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<document id="3DBDE66B5BDD8C49DADFB61C9D3C172A" ID-CLB-Dataset="160067" ID-DOI="10.11646/zootaxa.5194.4.1" ID-GBIF-Dataset="30119506-845f-4008-9c90-209a5369b7f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7156853" ID-ZooBank="EB6859A0-935C-44BF-9B2C-7FDF6FE76793" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665139293186" checkinUser="plazi" docAuthor="Kubicki, Brian, Reyes, Aura &amp; Arias, Erick" docDate="2022" docId="03C687EDFFC2FFFFA4C542EB2D1DFDA3" docLanguage="en" docName="zootaxa.5194.4.1.pdf" docOrigin="Zootaxa 5194 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nototriton (Nototriton) guanacaste Good &amp; Wake 1993" docType="treatment" docVersion="7" lastPageNumber="489" masterDocId="FFFFFF95FFE4FFD9A452427C2F1FFFD8" masterDocTitle="Revised taxonomy and distributions of Costa Rican moss salamanders (Caudata: Plethodontidae: Nototriton), with descriptions of new taxa" masterLastPageNumber="496" masterPageNumber="451" pageNumber="489" updateTime="1699389481015" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D5E3096E89439090B6F28019DBF9B9A5">Revised taxonomy and distributions of Costa Rican moss salamanders (Caudata: Plethodontidae: Nototriton), with descriptions of new taxa</mods:title>
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<mods:namePart id="6265917E5A8BC7608298D1A31FAF0FC3">Kubicki, Brian</mods:namePart>
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<mods:namePart id="F837810A060D4BB57491BD125AAB4ED5">Reyes, Aura</mods:namePart>
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<mods:affiliation id="DB25F5BDE82E9F8BB0BDD6F37AF3B226">Costa Rican Amphibian Research Center, Guayacán, Provincia de Limón, Costa Rica. &amp; auralr 22 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5845 - 0528</mods:affiliation>
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<mods:namePart id="8BA318EC8600DC0AC8743EF6FB5CE28D">Arias, Erick</mods:namePart>
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<mods:affiliation id="547DD2815C99A9E6B83948170C97D7D2">Departamento de Zoología, Instituto de Biología, UNAM, AP 70 - 153 Ciudad Universitaria, CP 04510, México, D. F., México. eapiedra @ gmail. com; https: // orcid. org / 0000 - 0002 - 4449 - 1070 &amp; Escuela de Biología, Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, Costa Rica.</mods:affiliation>
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<treatment id="03C687EDFFC2FFFFA4C542EB2D1DFDA3" ID-DOI="http://doi.org/10.5281/zenodo.7156849" ID-GBIF-Taxon="202157139" ID-Zenodo-Dep="7156849" LSID="urn:lsid:plazi:treatment:03C687EDFFC2FFFFA4C542EB2D1DFDA3" httpUri="http://treatment.plazi.org/id/03C687EDFFC2FFFFA4C542EB2D1DFDA3" lastPageNumber="489" pageId="38" pageNumber="489">
<subSubSection id="C3756570FFC2FFFFA4C542EB2EFFFF0D" pageId="38" pageNumber="489" type="nomenclature">
<paragraph id="8BD036FBFFC2FFFFA4C542EB2C5EFF69" blockId="38.[151,833,151,214]" box="[151,833,151,177]" pageId="38" pageNumber="489">
<heading id="D0988197FFC2FFFFA4C542EB2C5EFF69" bold="true" box="[151,833,151,177]" fontSize="11" level="1" pageId="38" pageNumber="489" reason="1">
<taxonomicName id="4C6F4D78FFC2FFFFA4C542EB2C5EFF69" ID-CoL="6SDDW" authority="Good &amp; Wake, 1993" authorityName="Good &amp; Wake" authorityYear="1993" box="[151,833,151,177]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="guanacaste" subGenus="Nototriton">
<emphasis id="B91BEAE9FFC2FFFFA4C542EB2C5EFF69" bold="true" box="[151,833,151,177]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA4C542EB2E0AFF69" bold="true" box="[151,277,151,177]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFC2FFFFA57442EB2EBBFF69" bold="true" box="[294,420,151,177]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
)
<emphasis id="B91BEAE9FFC2FFFFA5E642EB2D22FF69" bold="true" box="[436,573,151,177]" italics="true" pageId="38" pageNumber="489">guanacaste</emphasis>
<bibRefCitation id="EFFE4B0AFFC2FFFFA61742EB2C5EFF69" author="Good, D. A. &amp; Wake, D. B." box="[581,833,151,177]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake, 1993</bibRefCitation>
</emphasis>
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</paragraph>
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<collectingRegion id="49ABF819FFC2FFFFA4C542C72E07FF0D" box="[151,280,187,213]" country="Costa Rica" name="Guanacaste" pageId="38" pageNumber="489">Guanacaste</collectingRegion>
moss salamander
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA4C5437C2C51FEEE" pageId="38" pageNumber="489" type="reference_group">
<paragraph id="8BD036FBFFC2FFFFA4C5437C2D89FEC0" blockId="38.[151,846,256,310]" box="[151,662,256,280]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C5437C2D89FEC0" box="[151,662,256,280]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C5437C2D8DFEC0" author="Good, D. A. &amp; Wake, D. B." box="[151,658,256,280]" page=")" pageId="38" pageNumber="489" year="1993">
<taxonomicName id="4C6F4D78FFC2FFFFA4C5437C2D8DFEC0" ID-CoL="6SDDW" authority="Good &amp; Wake (1993: 138)" authorityName="Good &amp; Wake" authorityYear="1993" box="[151,658,256,280]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="guanacaste">
<emphasis id="B91BEAE9FFC2FFFFA4C5437C2E65FECF" box="[151,378,256,279]" italics="true" pageId="38" pageNumber="489">Nototriton guanacaste</emphasis>
<bibRefCitation id="EFFE4B0AFFC2FFFFA5D3437C2D8DFEC0" author="Good, D. A. &amp; Wake, D. B." box="[385,658,256,280]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake (1993: 138)</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C543632C51FEEE" box="[151,846,286,310]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C543632C55FEEE" author="Dubois, A. &amp; Raffaelli, J." box="[151,842,286,310]" page=")" pageId="38" pageNumber="489" year="2012">
<taxonomicName id="4C6F4D78FFC2FFFFA4C543632C55FEEE" ID-CoL="6SDDW" authority=": Dubois &amp; Raffaelli (2012: 141)" authorityName="Dubois &amp; Raffaelli" authorityPageNumber="141" authorityYear="2012" box="[151,842,286,310]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="guanacaste" subGenus="Nototriton">
<emphasis id="B91BEAE9FFC2FFFFA4C543632E1EFEEE" box="[151,257,287,310]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFC2FFFFA54243632E65FEEE" box="[272,378,287,310]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
)
<emphasis id="B91BEAE9FFC2FFFFA5DA43632EE4FEEE" box="[392,507,287,310]" italics="true" pageId="38" pageNumber="489">guanacaste</emphasis>
:
<bibRefCitation id="EFFE4B0AFFC2FFFFA65B43622C55FEEE" author="Dubois, A. &amp; Raffaelli, J." box="[521,842,286,310]" pageId="38" pageNumber="489" pagination="77 - 161" refId="ref26197" refString="Dubois, A. &amp; Raffaelli, J. (2012) A new ergotaxonomy of the order Urodela Dumeril, 1805 (Amphibia, Batrachia). Alytes, 28, 77 - 161." type="journal article" year="2012">Dubois &amp; Raffaëlli (2012: 141)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BD036FBFFC2FFFFA4C543192B6BFE1F" blockId="38.[151,1437,356,635]" pageId="38" pageNumber="489">
<materialsCitation id="3B073CA6FFC2FFFFA4C543192B6BFE1F" ID-GBIF-Occurrence="3921375332" collectingDate="1987-08-24" collectionCode="MVZ" collectorName="D. C. Cannattela &amp; D. A. Good" country="Costa Rica" county="Distrito de Mayorga" elevation="1580" location="Cacao Volcano" municipality="Guanacaste National Park" pageId="38" pageNumber="489" specimenCode="MVZ 207111" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="B91BEAE9FFC2FFFFA4C543192E14FEA7" bold="true" box="[151,267,357,383]" pageId="38" pageNumber="489">
<typeStatus id="54D48859FFC2FFFFA4C543192E19FEA7" box="[151,262,357,383]" pageId="38" pageNumber="489" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBC99E80FFC2FFFFA54043192EB3FEA7" box="[274,428,357,383]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="38" pageNumber="489" type="Museum">MVZ 207111</specimenCode>
, by original designation, an adult male from
<collectingCountry id="F378766BFFC2FFFFA7F243192B00FEA6" box="[928,1055,356,383]" name="Costa Rica" pageId="38" pageNumber="489">Costa Rica</collectingCountry>
:
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Provincia de
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</collectingRegion>
: Cantón de
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:
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Distrito
<collectorName id="269A532DFFC2FFFFA5D243F42D11FE7B" box="[384,526,392,419]" pageId="38" pageNumber="489">de Mayorga</collectorName>
</collectingCounty>
:
<collectingMunicipality id="6BB4AC81FFC2FFFFA64943F52C55FE7A" box="[539,842,392,419]" pageId="38" pageNumber="489">Guanacaste National Park</collectingMunicipality>
: upper slopes of the
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,
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a.s.l.
</elevation>
, collected by
<collectorName id="269A532DFFC2FFFFA57443D12EFEFE1E" box="[294,481,428,455]" pageId="38" pageNumber="489">D. C. Cannattela</collectorName>
and
<collectorName id="269A532DFFC2FFFFA64543D12D87FE1E" box="[535,664,428,455]" pageId="38" pageNumber="489">D. A. Good</collectorName>
on
<date id="FFD1103BFFC2FFFFA69043D12C6BFE1F" box="[706,884,429,455]" pageId="38" pageNumber="489" value="1987-08-24">
<collectingDate id="EF95E9D3FFC2FFFFA69043D12C6BFE1F" box="[706,884,429,455]" pageId="38" pageNumber="489" value="1987-08-24">24 August 1987</collectingDate>
</date>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA7D043D02B77FE1F" author="Good, D. A. &amp; Wake, D. B." box="[898,1128,428,455]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake 1993</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA49543AD2BB1FDD7" pageId="38" pageNumber="489" type="etymology">
<paragraph id="8BD036FBFFC2FFFFA49543AD2BB1FDD7" blockId="38.[151,1437,356,635]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA49543AD2E51FE33" bold="true" box="[199,334,465,491]" pageId="38" pageNumber="489">Etymology.</emphasis>
The specific epithet is a noun in apposition and was chosen in celebration of establishment of Guanacaste National Park, where this species is known to be restricted (
<bibRefCitation id="EFFE4B0AFFC2FFFFA7EF43882BBDFDD7" author="Good, D. A. &amp; Wake, D. B." box="[957,1186,500,527]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">
Good &amp;
<collectingCountry id="F378766BFFC2FFFFA07243882B40FDD6" box="[1056,1119,500,526]" name="Wake Island" pageId="38" pageNumber="489">Wake</collectingCountry>
1993
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA49540642D1DFDA3" pageId="38" pageNumber="489" type="distribution">
<paragraph id="8BD036FBFFC2FFFFA49540642D1DFDA3" blockId="38.[151,1437,356,635]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA49540642E7EFDEA" bold="true" box="[199,353,536,562]" pageId="38" pageNumber="489">Distribution.</emphasis>
<taxonomicName id="4C6F4D78FFC2FFFFA53440652D7EFDEA" box="[358,609,537,562]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="guanacaste">
<emphasis id="B91BEAE9FFC2FFFFA53440652D7EFDEA" box="[358,609,537,562]" italics="true" pageId="38" pageNumber="489">Nototriton guanacaste</emphasis>
</taxonomicName>
is currently only known to inhabit two small patches of cloud forest within
<collectingRegion id="49ABF819FFC2FFFFA4C540412E07FD8F" box="[151,280,573,599]" country="Costa Rica" name="Guanacaste" pageId="38" pageNumber="489">Guanacaste</collectingRegion>
National Park, both located on the upper slopes of the Orosi and Cacao volcanoes between roughly
<quantity id="4C979B1EFFC2FFFFA4C5401C2E4EFDA2" box="[151,337,608,635]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="1.65" metricValueMin="1.4" pageId="38" pageNumber="489" unit="m" value="1525.0" valueMax="1650.0" valueMin="1400.0">1400–1650 masl</quantity>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA532401D2EE9FDA3" author="Savage, J. M." box="[352,502,609,635]" pageId="38" pageNumber="489" refId="ref27408" refString="Savage, J. M. (2002) The Amphibians and Reptiles of Costa Rica: A Herpetofauna between Two Continents, between Two Seas. The University of Chicago Press, Chicago, Illinois, 934 pp." type="book" year="2002">Savage 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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