155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="03231B7CDA7F2690EE4F87A360F9715B" ID-DOI="10.5281/zenodo.276822" ID-GBIF-Dataset="f5489381-8fe8-48a0-9fb6-5addeebaa3f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="276822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237113383" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03C687E4FFD4FF99FF1BEF3C1296FCDE" docLanguage="en" docName="zt02763p033.pdf" docOrigin="Zootaxa 2763" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hlavaciellus bifoveatus Jałoszyński, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="13" masterDocId="FFFFFF9CFFDFFF95FF8CEB58101EFFCA" masterDocTitle="Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="12" updateTime="1698239820214" updateUser="plazi">
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<mods:title id="5C5DD05EE7EA28414C198283D3A2018F">Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="E8469189186118AB235165D999113A93" type="personal">
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<mods:roleTerm id="FA7043D7914CD9FCE315120FD048BA5B">Author</mods:roleTerm>
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<mods:namePart id="56D4FB9986FCBF19B15A621BC5AB3667">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="391EA33EE883116B0A837A8258D96816">text</mods:typeOfResource>
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<mods:title id="ACE9164F293B6156C0FBF548690ECE81">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="967C20EBE960FD9E674DAC74B8A4F3DC">
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<mods:date id="1017CB411128634D79A0F536055FEB90">2011</mods:date>
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<mods:detail id="4A52187C191A60BC0C0250B2274A1B5B" type="volume">
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<mods:number id="AE7F9775AACEC516D9BFF90AEBE45DE1">2763</mods:number>
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<mods:start id="98BEE36B742635A53C97A0F5E52FC2AC">1</mods:start>
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<mods:classification id="C7C1BF6FCCFF34633B198A6EE371ED0C">journal article</mods:classification>
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<mods:identifier id="82567C19C28ED536DC0C8F1C34CAA8CB" type="DOI">10.5281/zenodo.276822</mods:identifier>
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<mods:identifier id="C7478EEB8E04673A99C0CE0D1AC31FA2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C687E4FFD4FF99FF1BEF3C1296FCDE" ID-DOI="http://doi.org/10.5281/zenodo.6192607" ID-GBIF-Taxon="119388734" ID-Zenodo-Dep="6192607" LSID="urn:lsid:plazi:treatment:03C687E4FFD4FF99FF1BEF3C1296FCDE" httpUri="http://treatment.plazi.org/id/03C687E4FFD4FF99FF1BEF3C1296FCDE" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<subSubSection id="C3756579FFD4FF9EFF1BEF3C116AFB54" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BD036F2FFD4FF9EFF1BEF3C11E4FBB7" blockId="11.[151,506,1123,1182]" box="[151,506,1123,1150]" pageId="11" pageNumber="12">
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<heading id="D098819EFFD4FF9EFF1BEF3C11E4FBB7" bold="true" box="[151,506,1123,1150]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<emphasis id="B91BEAE0FFD4FF9EFF1BEF3C11E4FBB7" bold="true" box="[151,506,1123,1150]" pageId="11" pageNumber="12">
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<taxonomicName id="4C6F4D71FFD4FF9EFF1BEF3C11AFFBB4" ID-CoL="3M5PQ" box="[151,433,1124,1150]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bifoveatus" status="sp. nov.">
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<emphasis id="B91BEAE0FFD4FF9EFF1BEF3C11AFFBB4" bold="true" box="[151,433,1124,1150]" italics="true" pageId="11" pageNumber="12">Hlavaciellus bifoveatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228579BFFD4FF9EFE34EF3B11E4FBB7" box="[440,506,1123,1149]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD036F2FFD4FF9EFF1BEFDE116AFB54" blockId="11.[151,506,1123,1182]" box="[151,372,1158,1182]" pageId="11" pageNumber="12">
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(
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<figureCitation id="13542A77FFD4FF9EFF13EFDE10F0FB54" box="[159,238,1158,1182]" captionStart="FIGURES 4 – 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="11" pageNumber="12">Figs. 9</figureCitation>
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,
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<figureCitation id="13542A77FFD4FF9EFF76EFDE1107FB54" box="[250,281,1158,1182]" captionStart="FIGURES 24 – 26" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[166,1429,853,1827]" captionTargetId="figure@12.[151,1436,844,1840]" captionTargetPageId="12" captionText="FIGURES 24 – 26. Dorsal view of holotypes of Hlavaciellus, males. 24, H. bifoveatus sp. n. 25, H. metrios sp. n. 26, H. miser sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276827/files/figure.png" pageId="11" pageNumber="12">24</figureCitation>
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, 31, 32)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD4FF9EFF1BEF88152FFAFA" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8BD036F2FFD4FF9EFF1BEF88152FFAFA" blockId="11.[151,1436,1231,2014]" pageId="11" pageNumber="12">
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<emphasis id="B91BEAE0FFD4FF9EFF1BEF881274FB22" bold="true" box="[151,618,1231,1256]" pageId="11" pageNumber="12">
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<typeStatus id="54D48850FFD4FF9EFF1BEF8810CCFB22" box="[151,210,1232,1256]" pageId="11" pageNumber="12">Type</typeStatus>
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material.
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<typeStatus id="54D48850FFD4FF9EFEC1EF8811A0FB22" box="[333,446,1232,1256]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3787662FFD4FF9EFE47EF97127BFB22" box="[459,613,1231,1256]" name="Malaysia" pageId="11" pageNumber="12">MALAYSIA</collectingCountry>
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:
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</emphasis>
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ɗ, "
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<collectionCode id="ED7EAE37FFD4FF9EFD11EF97131DFB2D" box="[669,771,1231,1255]" pageId="11" pageNumber="12">SABAH</collectionCode>
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: Crocker Ra. \
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<quantity id="4C979B17FFD4FF9EFC22EF971418FB2D" box="[942,1030,1231,1255]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="1600.0">1600 m</quantity>
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, km 51 rte \ Kota Kinabalu– \ Tambunan,
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<date id="FFD11032FFD4FF9EFF66EFAC115FFAC6" box="[234,321,1268,1292]" pageId="11" pageNumber="12" value="1987-05-18">18.V.87</date>
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\ Burckhardt - Löbl" [white, printed], "
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<taxonomicName id="4C6F4D71FFD4FF9EFC8FEFAC13DBFAC6" box="[771,965,1268,1292]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BEAE0FFD4FF9EFC8FEFAC13DBFAC6" box="[771,965,1268,1292]" italics="true" pageId="11" pageNumber="12">HLAVACIELLUS</emphasis>
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</taxonomicName>
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\
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<taxonomicName id="4C6F4D71FFD4FF9EFC51EFAC144EFAC6" box="[989,1104,1268,1292]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bifoveatus">
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<emphasis id="B91BEAE0FFD4FF9EFC51EFAC144EFAC6" box="[989,1104,1268,1292]" italics="true" pageId="11" pageNumber="12">bifoveatus</emphasis>
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</taxonomicName>
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m. \ det. P. JAŁOSZYŃSKI, '09 \
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<typeStatus id="54D48850FFD4FF9EFF42EE401169FAFA" box="[206,375,1304,1328]" pageId="11" pageNumber="12" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (
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<collectionCode id="ED7EAE37FFD4FF9EFDA5EE40129BFAFA" box="[553,645,1304,1328]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="12">MHNG</collectionCode>
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).
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<emphasis id="B91BEAE0FFD4FF9EFD1BEE4113AAFAFB" bold="true" box="[663,948,1304,1329]" pageId="11" pageNumber="12">
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<typeStatus id="54D48850FFD4FF9EFD1BEE411316FAFB" box="[663,776,1305,1329]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
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:
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<collectingCountry id="F3787662FFD4FF9EFC99EE4013B1FAFB" box="[789,943,1304,1329]" name="Malaysia" pageId="11" pageNumber="12">MALAYSIA</collectingCountry>
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:
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</emphasis>
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ɗ, same data as
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<typeStatus id="54D48850FFD4FF9EFBFDEE4014CDFAFA" box="[1137,1235,1304,1328]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED7EAE37FFD4FF9EFB6DEE40153AFAFA" box="[1249,1316,1304,1328]" pageId="11" pageNumber="12">PCPJ</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD4FF9EFF4BEE651281FAB3" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BD036F2FFD4FF9EFF4BEE651281FAB3" blockId="11.[151,1436,1231,2014]" pageId="11" pageNumber="12">
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<emphasis id="B91BEAE0FFD4FF9EFF4BEE65115FFA9C" bold="true" box="[199,321,1341,1366]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
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Vertex in males with pair of shallow transverse lateral impressions broadly separated in middle; antennomere III nearly twice as long as broad.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD4FF99FF4BEEDD11BCFD6D" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
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<paragraph id="8BD036F2FFD4FF9EFF4BEEDD1120FA0B" blockId="11.[151,1436,1231,2014]" pageId="11" pageNumber="12">
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<emphasis id="B91BEAE0FFD4FF9EFF4BEEDD1147FA54" bold="true" box="[199,345,1413,1438]" pageId="11" pageNumber="12">Description.</emphasis>
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BL 2.05–2.18 (2.11). Body of male (
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<figureCitation id="13542A77FFD4FF9EFC89EEDD1346FA57" box="[773,856,1413,1437]" captionStart="FIGURES 24 – 26" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[166,1429,853,1827]" captionTargetId="figure@12.[151,1436,844,1840]" captionTargetPageId="12" captionText="FIGURES 24 – 26. Dorsal view of holotypes of Hlavaciellus, males. 24, H. bifoveatus sp. n. 25, H. metrios sp. n. 26, H. miser sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276827/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
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) strongly convex, pigmentation light brown, vestiture yellowish.
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</paragraph>
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<paragraph id="8BD036F2FFD4FF9EFF4BEE95156CF887" blockId="11.[151,1436,1231,2014]" pageId="11" pageNumber="12">
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Head (
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<figureCitation id="13542A77FFD4FF9EFE9DEE95114AFA2F" box="[273,340,1485,1509]" captionStart="FIGURES 4 – 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
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) very large, HL 0.35–0.38 (0.36), HW 0.53–0.55 (0.54); vertex long and convex, its anterior margin broadly subtriangular and delimited from supraantennal tubercles and frons by pair of large, shallow and subtriangular impressions broadly separated in middle; frons and clypeus confluent, distinctly convex; supraantennal tubercles prominent, indistinctly separated from median part of frons; eyes large, strongly convex and coarsely faceted. Punctures and setae as in
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<figureCitation id="13542A77FFD4FF9EFE79ED051224F9BF" box="[501,570,1629,1653]" captionStart="FIGURES 4 – 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
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. Antennae very long and slender, as long as 0.78 BL, gradually thickened up to antennomere XI, uniformly covered with thin, long, dense and suberect setae, AnL 1.60–1.68 (1.64), antennomere I twice as long as broad; II much narrower and shorter than I, 1.3x as long as broad; III slightly broader and much longer than II, 1.8x as long as broad; IV slightly larger than III, 1.8x as long as broad; V distinctly broader but only minimally longer than IV, 1.7x as long as broad; VI distinctly longer and broader than V, 1.6x as long as broad; VII–IX nearly equal in length and width, each distinctly larger than VI and about 1.6x as long as broad; X slightly larger than IX, 1.6x as long as broad; XI slightly broader than X, distinctly shorter than IX–X together.
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</paragraph>
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<paragraph id="8BD036F2FFD4FF99FF4BEC0214E6FF3D" blockId="11.[151,1436,1231,2014]" lastBlockId="12.[151,1437,151,788]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Pronotum in dorsal view subrectangular with weakly convex anterior margin, nearly equally broad from base up to anterior fourth, PL 0.63–0.68 (0.65), PW 0.85–0.88 (0.86); sides strongly rounded in anterior fifth or fourth, then nearly straight and parallel with barely discernible constriction near posterior third; hind angles slightly sharp but broadly rounded; posterior margin deeply bi-emarginate; ante-basal pits very shallow and diffused, each equally distant from posterior and lateral margin of pronotum; lateral edges not swollen. Pronotal disc glossy, in middle covered with very fine but distinct punctures separated by spaces 1–1.5x as wide as puncture diameters, punctures close to margins of pronotum are smaller. Setae long, moderately dense, suberect to erect.</paragraph>
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<paragraph id="8BD036F2FFD3FF99FF4BEA5C122CFE3E" blockId="12.[151,1437,151,788]" pageId="12" pageNumber="13">
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Elytra oval, relatively long, broadest near anterior third, EL 1.08–1.13 (1.10), EW 0.93, EI 1.16–1.22; basal pit on each elytron small but distinct, located close to middle of elytral base within adsutural half of elytron; subhumeral lines very distinct, as long as 0.3x EL, very slightly divergent, each developed as sharply marked border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron only slightly more distinct than those on pronotum, nearly equally dense but slightly larger, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum.
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<collectingCountry id="F3787662FFD3FF99FF1BEA8410D1FE3E" box="[151,207,476,500]" name="India" pageId="12" pageNumber="13">Hind</collectingCountry>
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wings entirely reduced, absent.
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</paragraph>
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<paragraph id="8BD036F2FFD3FF99FF4BE958132CFDDD" blockId="12.[151,1437,151,788]" box="[199,818,511,536]" pageId="12" pageNumber="13">Legs slender and long, tibiae straight or nearly straight.</paragraph>
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<paragraph id="8BD036F2FFD3FF99FF4BE97C14F3FDF6" blockId="12.[151,1437,151,788]" box="[199,1261,548,573]" pageId="12" pageNumber="13">Metaventrite with very large and deep postmesocoxal impressions, occupying 2/3 of its length.</paragraph>
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<paragraph id="8BD036F2FFD3FF99FF4BE91F1543FD4E" blockId="12.[151,1437,151,788]" pageId="12" pageNumber="13">Aedeagus (Figs. 31, 32) very slender, AeL 0.43; median lobe with subtriangular, rounded apical part; internal armature asymmetrical; parameres in lateral view moderately broad, with moderately strongly curved apices.</paragraph>
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<paragraph id="8BD036F2FFD3FF99FF4BE9C811BCFD6D" blockId="12.[151,1437,151,788]" box="[199,418,655,680]" pageId="12" pageNumber="13">
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||
<emphasis id="B91BEAE0FFD3FF99FF4BE9C8113BFD62" bold="true" box="[199,293,656,680]" pageId="12" pageNumber="13">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD3FF99FF4BE9EB12DEFD06" box="[199,704,691,716]" pageId="12" pageNumber="13" type="distribution">
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||
<paragraph id="8BD036F2FFD3FF99FF4BE9EB12DEFD06" blockId="12.[151,1437,151,788]" box="[199,704,691,716]" pageId="12" pageNumber="13">
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<emphasis id="B91BEAE0FFD3FF99FF4BE9EB117EFD06" bold="true" box="[199,352,691,716]" pageId="12" pageNumber="13">Distribution.</emphasis>
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East
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<collectingCountry id="F3787662FFD3FF99FE12E9EC1217FD06" box="[414,521,692,716]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
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: Borneo, Sabah.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD3FF99FF4BE98013A3FD25" box="[199,957,727,752]" pageId="12" pageNumber="13" type="etymology">
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<paragraph id="8BD036F2FFD3FF99FF4BE98013A3FD25" blockId="12.[151,1437,151,788]" box="[199,957,727,752]" pageId="12" pageNumber="13">
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<emphasis id="B91BEAE0FFD3FF99FF4BE9801150FD3A" bold="true" box="[199,334,728,752]" pageId="12" pageNumber="13">Etymology.</emphasis>
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The name refers to the pair of impressions on the head.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD3FF99FF4BE9A41296FCDE" box="[199,648,764,788]" pageId="12" pageNumber="13" type="discussion">
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||
<paragraph id="8BD036F2FFD3FF99FF4BE9A41296FCDE" blockId="12.[151,1437,151,788]" box="[199,648,764,788]" pageId="12" pageNumber="13">
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<emphasis id="B91BEAE0FFD3FF99FF4BE9A41125FCDE" bold="true" box="[199,315,764,788]" pageId="12" pageNumber="13">Remarks.</emphasis>
|
||
See remarks for
|
||
<taxonomicName id="4C6F4D71FFD3FF99FE76E9A5129FFCDE" box="[506,641,765,788]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="B91BEAE0FFD3FF99FE76E9A5129FFCDE" box="[506,641,765,788]" italics="true" pageId="12" pageNumber="13">H. smetanai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |