50FEBD" blockId="25.[151,1436,150,357]" pageId="25" pageNumber="476">
<emphasis id="B91BEAE9FFFDFFC0A495435B2D8AFE99" bold="true" box="[199,661,294,321]" pageId="25" pageNumber="476">Noteworthy observations or remarks.</emphasis>
<taxonomicName id="4C6F4D78FFFDFFC0A6FE435B2C64FE98" authorityName=": Dubois &amp; Raffaelli" authorityYear="2012" box="[684,891,295,320]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="tapanti">
<emphasis id="B91BEAE9FFFDFFC0A6FE435B2C64FE98" box="[684,891,295,320]" italics="true" pageId="25" pageNumber="476">Nototriton tapanti</emphasis>
</taxonomicName>
was only known from a single specimen, the
<typeStatus id="54D48859FFFDFFC0A4C543372FE3FEBD" box="[151,252,331,357]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
, until additional specimens were discovered and collected during this study.
</paragraph>
</subSubSection>
</treatment>
</document>
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