445 lines
61 KiB
XML
445 lines
61 KiB
XML
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<mods:title id="9EEE6C2483B309C097B71E0879C45AC6">Systematic revision of the genus Phaeaphodius Reitter, 1892 (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:namePart id="49D632A328416E928C7901E76FEE475F">Dellacasa, Marco</mods:namePart>
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<mods:title id="DD7C32A0E0E886A6F84D6FAFA5CFF067">Zootaxa</mods:title>
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<paragraph id="776F3686C2710F7CFF31FB49F32115FB" blockId="1.[151,576,1151,1178]" box="[151,576,1151,1178]" pageId="1" pageNumber="144">
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<heading id="2C2781EAC2710F7CFF31FB49F32115FB" bold="true" box="[151,576,1151,1178]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
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<emphasis id="45A4EA94C2710F7CFF31FB49F32115FB" bold="true" box="[151,576,1151,1178]" pageId="1" pageNumber="144">
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Genus
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<taxonomicName id="B0D04D05C2710F7CFF48FBB6F32115FB" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[238,576,1151,1178]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
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<emphasis id="45A4EA94C2710F7CFF48FBB6F0FA15F8" bold="true" box="[238,411,1151,1177]" italics="true" pageId="1" pageNumber="144">Phaeaphodius</emphasis>
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<bibRefCitation id="13414B77C2710F7CFE04FB49F32115FB" author="Reitter" box="[418,576,1152,1178]" pageId="1" pageNumber="144" refString="Reitter, E. (1892) Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brunn, 30, 140 - 262." type="journal article" year="1892">Reitter, 1892</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="3FCA650DC2710F7CFF31FB08F39D1475" pageId="1" pageNumber="144" type="reference_group">
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<paragraph id="776F3686C2710F7CFF31FB08F41715B6" blockId="1.[151,1398,1217,1301]" box="[151,1398,1217,1239]" pageId="1" pageNumber="144">
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<treatmentCitationGroup id="57C011A8C2710F7CFF31FB08F41715B6" box="[151,1398,1217,1239]" pageId="1" pageNumber="144">
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<taxonomicName id="B0D04D05C2710F7CFF31FB08F33F15B6" authority="Reitter, 1892: 205" authorityName="Reitter" authorityPageNumber="205" authorityYear="1892" box="[151,606,1217,1239]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="subGenus" subGenus="Phaeaphodius">
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<emphasis id="45A4EA94C2710F7CFF31FB08F19615B6" box="[151,247,1217,1239]" italics="true" pageId="1" pageNumber="144">Aphodius</emphasis>
|
||
(
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||
<emphasis id="45A4EA94C2710F7CFEA3FB08F0F715B6" box="[261,406,1217,1239]" italics="true" pageId="1" pageNumber="144">Phaeaphodius</emphasis>
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)
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||
<treatmentCitation id="F6711097C2710F7CFE03FB08F33F15B6" author="Reitter" box="[421,606,1217,1239]" page="205" pageId="1" pageNumber="144" year="1892">
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<bibRefCitation id="13414B77C2710F7CFE03FB08F34A15B6" author="Reitter" box="[421,555,1217,1239]" pageId="1" pageNumber="144" refString="Reitter, E. (1892) Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brunn, 30, 140 - 262." type="journal article" year="1892">Reitter, 1892</bibRefCitation>
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||
: 205
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||
</treatmentCitation>
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||
</taxonomicName>
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||
;
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<treatmentCitation id="F6711097C2710F7CFDCFFB08F25615B6" author="Balthasar" box="[617,823,1217,1239]" page="281" pageId="1" pageNumber="144" year="1964">
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<bibRefCitation id="13414B77C2710F7CFDCFFB08F26415B6" author="Balthasar" box="[617,773,1217,1239]" pageId="1" pageNumber="144" refString="Balthasar, V. (1964) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 3. Aphodiidae. Verlag der Tschechoslowakische Akademie der Wissenschaften, Prague, 652 pp." type="book" year="1964">Balthasar 1964</bibRefCitation>
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||
: 281
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="F6711097C2710F7CFCE4FB08F57515B6" author="Dellacasa" box="[834,1044,1217,1239]" page="588" pageId="1" pageNumber="144" year="1993">
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<bibRefCitation id="13414B77C2710F7CFCE4FB08F28015B6" author="Dellacasa" box="[834,993,1217,1239]" pageId="1" pageNumber="144" refString="Dellacasa, G. (1993) Systematic redefinition of Aphodius subgenera Aphodiellus A. Schmidt, 1913, Lunaphodius Balthasar, 1964, Phaeaphodius Reitter, 1892 and their type species. Memorie della Societa entomologica italiana, 71 (1992), 581 - 592." type="journal article" year="1993">Dellacasa 1993</bibRefCitation>
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||
: 588
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="F6711097C2710F7CFBB8FB08F41315B6" author="Dellacasa" box="[1054,1394,1217,1239]" page="133" pageId="1" pageNumber="144" year="2006">
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<bibRefCitation id="13414B77C2710F7CFBB8FB08F42115B6" author="Dellacasa" box="[1054,1344,1217,1239]" pageId="1" pageNumber="144" refString="Dellacasa, M. & Dellacasa, G. (2006) Scarabaeidae: Aphodiinae: Aphodiini (Aphodiina). In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 105 - 142." type="book" year="2006">Dellacasa & Dellacasa 2006</bibRefCitation>
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: 133
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="776F3686C2710F7CFF31FB29F58A1597" blockId="1.[151,1398,1217,1301]" box="[151,1259,1248,1270]" pageId="1" pageNumber="144">
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<treatmentCitationGroup id="57C011A8C2710F7CFF31FB29F58A1597" box="[151,1259,1248,1270]" pageId="1" pageNumber="144">
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<taxonomicName id="B0D04D05C2710F7CFF31FB29F33A1597" authority="Balthasar, 1932: 109" authorityName="Balthasar" authorityPageNumber="109" authorityYear="1932" box="[151,603,1248,1270]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="subGenus" subGenus="Limaroides">
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<emphasis id="45A4EA94C2710F7CFF31FB29F1961597" box="[151,247,1248,1270]" italics="true" pageId="1" pageNumber="144">Aphodius</emphasis>
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||
(
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<emphasis id="45A4EA94C2710F7CFEA3FB29F0161597" box="[261,375,1248,1270]" italics="true" pageId="1" pageNumber="144">Limaroides</emphasis>
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||
)
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||
<treatmentCitation id="F6711097C2710F7CFE20FB29F33A1597" author="Balthasar" box="[390,603,1248,1270]" page="109" pageId="1" pageNumber="144" year="1932">
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<bibRefCitation id="13414B77C2710F7CFE20FB29F3491597" author="Balthasar" box="[390,552,1248,1270]" pageId="1" pageNumber="144" refString="Balthasar, V. (1932) Zehn neue Arten und Formen der Gattung Aphodius Ill. (18. Beitrag zur Kenntnis der Scarabaeidae des palaarktischen Faunen-Gebietes). Wiener entomologische Zeitung, 49, 109 - 125." type="journal article" year="1932">Balthasar, 1932</bibRefCitation>
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||
: 109
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||
</treatmentCitation>
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||
</taxonomicName>
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||
;
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||
<treatmentCitation id="F6711097C2710F7CFDC3FB29F2C21597" author="Stebnicka" box="[613,931,1248,1270]" page="725" pageId="1" pageNumber="144" year="1991">
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<bibRefCitation id="13414B77C2710F7CFDC3FB29F2151597" author="Stebnicka" box="[613,884,1248,1270]" pageId="1" pageNumber="144" refString="Stebnicka, Z. & Galante, E. (1991) Studies on some Aphodiinae (Coleoptera, Scarabaeidae) from the Museum d'Histoire naturelle in Geneve. Revue Suisse de Zoologie, 98, 725 - 729. http: // dx. doi. org / 10.5962 / bhl. part. 79809" type="journal article" year="1991">Stebnicka & Galante 1991</bibRefCitation>
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||
: 725
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</treatmentCitation>
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(as synonym of
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||
<taxonomicName id="B0D04D05C2710F7CFBEAFB29F5BC1597" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[1100,1245,1248,1270]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
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||
<emphasis id="45A4EA94C2710F7CFBEAFB29F5BC1597" box="[1100,1245,1248,1270]" italics="true" pageId="1" pageNumber="144">Phaeaphodius</emphasis>
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</taxonomicName>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="776F3686C2710F7CFF31FB37F39D1475" blockId="1.[151,1398,1217,1301]" box="[151,764,1278,1301]" pageId="1" pageNumber="144">
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||
<treatmentCitationGroup id="57C011A8C2710F7CFF31FB37F39D1475" box="[151,764,1278,1301]" pageId="1" pageNumber="144">
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<taxonomicName id="B0D04D05C2710F7CFF31FB37F0821475" authority="Paulian 1942: 85" authorityName="Paulian" authorityPageNumber="85" authorityYear="1942" box="[151,483,1278,1300]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
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||
<emphasis id="45A4EA94C2710F7CFF31FB37F0491475" box="[151,296,1278,1300]" italics="true" pageId="1" pageNumber="144">Phaeaphodius</emphasis>
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||
:
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<treatmentCitation id="F6711097C2710F7CFE93FB37F0821475" author="Paulian" box="[309,483,1278,1300]" page="85" pageId="1" pageNumber="144" year="1942">
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<bibRefCitation id="13414B77C2710F7CFE93FB37F0DC1475" author="Paulian" box="[309,445,1278,1300]" pageId="1" pageNumber="144" refString="Paulian, R. (1942) Coleopteres Scarabaeidae: Aphodiinae. Exploration du Parc National Albert. Mission G. F. de Witte (1933 - 1935). Institut des Parcs Nationaux du Congo Belge, Bruxelles, 35, 1 - 143." type="journal article" year="1942">Paulian 1942</bibRefCitation>
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||
: 85
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||
</treatmentCitation>
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||
</taxonomicName>
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||
;
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||
<treatmentCitation id="F6711097C2710F7CFE48FB37F3981475" author="Dellacasa" box="[494,761,1278,1301]" page="238" pageId="1" pageNumber="144" year="2001">
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<bibRefCitation id="13414B77C2710F7CFE48FB37F3A71475" author="Dellacasa" box="[494,710,1278,1301]" pageId="1" pageNumber="144" refString="Dellacasa, G., Bordat, P. & Dellacasa, M. (2001) A revisional essay of world genus-group taxa of Aphodiinae. Memorie della Societa entomologica italiana, 79 (2000), 1 - 482." type="journal article" year="2001">
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||
Dellacasa
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||
<emphasis id="45A4EA94C2710F7CFDF0FAC9F3EB1475" box="[598,650,1278,1301]" italics="true" pageId="1" pageNumber="144">et al.</emphasis>
|
||
2001
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||
</bibRefCitation>
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||
: 238
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="3FCA650DC2710F7CFF31FAF3F3051417" pageId="1" pageNumber="144" type="materials_examined">
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<paragraph id="776F3686C2710F7CFF31FAF3F3051417" blockId="1.[151,1437,1338,2032]" pageId="1" pageNumber="144">
|
||
<emphasis id="45A4EA94C2710F7CFF31FAF3F0551432" bold="true" box="[151,308,1338,1363]" pageId="1" pageNumber="144">Type species</emphasis>
|
||
.
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||
<taxonomicName id="B0D04D05C2710F7CFEEAFAF2F08D1432" box="[332,492,1339,1363]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="45A4EA94C2710F7CFEEAFAF2F08D1432" box="[332,492,1339,1363]" italics="true" pageId="1" pageNumber="144">Phaeaphodius</emphasis>
|
||
</taxonomicName>
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||
:
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||
<taxonomicName id="B0D04D05C2710F7CFDA2FAF2F2191432" authority="Harold, 1871" authorityName="Harold" authorityYear="1871" box="[516,888,1338,1363]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="solskyi">
|
||
<emphasis id="45A4EA94C2710F7CFDA2FAF2F3AB1432" box="[516,714,1339,1363]" italics="true" pageId="1" pageNumber="144">Aphodius solskyi</emphasis>
|
||
<bibRefCitation id="13414B77C2710F7CFD7DFAF3F2191432" author="Harold" box="[731,888,1338,1363]" pageId="1" pageNumber="144" refString="Harold, E. (1871) Beitrage zur Kenntniss einiger coprophagen Lamellicornien. (Siebentes Stuck). Berliner entomologische Zeitschrift, 15, 249 - 287. http: // dx. doi. org / 10.1002 / mmnd. 18710150229" type="journal article" year="1871">Harold, 1871</bibRefCitation>
|
||
</taxonomicName>
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||
(original designation).
|
||
<taxonomicName id="B0D04D05C2710F7CFB3DFAF2F4781432" box="[1179,1305,1339,1363]" class="Insecta" family="Aphodiidae" genus="Limaroides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="45A4EA94C2710F7CFB3DFAF2F4781432" box="[1179,1305,1339,1363]" italics="true" pageId="1" pageNumber="144">Limaroides</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="B0D04D05C2710F7CFA94FAF2F0B91417" authority="Balthasar, 1932" authorityName="Balthasar" authorityYear="1932" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="kiulungensis">
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||
<emphasis id="45A4EA94C2710F7CFA94FAF2F0421417" italics="true" pageId="1" pageNumber="144">Aphodius kiulungensis</emphasis>
|
||
<bibRefCitation id="13414B77C2710F7CFE8CFA94F0B91417" author="Balthasar" box="[298,472,1373,1398]" pageId="1" pageNumber="144" refString="Balthasar, V. (1932) Zehn neue Arten und Formen der Gattung Aphodius Ill. (18. Beitrag zur Kenntnis der Scarabaeidae des palaarktischen Faunen-Gebietes). Wiener entomologische Zeitung, 49, 109 - 125." type="journal article" year="1932">Balthasar, 1932</bibRefCitation>
|
||
</taxonomicName>
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||
(monotypy)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3FCA650DC2710F7CFF61FA48F0E316AC" pageId="1" pageNumber="144" type="diagnosis">
|
||
<paragraph id="776F3686C2710F7CFF61FA48F0E316AC" blockId="1.[151,1437,1338,2032]" pageId="1" pageNumber="144">
|
||
<emphasis id="45A4EA94C2710F7CFF61FA48F05B14FB" bold="true" box="[199,314,1409,1434]" pageId="1" pageNumber="144">Diagnosis</emphasis>
|
||
. Small or medium size species (length 4.0–7.0 mm), moderately convex, shiny, glabrous. Color black or brown; sometimes elytra yellow with dark spots, rarely testaceous; antennae and legs dark, latter with paler tarsi. Head with epistome gibbous, transversely subcarinate, coarsely subrugosely punctate; clypeus sinuate at middle, generally round at sides, thickly bordered, edge glabrous; genae obtusely round, densely setose, protruding from the eyes; frontal suture usually tuberculate. Pronotum transverse, moderately convex, punctate; sides feebly round; hind angles distinctly, sometimes inward-sinuately truncate; base more or less completely bordered. Scutellum small, triangularly elongate or pentagonal with subparallel sides toward base. Elytra elongate, faintly widened posteriorly; striae distinctly punctate; interstriae flat or moderately convex. Protibiae distally tridentate and proximally serrulate on outer margin; upper side very finely and sparsely punctate. Mesotibiae and metatibiae with strong, transverse carinae on outer face; apically fimbriate with elongate and irregularly unequal, exceptionally short and progressively unequal spinules. Pygidium with dense, elongate, recumbent pubescence; apical margin finely, densely setose. Sexual dimorphism in males: frontal suture with prominent tubercles; pronotum not narrowed apically; protibiae spur and mesotibiae inferior spur hooked apically. Aedeagus slender, with elongate parameres. Epipharynx feebly sinuate at anterior margin and widely round at sides; epitorma broadly conical; corypha with protruding slender apical celtes; chaetopedia entirely densely pubescent and with several subregular belts of spines, much shorter, stout, densely arranged near the base of epitorma; chaetopariae dense and moderately elongate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3FCA650DC2710F7CFF61F81EF3441691" box="[199,549,2007,2032]" pageId="1" pageNumber="144" type="distribution">
|
||
<paragraph id="776F3686C2710F7CFF61F81EF3441691" blockId="1.[151,1437,1338,2032]" box="[199,549,2007,2032]" pageId="1" pageNumber="144">
|
||
<emphasis id="45A4EA94C2710F7CFF61F81EF0381691" bold="true" box="[199,345,2007,2032]" pageId="1" pageNumber="144">Distribution</emphasis>
|
||
. Holarctic region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3FCA650DC2720F7FFF31FF51F37811D0" box="[151,537,151,178]" pageId="2" pageNumber="145" type="discussion">
|
||
<paragraph id="776F3686C2720F7FFF31FF51F37811D0" blockId="2.[151,537,151,178]" box="[151,537,151,178]" pageId="2" pageNumber="145">
|
||
<heading id="2C2781EAC2720F7FFF31FF51F37811D0" bold="true" box="[151,537,151,178]" fontSize="11" level="1" pageId="2" pageNumber="145" reason="1">
|
||
<emphasis id="45A4EA94C2720F7FFF31FF51F37811D0" bold="true" box="[151,537,151,178]" pageId="2" pageNumber="145">
|
||
Key to species of
|
||
<taxonomicName id="B0D04D05C2720F7FFECBFF5EF37811D0" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[365,537,151,177]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="45A4EA94C2720F7FFECBFF5EF37811D0" bold="true" box="[365,537,151,177]" italics="true" pageId="2" pageNumber="145">Phaeaphodius</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3FCA650DC2720F7FFF31FF17F4FD142F" pageId="2" pageNumber="145" type="key">
|
||
<keyStep id="CC242E23C2720F7FFF31FF17F4FA104F" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FF17F4FD1071" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FF17F4FD1071" pageId="2" pageNumber="145">
|
||
1. Clypeus denticulate at sides of median sinuosity (
|
||
<figureCitation id="EFEB2A03C2720F7FFD69FF17F27B1192" box="[719,794,222,243]" captionStart="FIGURES 44 – 62" captionStartId="16.[151,264,1818,1841]" captionTargetBox="[164,1429,200,1785]" captionTargetId="figure@16.[152,1435,193,1797]" captionTargetPageId="16" captionText="FIGURES 44 – 62. Phaeaphodius rectus (Motschulsky, 1866) (Russia, Eastern Siberia, Irkutsk environs). 44, epipharynx; 45 – 46, aedeagus (dorsal and lateral views); 47 – 57, habitus (morphological details, scale bar, and color variations). Phaeaphodius roddi (Koshantschikov, 1911) (Barnaul, lectotype). 58, epipharynx; 59 – 60, aedeagus (dorsal and lateral views); 61 – 62, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254697/files/figure.png" pageId="2" pageNumber="145">Fig. 62</figureCitation>
|
||
). Shiny black. Length 5.0–6.0 mm.
|
||
<collectingCountry id="0FC77616C2720F7FFB24FF17F5951192" box="[1154,1268,222,243]" name="Kazakhstan" pageId="2" pageNumber="145">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="0FC77616C2720F7FFAA6FF17F4211192" box="[1280,1344,222,243]" name="Russia" pageId="2" pageNumber="145">Russia</collectingCountry>
|
||
(western Siberia)....................................................................
|
||
<emphasis id="45A4EA94C2720F7FFBC1FF32F5CE1071" box="[1127,1199,251,272]" italics="true" pageId="2" pageNumber="145">P. ro d d i</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFB18FF32F4F51071" author="Koshantschikov" box="[1214,1428,251,272]" pageId="2" pageNumber="145" refString="Koshantschikov, W. (1911) Zweiter Beitrag zur Kenntnis der Aphodiini. Russkoe Entomologicheskoe Obozreniye, 11, 18 - 25." type="journal article" year="1911">Koshantschikov, 1911</bibRefCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FED0F4FA104F" blockId="2.[151,1437,222,1358]" box="[151,1435,281,302]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FED0F4FA104F" box="[151,1435,281,302]" pageId="2" pageNumber="145">
|
||
- Clypeus round at sides of median sinuosity (
|
||
<figureCitation id="EFEB2A03C2720F7FFD2DFED0F386104F" box="[651,743,281,302]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1908,1931]" captionTargetBox="[214,1360,208,1863]" captionTargetId="figure@3.[201,1372,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Phaeaphodius costalis (Gebler, 1848) (Kazakhstan, Kurgaldzhino, Lake Sultan). 1, epipharynx; 2 – 3. habitus (scale bar and morphological details); 4 – 5. aedeagus (dorsal and lateral views). Phaeaphodius dauricus (Harold, 1863) (Turkmenia mer., Tedžen). 6, epipharynx; 7 – 10, habitus (morphological details, scale bar, and color variations); 11 – 12, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254693/files/figure.png" pageId="2" pageNumber="145">Figs. 3, 7</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD56FED0F24E104F" box="[752,815,281,302]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">15, 19</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFC9EFED0F217104F" box="[824,886,281,302]" captionStart="FIGURES 24 – 33" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[200,1387,209,1899]" captionTargetId="figure@9.[185,1401,193,1911]" captionTargetPageId="9" captionText="FIGURES 24 – 33. Phaeaphodius kiulungensis (Balthasar, 1932) (China, Sichuan, Baoxing, 100 km N of Yaan). 24, epipharynx; 25 – 26. habitus (scale bar and morphological details); 27 – 28. aedeagus (dorsal and lateral views). Phaeaphodius novikovi (Kabakov, 1998) (Ukraine, Donetskaya Oblasty, Telmanovskij, Samsonovo z. - k., “ Khomutovskaya Step ”, paratype). 29, epipharynx; 30 – 31, habitus (morphological details and scale bar); 32 – 33, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254695/files/figure.png" pageId="2" pageNumber="145">26, 30</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFC26FED0F2DF104F" box="[896,958,281,302]" captionStart="FIGURES 34 – 43" captionStartId="11.[151,264,1911,1934]" captionTargetBox="[169,1424,206,1876]" captionTargetId="figure@11.[150,1436,193,1890]" captionTargetPageId="11" captionText="FIGURES 34 – 43. Phaeaphodius plutenkoi (Král, 2002) (China, Shaanxi, Taibashan Mountains, N Hoazhenzi, 2460 m). 34, epipharynx; 35 – 36, habitus (scale bar and morphological details); 37 – 38, aedeagus (dorsal and lateral views). Phaeaphodius pupillus (Balthasar, 1961) (Afghanistan, Kandahar-Kuna). 39. epipharynx; 40 – 41, habitus (morphological details and scale bar); 42 – 43, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254696/files/figure.png" pageId="2" pageNumber="145">36, 40</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFC6EFED0F283104F" box="[968,994,281,302]" captionStart="FIGURES 44 – 62" captionStartId="16.[151,264,1818,1841]" captionTargetBox="[164,1429,200,1785]" captionTargetId="figure@16.[152,1435,193,1797]" captionTargetPageId="16" captionText="FIGURES 44 – 62. Phaeaphodius rectus (Motschulsky, 1866) (Russia, Eastern Siberia, Irkutsk environs). 44, epipharynx; 45 – 46, aedeagus (dorsal and lateral views); 47 – 57, habitus (morphological details, scale bar, and color variations). Phaeaphodius roddi (Koshantschikov, 1911) (Barnaul, lectotype). 58, epipharynx; 59 – 60, aedeagus (dorsal and lateral views); 61 – 62, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254697/files/figure.png" pageId="2" pageNumber="145">47</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFC4AFED0F564104F" box="[1004,1029,281,302]" captionStart="FIGURES 63 – 67" captionStartId="17.[151,264,1844,1867]" captionTargetBox="[159,1431,1320,1819]" captionTargetId="figure@17.[150,1433,1308,1827]" captionTargetPageId="17" captionText="FIGURES 63 – 67. Phaeaphodius tauricola (Hrubant, 1961) (Turkey, Vilayet Erzurum, Kireçli Geçidi, 2400 m). 63, epipharynx; 64 – 65, aedeagus (dorsal and lateral views); 66 – 67, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254698/files/figure.png" pageId="2" pageNumber="145">67</figureCitation>
|
||
)............................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FEFFF4FA1009" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FEFFF4FA102A" blockId="2.[151,1437,222,1358]" box="[151,1435,310,331]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FEFFF4FA102A" box="[151,1435,310,331]" pageId="2" pageNumber="145">
|
||
2 (1). Elytral declivity with short pubescence (
|
||
<figureCitation id="EFEB2A03C2720F7FFDC2FEFFF3D2102A" box="[612,691,310,331]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD18FEFFF3B6102A" box="[702,727,310,331]" captionStart="FIGURES 34 – 43" captionStartId="11.[151,264,1911,1934]" captionTargetBox="[169,1424,206,1876]" captionTargetId="figure@11.[150,1436,193,1890]" captionTargetPageId="11" captionText="FIGURES 34 – 43. Phaeaphodius plutenkoi (Král, 2002) (China, Shaanxi, Taibashan Mountains, N Hoazhenzi, 2460 m). 34, epipharynx; 35 – 36, habitus (scale bar and morphological details); 37 – 38, aedeagus (dorsal and lateral views). Phaeaphodius pupillus (Balthasar, 1961) (Afghanistan, Kandahar-Kuna). 39. epipharynx; 40 – 41, habitus (morphological details and scale bar); 42 – 43, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254696/files/figure.png" pageId="2" pageNumber="145">36</figureCitation>
|
||
)........................................................ 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FE9AF4FA1009" blockId="2.[151,1437,222,1358]" box="[151,1435,339,360]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FE9AF4FA1009" box="[151,1435,339,360]" pageId="2" pageNumber="145">
|
||
- Elytral declivity glabrous (
|
||
<figureCitation id="EFEB2A03C2720F7FFE44FE9AF35F1009" box="[482,574,339,360]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1908,1931]" captionTargetBox="[214,1360,208,1863]" captionTargetId="figure@3.[201,1372,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Phaeaphodius costalis (Gebler, 1848) (Kazakhstan, Kurgaldzhino, Lake Sultan). 1, epipharynx; 2 – 3. habitus (scale bar and morphological details); 4 – 5. aedeagus (dorsal and lateral views). Phaeaphodius dauricus (Harold, 1863) (Turkmenia mer., Tedžen). 6, epipharynx; 7 – 10, habitus (morphological details, scale bar, and color variations); 11 – 12, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254693/files/figure.png" pageId="2" pageNumber="145">Figs. 3, 7</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFDEEFE9AF3031009" box="[584,610,339,360]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">19</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFDCDFE9AF3CB1009" box="[619,682,339,360]" captionStart="FIGURES 24 – 33" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[200,1387,209,1899]" captionTargetId="figure@9.[185,1401,193,1911]" captionTargetPageId="9" captionText="FIGURES 24 – 33. Phaeaphodius kiulungensis (Balthasar, 1932) (China, Sichuan, Baoxing, 100 km N of Yaan). 24, epipharynx; 25 – 26. habitus (scale bar and morphological details); 27 – 28. aedeagus (dorsal and lateral views). Phaeaphodius novikovi (Kabakov, 1998) (Ukraine, Donetskaya Oblasty, Telmanovskij, Samsonovo z. - k., “ Khomutovskaya Step ”, paratype). 29, epipharynx; 30 – 31, habitus (morphological details and scale bar); 32 – 33, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254695/files/figure.png" pageId="2" pageNumber="145">26, 30</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD15FE9AF3AC1009" box="[691,717,339,360]" captionStart="FIGURES 34 – 43" captionStartId="11.[151,264,1911,1934]" captionTargetBox="[169,1424,206,1876]" captionTargetId="figure@11.[150,1436,193,1890]" captionTargetPageId="11" captionText="FIGURES 34 – 43. Phaeaphodius plutenkoi (Král, 2002) (China, Shaanxi, Taibashan Mountains, N Hoazhenzi, 2460 m). 34, epipharynx; 35 – 36, habitus (scale bar and morphological details); 37 – 38, aedeagus (dorsal and lateral views). Phaeaphodius pupillus (Balthasar, 1961) (Afghanistan, Kandahar-Kuna). 39. epipharynx; 40 – 41, habitus (morphological details and scale bar); 42 – 43, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254696/files/figure.png" pageId="2" pageNumber="145">40</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD71FE9AF3901009" box="[727,753,339,360]" captionStart="FIGURES 44 – 62" captionStartId="16.[151,264,1818,1841]" captionTargetBox="[164,1429,200,1785]" captionTargetId="figure@16.[152,1435,193,1797]" captionTargetPageId="16" captionText="FIGURES 44 – 62. Phaeaphodius rectus (Motschulsky, 1866) (Russia, Eastern Siberia, Irkutsk environs). 44, epipharynx; 45 – 46, aedeagus (dorsal and lateral views); 47 – 57, habitus (morphological details, scale bar, and color variations). Phaeaphodius roddi (Koshantschikov, 1911) (Barnaul, lectotype). 58, epipharynx; 59 – 60, aedeagus (dorsal and lateral views); 61 – 62, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254697/files/figure.png" pageId="2" pageNumber="145">47</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD5DFE9AF2741009" box="[763,789,339,360]" captionStart="FIGURES 63 – 67" captionStartId="17.[151,264,1844,1867]" captionTargetBox="[159,1431,1320,1819]" captionTargetId="figure@17.[150,1433,1308,1827]" captionTargetPageId="17" captionText="FIGURES 63 – 67. Phaeaphodius tauricola (Hrubant, 1961) (Turkey, Vilayet Erzurum, Kireçli Geçidi, 2400 m). 63, epipharynx; 64 – 65, aedeagus (dorsal and lateral views); 66 – 67, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254698/files/figure.png" pageId="2" pageNumber="145">67</figureCitation>
|
||
)................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FEB8F4FA10BF" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FEB8F4FA10C2" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FEB8F4FA10C2" pageId="2" pageNumber="145">
|
||
3 (2). Length 6.0–7.0 mm; pronotum, in males, with small median foveola at the anterior margin (
|
||
<figureCitation id="EFEB2A03C2720F7FFBE9FEB8F5F610E7" box="[1103,1175,369,390]" captionStart="FIGURES 34 – 43" captionStartId="11.[151,264,1911,1934]" captionTargetBox="[169,1424,206,1876]" captionTargetId="figure@11.[150,1436,193,1890]" captionTargetPageId="11" captionText="FIGURES 34 – 43. Phaeaphodius plutenkoi (Král, 2002) (China, Shaanxi, Taibashan Mountains, N Hoazhenzi, 2460 m). 34, epipharynx; 35 – 36, habitus (scale bar and morphological details); 37 – 38, aedeagus (dorsal and lateral views). Phaeaphodius pupillus (Balthasar, 1961) (Afghanistan, Kandahar-Kuna). 39. epipharynx; 40 – 41, habitus (morphological details and scale bar); 42 – 43, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254696/files/figure.png" pageId="2" pageNumber="145">Fig. 36</figureCitation>
|
||
). Dorsal color black, elytra dirty yellow with dark brown, shadowy-margined spots.
|
||
<collectingCountry id="0FC77616C2720F7FFCA6FE47F25810C2" box="[768,825,398,419]" name="China" pageId="2" pageNumber="145">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="B514F864C2720F7FFCE0FE47F2E610C2" box="[838,903,398,419]" country="China" name="Gansu" pageId="2" pageNumber="145">Gansu</collectingRegion>
|
||
,
|
||
<collectingRegion id="B514F864C2720F7FFC36FE47F2BE10C2" box="[912,991,398,419]" country="China" name="Shaanxi" pageId="2" pageNumber="145">Shaanxi</collectingRegion>
|
||
)................
|
||
<taxonomicName id="B0D04D05C2720F7FFB08FE47F4FA10C2" authority="Kral, 2002" authorityName="Kral" authorityYear="2002" box="[1198,1435,398,419]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="plutenkoi">
|
||
<emphasis id="45A4EA94C2720F7FFB08FE47F47E10C2" box="[1198,1311,398,419]" italics="true" pageId="2" pageNumber="145">P. plutenkoi</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFA8AFE47F4F410C2" author="Kral" box="[1324,1429,398,419]" pageId="2" pageNumber="145" refString="Kral, D. (2002) A review of Chinese Aphodius species. Part 6: A. (Phaeaphodius) plutenkoi sp. n. from Shaanxi. Acta Societatis Zoologicae Bohemicae, 66, 47 - 50." type="journal article" year="2002">Král, 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FE62F4FA10BF" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FE62F4FA10BF" pageId="2" pageNumber="145">
|
||
- Length 5.5–6.0 mm; pronotum, in males, without anterior foveola (
|
||
<figureCitation id="EFEB2A03C2720F7FFCC9FE62F2D610A1" box="[879,951,427,448]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">Fig. 15</figureCitation>
|
||
). Dorsal color piceous.
|
||
<collectingCountry id="0FC77616C2720F7FFB06FE62F5B810A1" box="[1184,1241,427,448]" name="Nepal" pageId="2" pageNumber="145">Nepal</collectingCountry>
|
||
(Annapurna Mountains)............................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFC52FE00F4FA10BF" authority="Ahrens & Stebnicka, 1997" authorityName="Ahrens & Stebnicka" authorityYear="1997" box="[1012,1435,457,478]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="himalorectus">
|
||
<emphasis id="45A4EA94C2720F7FFC52FE00F5E610BF" box="[1012,1159,457,478]" italics="true" pageId="2" pageNumber="145">P. himalorectus</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFB32FE00F4F510BF" author="Ahrens" box="[1172,1428,457,478]" pageId="2" pageNumber="145" refString="Ahrens, D. & Stebnicka, Z. (1997) On the Aphodiinae of the Nepal-Himalayas. Stuttgarter Beitrage zur Naturkunde, Serie A, 552, 1 - 17." type="journal article" year="1997">Ahrens & Stebnicka, 1997</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FE2FF4FA1332" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FE2FF4FD1379" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FE2FF4FD1379" pageId="2" pageNumber="145">
|
||
4 (2). Base of pronotum not bordered; metatibiae superior spur shorter than first tarsomere; China (Fujian, Sichuan, Xizang)................................................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFBC7FDCAF4FD1379" authority="Balthasar, 1932" authorityName="Balthasar" authorityYear="1932" box="[1121,1436,515,536]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="kiulungensis">
|
||
<emphasis id="45A4EA94C2720F7FFBC7FDCAF5911379" box="[1121,1264,515,536]" italics="true" pageId="2" pageNumber="145">P. kiulungensis</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFB5BFDCAF4F41379" author="Balthasar" box="[1277,1429,515,536]" pageId="2" pageNumber="145" refString="Balthasar, V. (1932) Zehn neue Arten und Formen der Gattung Aphodius Ill. (18. Beitrag zur Kenntnis der Scarabaeidae des palaarktischen Faunen-Gebietes). Wiener entomologische Zeitung, 49, 109 - 125." type="journal article" year="1932">Balthasar, 1932</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FDE8F4FA1332" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FDE8F4FA1332" pageId="2" pageNumber="145">- Base of pronotum bordered, sometimes the border interrupted on each side before hind angles; metatibiae superior spur as long as first tarsomere or longer............................................................................. 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FD95F4FA13EF" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FD95F4FA1310" blockId="2.[151,1437,222,1358]" box="[151,1435,604,625]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FD95F4FA1310" box="[151,1435,604,625]" pageId="2" pageNumber="145">
|
||
5 (4). Basal border of pronotum interrupted on each side before hind angles (
|
||
<figureCitation id="EFEB2A03C2720F7FFC21FD95F2B61310" box="[903,983,604,625]" captionStart="FIGURES 44 – 62" captionStartId="16.[151,264,1818,1841]" captionTargetBox="[164,1429,200,1785]" captionTargetId="figure@16.[152,1435,193,1797]" captionTargetPageId="16" captionText="FIGURES 44 – 62. Phaeaphodius rectus (Motschulsky, 1866) (Russia, Eastern Siberia, Irkutsk environs). 44, epipharynx; 45 – 46, aedeagus (dorsal and lateral views); 47 – 57, habitus (morphological details, scale bar, and color variations). Phaeaphodius roddi (Koshantschikov, 1911) (Barnaul, lectotype). 58, epipharynx; 59 – 60, aedeagus (dorsal and lateral views); 61 – 62, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254697/files/figure.png" pageId="2" pageNumber="145">Figs. 47</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFC47FD95F29B1310" box="[993,1018,604,625]" captionStart="FIGURES 63 – 67" captionStartId="17.[151,264,1844,1867]" captionTargetBox="[159,1431,1320,1819]" captionTargetId="figure@17.[150,1433,1308,1827]" captionTargetPageId="17" captionText="FIGURES 63 – 67. Phaeaphodius tauricola (Hrubant, 1961) (Turkey, Vilayet Erzurum, Kireçli Geçidi, 2400 m). 63, epipharynx; 64 – 65, aedeagus (dorsal and lateral views); 66 – 67, habitus (scale bar and morphological details)." httpUri="https://zenodo.org/record/254698/files/figure.png" pageId="2" pageNumber="145">67</figureCitation>
|
||
)................................ 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FDB0F4FA13EF" blockId="2.[151,1437,222,1358]" box="[151,1435,633,654]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FDB0F4FA13EF" box="[151,1435,633,654]" pageId="2" pageNumber="145">
|
||
- Basal border of pronotum entire (
|
||
<figureCitation id="EFEB2A03C2720F7FFD87FDB0F31F13EF" box="[545,638,633,654]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1908,1931]" captionTargetBox="[214,1360,208,1863]" captionTargetId="figure@3.[201,1372,195,1876]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Phaeaphodius costalis (Gebler, 1848) (Kazakhstan, Kurgaldzhino, Lake Sultan). 1, epipharynx; 2 – 3. habitus (scale bar and morphological details); 4 – 5. aedeagus (dorsal and lateral views). Phaeaphodius dauricus (Harold, 1863) (Turkmenia mer., Tedžen). 6, epipharynx; 7 – 10, habitus (morphological details, scale bar, and color variations); 11 – 12, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254693/files/figure.png" pageId="2" pageNumber="145">Figs. 3, 7</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD21FDB0F3C013EF" box="[647,673,633,654]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">19</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD0DFDB0F3A413EF" box="[683,709,633,654]" captionStart="FIGURES 24 – 33" captionStartId="9.[151,264,1920,1943]" captionTargetBox="[200,1387,209,1899]" captionTargetId="figure@9.[185,1401,193,1911]" captionTargetPageId="9" captionText="FIGURES 24 – 33. Phaeaphodius kiulungensis (Balthasar, 1932) (China, Sichuan, Baoxing, 100 km N of Yaan). 24, epipharynx; 25 – 26. habitus (scale bar and morphological details); 27 – 28. aedeagus (dorsal and lateral views). Phaeaphodius novikovi (Kabakov, 1998) (Ukraine, Donetskaya Oblasty, Telmanovskij, Samsonovo z. - k., “ Khomutovskaya Step ”, paratype). 29, epipharynx; 30 – 31, habitus (morphological details and scale bar); 32 – 33, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254695/files/figure.png" pageId="2" pageNumber="145">30</figureCitation>
|
||
,
|
||
<figureCitation id="EFEB2A03C2720F7FFD69FDB0F38913EF" box="[719,744,633,654]" captionStart="FIGURES 34 – 43" captionStartId="11.[151,264,1911,1934]" captionTargetBox="[169,1424,206,1876]" captionTargetId="figure@11.[150,1436,193,1890]" captionTargetPageId="11" captionText="FIGURES 34 – 43. Phaeaphodius plutenkoi (Král, 2002) (China, Shaanxi, Taibashan Mountains, N Hoazhenzi, 2460 m). 34, epipharynx; 35 – 36, habitus (scale bar and morphological details); 37 – 38, aedeagus (dorsal and lateral views). Phaeaphodius pupillus (Balthasar, 1961) (Afghanistan, Kandahar-Kuna). 39. epipharynx; 40 – 41, habitus (morphological details and scale bar); 42 – 43, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254696/files/figure.png" pageId="2" pageNumber="145">40</figureCitation>
|
||
)....................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FD5FF4FD123A" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FD5FF4FA13A8" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FD5FF4FA13A8" pageId="2" pageNumber="145">
|
||
6 (5). Basal border of pronotum thin, superficial, and crenulate by a contiguous series of moderately-large punctures; elytral striae subcrenulate. Dorsal color black. Length 4.5–5.0 mm. Armenia, Iran, Turkey................
|
||
<taxonomicName id="B0D04D05C2720F7FFB2AFD7DF4FA13A8" authority="Hrubant, 1961" authorityName="Hrubant" authorityYear="1961" box="[1164,1435,692,713]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="tauricola">
|
||
<emphasis id="45A4EA94C2720F7FFB2AFD7DF59A13A8" box="[1164,1275,692,713]" italics="true" pageId="2" pageNumber="145">P. tauricola</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFAAEFD7DF4F413A8" author="Hrubant" box="[1288,1429,692,713]" pageId="2" pageNumber="145" refString="Hrubant, M. (1961) Eine neue Art der Gattung Aphodius Ill. Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fur Tierkunde in Dresden, 26, 97 - 100." type="journal article" year="1961">Hrubant, 1961</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FD18F4FD123A" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FD18F4FD123A" pageId="2" pageNumber="145">
|
||
- Basal border of pronotum strongly impressed, not crenulate; elytral striae distinctly crenulate. Dorsal color black, elytra yellow or castaneous with dark longitudinal bands more or less widened; rarely entirely dirty yellow or castaneous. Length 5.0–6.0 mm. China, Iran, Japan, Kazakhstan, Kyrgyzstan, Mongolia,
|
||
<collectingCountry id="0FC77616C2720F7FFC9CFCC5F21B1240" box="[826,890,780,801]" name="Russia" pageId="2" pageNumber="145">Russia</collectingCountry>
|
||
(
|
||
<collectingRegion id="B514F864C2720F7FFC2FFCC5F2A21240" box="[905,963,780,801]" country="Cameroon" name="North" pageId="2" pageNumber="145">North</collectingRegion>
|
||
European Territory,
|
||
<location id="720F605DC2720F7FFB36FCC5F4F91240" LSID="urn:lsid:plazi:treatment:FF798790C2710F7FFF31FB49F4FD142F:720F605DC2720F7FFB36FCC5F4F91240" box="[1168,1432,780,801]" country="Russia" name="Central European Territory" pageId="2" pageNumber="145" stateProvince="North">Central European Territory</location>
|
||
,
|
||
<collectingRegion id="B514F864C2720F7FFF47FCE0F06B125F" box="[225,266,809,830]" country="Cameroon" name="East" pageId="2" pageNumber="145">East</collectingRegion>
|
||
Siberia,
|
||
<collectingRegion id="B514F864C2720F7FFEC6FCE0F0EE125F" box="[352,399,809,830]" country="Cameroon" name="West" pageId="2" pageNumber="145">West</collectingRegion>
|
||
Siberia, Far
|
||
<collectingRegion id="B514F864C2720F7FFDADFCE0F359125F" box="[523,568,809,830]" country="Cameroon" name="East" pageId="2" pageNumber="145">East</collectingRegion>
|
||
), South Korea, Taiwan, Ukraine,
|
||
<collectingCountry id="0FC77616C2720F7FFCDEFCE0F50B125F" box="[888,1130,809,830]" name="United States of America" pageId="2" pageNumber="145">United States of America</collectingCountry>
|
||
(
|
||
<collectingRegion id="B514F864C2720F7FFBD1FCE0F5A2125F" box="[1143,1219,809,830]" country="United States of America" name="Oregon" pageId="2" pageNumber="145">Oregon</collectingRegion>
|
||
,
|
||
<collectingRegion id="B514F864C2720F7FFB6AFCE0F421125F" box="[1228,1344,809,830]" country="United States of America" name="Washington" pageId="2" pageNumber="145">Washington</collectingRegion>
|
||
)...................................................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFBD8FC8FF4FD123A" authority="Motschulsky, 1866" authorityName="Motschulsky" authorityYear="1866" box="[1150,1436,838,859]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="45A4EA94C2720F7FFBD8FC8FF5AE123A" box="[1150,1231,838,859]" italics="true" pageId="2" pageNumber="145">P. rectus</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFB7AFC8FF4F4123A" author="Motschulsky" box="[1244,1429,838,859]" pageId="2" pageNumber="145" refString="Motschulsky, V. de (1866) Catalogue des insectes recus du Japon. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 39, 163 - 200." type="journal article" year="1866">Motschulsky, 1866</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FCADF4FA128F" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FCADF4FD12D2" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FCADF4FD12D2" pageId="2" pageNumber="145">
|
||
7 (5). Clypeus distinctly sinuate at middle (
|
||
<figureCitation id="EFEB2A03C2720F7FFDE9FCADF3F91218" box="[591,664,868,889]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="2" pageNumber="145">Fig. 19</figureCitation>
|
||
); genae distinctly protruding from the eyes; protibiae spur, in males, stout and strongly hooked apically. Dorsal surface dull. Length 5.5–6.0 mm. Russia (Siberia, Altai, and Ural Mountains)........................................................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFB25FC57F4FD12D2" authority="Reitter, 1907" authorityName="Reitter" authorityYear="1907" box="[1155,1436,926,947]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="jouravliowi">
|
||
<emphasis id="45A4EA94C2720F7FFB25FC57F46B12D2" box="[1155,1290,926,947]" italics="true" pageId="2" pageNumber="145">P. jouravliowi</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFAB0FC57F4F412D2" author="Reitter" box="[1302,1429,926,947]" pageId="2" pageNumber="145" refString="Reitter, E. (1907) Eine Serie neuer Aphodius - Arten aus der palaarktischen Fauna. Deutsche entomologische Zeitschrift, 1907, 407 - 411." type="journal article" year="1907">Reitter, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FC75F4FA128F" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FC75F4FA128F" pageId="2" pageNumber="145">- Clypeus feebly sinuate at middle; genae feebly protruding from the eyes; protibiae spur, in males, rather slender and apically feebly hooked. Dorsal surface more or less shiny. Length 4.0–6.5 mm........................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FC3FF4FD1527" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FC3FF4FD1548" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FC3FF4FD1548" pageId="2" pageNumber="145">
|
||
8 (7). Epistome dually, irregularly but not rugosely punctate; elytral interstriae almost imperceptibly punctate. Length 5.0–6.5 mm. Russia, Ukraine.................................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFB2AFBDDF4FD1548" authority="Kabakov, 1998" authorityName="Kabakov" authorityYear="1998" box="[1164,1436,1044,1065]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="novikovi">
|
||
<emphasis id="45A4EA94C2720F7FFB2AFBDDF5951548" box="[1164,1268,1044,1065]" italics="true" pageId="2" pageNumber="145">P. novikovi</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFAA7FBDDF4F41548" author="Kabakov" box="[1281,1429,1044,1065]" pageId="2" pageNumber="145" refString="Kabakov, O. N. (1998) Obzor podroda Phaeaphodius Reitt. i gruppy vidov Aphodius zangi A. Schm. Rossii, Ukrainy i sopredelnykh stran. Izvestiya Kharkovskogo Entomologicheskogo Obshestva, 6, 5 - 11." type="journal article" year="1998">Kabakov, 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FBF8F4FD1527" blockId="2.[151,1437,222,1358]" box="[151,1436,1073,1094]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FBF8F4FD1527" box="[151,1436,1073,1094]" pageId="2" pageNumber="145">- Epistome rugosely punctate; elytral interstriae finely but distinctly punctate. Length 4.0–6.0 mm...................... 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FB87F4FD15FF" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FB87F4FD15E0" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FB87F4FD15E0" pageId="2" pageNumber="145">
|
||
9 (8). Shape moderately convex, elytra somewhat broadened posteriorly with striae feebly but distinctly crenulate. Length 5.0–6.0 mm. China, Kazakhstan, Mongolia, Russia, Siberia, Taiwan, Turkmenistan...................
|
||
<taxonomicName id="B0D04D05C2720F7FFB0FFBA5F4FD15E0" authority="Gebler, 1848" authorityName="Gebler" authorityYear="1848" box="[1193,1436,1132,1153]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="costalis">
|
||
<emphasis id="45A4EA94C2720F7FFB0FFBA5F46815E0" box="[1193,1289,1132,1153]" italics="true" pageId="2" pageNumber="145">P. costalis</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFAB0FBA5F4F415E0" author="Gebler" box="[1302,1429,1132,1153]" pageId="2" pageNumber="145" refString="Gebler, F. A. (1848) Verzeichniss der im Kolywano-Woskresenskischen Huttenbezirke Sud-West Sibiriens beobachteten Kaefer mit Bemerkungen und Beschreibungen. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 21, 3 - 85." type="journal article" year="1848">Gebler, 1848</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FB40F4FD15FF" blockId="2.[151,1437,222,1358]" box="[151,1436,1161,1182]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FB40F4FD15FF" box="[151,1436,1161,1182]" pageId="2" pageNumber="145">- Shape strongly convex, subcylindrical, elytra almost subparallel-sided with striae not crenulated. Length 4.0–6.0 mm..... 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="CC242E23C2720F7FFF31FB6FF4FD142F" pageId="2" pageNumber="145">
|
||
<paragraph id="776F3686C2720F7FFF31FB6FF4FA1597" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FB6FF4FA1597" pageId="2" pageNumber="145">
|
||
10 (9). Length 5.0–6.0 mm, scutellum superficially punctate on basal half, first metatarsi somewhat shorter than superior spur of tibia, epipharynx with several short and rather stout celtes at apex of corypha. Dorsal color shiny black, sometimes elytra piceous, rarely dirty yellowish brown. Central and eastern Asia...................................
|
||
<taxonomicName id="B0D04D05C2720F7FFB3DFB28F4FA1597" authority="Harold, 1863" authorityName="Harold" authorityYear="1863" box="[1179,1435,1249,1270]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="dauricus">
|
||
<emphasis id="45A4EA94C2720F7FFB3DFB28F4661597" box="[1179,1287,1249,1270]" italics="true" pageId="2" pageNumber="145">P. dauricus</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFAB5FB28F4F51597" author="Harold" box="[1299,1428,1249,1270]" pageId="2" pageNumber="145" refString="Harold, E. (1863) Beitrage zur Kenntniss einiger coprophagen Lamellicornien. (Funftes Stuck). Berliner entomologische Zeitschrift, 7, 327 - 389." type="journal article" year="1863">Harold, 1863</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="776F3686C2720F7FFF31FB37F4FD142F" blockId="2.[151,1437,222,1358]" pageId="2" pageNumber="145">
|
||
<keyLead id="CC2195B3C2720F7FFF31FB37F4FD142F" pageId="2" pageNumber="145">
|
||
- Length 4.0–4.5 mm, scutellum coarsely punctate on basal half, first metatarsi somewhat longer than superior spur of tibia, epipharynx with long stout celtes among other at apex of corypha. Head and pronotum piceous, elytra reddish brown.
|
||
<collectingCountry id="0FC77616C2720F7FFAEAFAD5F070142F" name="Afghanistan" pageId="2" pageNumber="145">Afghanistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="0FC77616C2720F7FFEBCFAF0F0E7142F" box="[282,390,1337,1358]" name="Uzbekistan" pageId="2" pageNumber="145">Uzbekistan</collectingCountry>
|
||
...............................................................
|
||
<taxonomicName id="B0D04D05C2720F7FFB2AFAF0F4FD142F" authority="Balthasar, 1961" authorityName="Balthasar" authorityYear="1961" box="[1164,1436,1337,1358]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="pupillus">
|
||
<emphasis id="45A4EA94C2720F7FFB2AFAF0F591142F" box="[1164,1264,1337,1358]" italics="true" pageId="2" pageNumber="145">P. pupillus</emphasis>
|
||
(
|
||
<bibRefCitation id="13414B77C2720F7FFB5BFAF0F4F4142F" author="Balthasar" box="[1277,1429,1337,1358]" pageId="2" pageNumber="145" refString="Balthasar, V. (1961) Neue Arten der Gattung Aphodius Ill. aus der palaearktischen Region. 102. Beitrag zur Kenntnis der Scarabaeiden, Coleoptera. Acta entomologica Musei nationalis Pragae, 34 (588), 359 - 381." type="journal article" year="1961">Balthasar, 1961</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |