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<document ID-DOI="http://doi.org/10.5281/zenodo.5576092" ID-GBIF-Dataset="84b5feba-a458-4f03-a5b5-35b9c1cc02e1" ID-GBIF-Taxon="128540026" ID-Zenodo-Dep="5576092" checkinTime="1492608070440" checkinUser="jeremy" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2013" docId="EC2A175AFFA4FF91AFC4D614FDBF9D43" docLanguage="en" docName="DankittipakulTavano_Singtripop2013FINAL.imd" docOrigin="Journal of Natural History 47 (23)" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:97EFA23C-7537-4156-8EF0-2D38CF889F5B" docStyle="DocumentStyle{}" docTitle="Jacaena lunulata Dankittipakul, Tavano &amp; Singtripop, 2013, sp. nov." docType="treatment" docVersion="24" lastPageId="20" lastPageNumber="1559" masterDocId="10136F22FFB5FF85AE6DD059FFD39809" masterDocTitle="Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)" masterLastPageNumber="1567" masterPageNumber="1539" pageId="17" pageNumber="1556" updateTime="1635896974979" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dankittipakul, Pakawin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tavano, Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Singtripop, Tippawan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>23</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1539</mods:start>
<mods:end>1567</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.zoobank.org/urn:lsid:zoobank.org:pub:97EFA23C-7537-4156-8EF0-2D38CF889F5B</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2012.763059</mods:identifier>
<mods:identifier type="GBIF-Dataset">84b5feba-a458-4f03-a5b5-35b9c1cc02e1</mods:identifier>
<mods:identifier type="Zenodo-Dep">556127</mods:identifier>
<mods:identifier type="ZooBank">97EFA23C-7537-4156-8EF0-2D38CF889F5B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5576092" ID-GBIF-Taxon="128540026" ID-Zenodo-Dep="5576092" LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43" httpUri="http://treatment.plazi.org/id/EC2A175AFFA4FF91AFC4D614FDBF9D43" lastPageId="20" lastPageNumber="1559" pageId="17" pageNumber="1556">
<subSubSection box="[425,701,1612,1637]" pageId="17" pageNumber="1556" type="nomenclature">
<paragraph blockId="17.[197,928,1612,1669]" box="[425,701,1612,1637]" pageId="17" pageNumber="1556">
<heading box="[425,701,1612,1637]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="1556" reason="4">
<taxonomicName box="[425,611,1612,1637]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="1556" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis bold="true" box="[425,611,1612,1637]" italics="true" pageId="17" pageNumber="1556">Jacaena lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[617,701,1613,1636]" pageId="17" pageNumber="1556" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[197,928,1644,1669]" pageId="17" pageNumber="1556" type="description">
<paragraph blockId="17.[197,928,1612,1669]" box="[197,928,1644,1669]" pageId="17" pageNumber="1556">
(
<figureCitation box="[205,308,1644,1669]" captionStart="Figure 1" captionStartId="3.[83,150,1062,1084]" captionTargetBox="[84,1041,131,1029]" captionTargetId="figure@3.[84,1041,131,1029]" captionTargetPageId="3" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." httpUri="https://zenodo.org/record/556128/files/figure.png" pageId="17" pageNumber="1556">Figures 1</figureCitation>
<collectionCode box="[308,330,1645,1669]" pageId="17" pageNumber="1556">B</collectionCode>
,
<collectionCode box="[341,363,1645,1668]" pageId="17" pageNumber="1556">H</collectionCode>
,
<figureCitation box="[377,391,1645,1669]" captionStart="Figure 2" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E, F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." pageId="17" pageNumber="1556">2</figureCitation>
<collectionCode box="[392,411,1645,1669]" pageId="17" pageNumber="1556">B</collectionCode>
,
<collectionCode box="[421,442,1645,1669]" pageId="17" pageNumber="1556">C</collectionCode>
,
<figureCitation box="[453,465,1645,1669]" captionStart="Figure 3" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A C, E); J. mihun, male, new material (D)." pageId="17" pageNumber="1556">3</figureCitation>
<collectionCode box="[466,488,1645,1669]" pageId="17" pageNumber="1556">A</collectionCode>
<collectionCode box="[503,522,1645,1669]" pageId="17" pageNumber="1556">C</collectionCode>
,
<collectionCode box="[533,553,1645,1668]" pageId="17" pageNumber="1556">E</collectionCode>
,
<figureCitation box="[564,578,1645,1669]" captionStart="Figure 4" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (D F); J. mihun, holotype (G I)." pageId="17" pageNumber="1556">4</figureCitation>
<collectionCode box="[578,598,1645,1669]" pageId="17" pageNumber="1556">D</collectionCode>
<collectionCode box="[614,630,1645,1669]" pageId="17" pageNumber="1556">F</collectionCode>
,
<figureCitation box="[641,654,1644,1668]" captionStart="Figure 6" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." pageId="17" pageNumber="1556">6</figureCitation>
<collectionCode box="[654,675,1644,1668]" pageId="17" pageNumber="1556">A</collectionCode>
<collectionCode box="[691,710,1644,1668]" pageId="17" pageNumber="1556">C</collectionCode>
,
<figureCitation box="[720,734,1645,1669]" captionStart="Figure 7" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." pageId="17" pageNumber="1556">7</figureCitation>
<collectionCode box="[734,754,1645,1669]" pageId="17" pageNumber="1556">C</collectionCode>
,
<figureCitation box="[766,779,1645,1669]" captionStart="Figure 8" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." pageId="17" pageNumber="1556">8</figureCitation>
<collectionCode box="[779,802,1645,1669]" pageId="17" pageNumber="1556">G</collectionCode>
,
<figureCitation box="[814,828,1645,1669]" captionStart="Figure 9" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." pageId="17" pageNumber="1556">9</figureCitation>
<collectionCode box="[828,850,1645,1669]" pageId="17" pageNumber="1556">D</collectionCode>
,
<collectionCode box="[861,881,1645,1668]" pageId="17" pageNumber="1556">E</collectionCode>
,
<figureCitation box="[893,920,1645,1669]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="17" pageNumber="1556">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1557" type="synonymic_list">
<paragraph blockId="18.[151,356,131,185]" pageId="18" pageNumber="1557">
<heading box="[151,356,131,154]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">
<taxonomicName box="[151,356,131,154]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="erawan">
<emphasis box="[151,356,131,154]" italics="true" pageId="18" pageNumber="1557">Sesieutes erawan</emphasis>
</taxonomicName>
</heading>
<bibRefCitation author="Deeleman-Reinhold CL." box="[381,698,129,154]" journalOrPublisher="Leiden, the Netherland: Brill" pageId="18" pageNumber="1557" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." title="Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]" type="book" year="2001">
<heading box="[381,615,129,153]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">Deeleman-Reinhold,</heading>
2001
</bibRefCitation>
: 464, ♂
<heading box="[847,902,129,153]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">from</heading>
<heading box="[928,972,129,153]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">Doi</heading>
<heading box="[998,1111,129,153]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">Inthanon.</heading>
<heading box="[151,351,161,185]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="1557" reason="4">Misidentification.</heading>
</paragraph>
<paragraph blockId="18.[151,1111,209,265]" pageId="18" pageNumber="1557">
<taxonomicName authority="Deeleman-Reinhold, 2001: 461" authorityName="Deeleman-Reinhold" authorityPageNumber="461" authorityYear="2001" box="[151,775,209,234]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis box="[151,412,210,234]" italics="true" pageId="18" pageNumber="1557">Sesieutes schwendingeri</emphasis>
<bibRefCitation author="Deeleman-Reinhold CL." box="[421,720,209,234]" journalOrPublisher="Leiden, the Netherland: Brill" pageId="18" pageNumber="1557" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." title="Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
: 461
</taxonomicName>
, ♂♂ from Doi Inthanon and Doi Suthep. Misidentification.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1557" type="materials_examined">
<paragraph blockId="18.[151,1111,318,446]" box="[151,303,318,343]" pageId="18" pageNumber="1557">
<emphasis box="[151,303,318,343]" italics="true" pageId="18" pageNumber="1557">
<typeStatus box="[151,205,320,343]" pageId="18" pageNumber="1557">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph blockId="18.[151,1111,318,446]" pageId="18" pageNumber="1557">
<materialsCitation ID-GBIF-Occurrence="1457261587" collectingDate="2006-01-11" collectionCode="MHNG" collectorName="P. Dankittipakul" country="Thailand" elevation="1640" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCode="PDC 5746" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province" typeStatus="holotype">
<typeStatus box="[151,262,358,382]" pageId="18" pageNumber="1557">Holotype</typeStatus>
:
<specimenCount box="[270,291,358,382]" pageId="18" pageNumber="1557" type="male"></specimenCount>
,
<collectingCountry box="[304,410,358,382]" name="Thailand" pageId="18" pageNumber="1557" value="Thailand">Thailand</collectingCountry>
:
<collectingRegion box="[419,658,358,382]" country="Thailand" name="Chiang Mai" pageId="18" pageNumber="1557">Chiang Mai Province</collectingRegion>
,
<collectingMunicipality box="[668,899,358,382]" pageId="18" pageNumber="1557">Chomthong District</collectingMunicipality>
,
<collectingMunicipality box="[909,1107,358,382]" pageId="18" pageNumber="1557">Doi Inthanon NP</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97AEFAD1DFFEE69997" box="[151,309,390,414]" country="Thailand" municipality="Chomthong District" name="Doi Inthanon" pageId="18" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Inthanon</location>
, evergreen hill forest,
<elevation box="[547,697,390,414]" metricMagnitude="3" metricUnit="m" metricValue="1.64" metricValueMax="1.68" metricValueMin="1.6" pageId="18" pageNumber="1557" unit="m" value="1640.0" valueMax="1680.0" valueMin="1600.0">
<quantity box="[547,697,390,414]" metricMagnitude="3" metricUnit="m" metricValue="1.64" metricValueMax="1.68" metricValueMin="1.6" pageId="18" pageNumber="1557" unit="m" value="1640.0" valueMax="1680.0" valueMin="1600.0">16001680 m</quantity>
</elevation>
, extraction of soil sample,
<date pageId="18" pageNumber="1557" value="2006-01-11">
<collectingDate pageId="18" pageNumber="1557" value="2006-01-11">11 January 2006</collectingDate>
</date>
,
<collectorName box="[218,405,422,446]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg. (
<collectionCode box="[465,559,422,446]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="18" pageNumber="1557">MHNG</collectionCode>
,
<specimenCode box="[571,692,422,446]" pageId="18" pageNumber="1557">PDC 5746</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[151,1111,467,842]" pageId="18" pageNumber="1557">
<materialsCitation ID-GBIF-Occurrence="1457261583" collectingDate="2000-03-25" collectingDateMax="2000-04-23" collectingDateMin="2000-03-25" collectionCode="MHNG" collectorName="P. Dankittipakul" country="Thailand" elevation="1640" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCode="PDC 5747" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Province" typeStatus="paratype">
<typeStatus box="[151,265,468,492]" pageId="18" pageNumber="1557">Paratypes</typeStatus>
: From type locality, pitfall trap,
<date box="[632,911,467,492]" pageId="18" pageNumber="1557" value="2000-03-25" valueMax="2000-04-23" valueMin="2000-03-25">
<collectingDate box="[632,911,467,492]" pageId="18" pageNumber="1557" value="2000-03-25" valueMax="2000-04-23" valueMin="2000-03-25">25 March23 April 2000</collectingDate>
</date>
,
<collectorName box="[923,1111,467,491]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount box="[209,236,499,523]" pageId="18" pageNumber="1557" type="female">1♀</specimenCount>
(
<collectionCode box="[256,350,499,523]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="18" pageNumber="1557">MHNG</collectionCode>
,
<specimenCode box="[366,492,499,523]" pageId="18" pageNumber="1557">PDC 5747</specimenCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1457261581" collectingDate="2005-09-10" collectionCode="TNHM" collectorName="P. Dankittipakul" country="Thailand" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiang Mai Province">
litter sample,
<collectingDate box="[678,903,499,523]" pageId="18" pageNumber="1557" value="2005-09-10">
<date box="[678,903,499,523]" pageId="18" pageNumber="1557" value="2005-09-10">10 September 2005</date>
</collectingDate>
,
<collectorName box="[919,1111,499,523]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount box="[207,241,531,555]" pageId="18" pageNumber="1557" type="male">2♂</specimenCount>
,
<specimenCount box="[258,285,531,555]" pageId="18" pageNumber="1557" type="female">1♀</specimenCount>
, (
<collectionCode box="[312,403,531,555]" collectionName="TNHM" pageId="18" pageNumber="1557">TNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1457261570" collectingDate="1999-10-09" collectionCode="THNM" collectorName="P. Dankittipakul" country="Thailand" elevation="1650" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCode="PDC 5748" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province">
evergreen hill forest,
<elevation box="[667,754,531,555]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="18" pageNumber="1557" unit="m" value="1650.0">
<quantity box="[667,754,531,555]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="18" pageNumber="1557" unit="m" value="1650.0">1650 m</quantity>
</elevation>
, litter sample,
<collectingDate box="[926,1106,531,555]" pageId="18" pageNumber="1557" value="1999-10-09">
<date box="[926,1106,531,555]" pageId="18" pageNumber="1557" value="1999-10-09">9 October 1999</date>
</collectingDate>
,
<collectorName box="[151,340,563,587]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount box="[405,439,563,587]" pageId="18" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode box="[456,547,563,587]" collectionName="THNM" pageId="18" pageNumber="1557">THNM</collectionCode>
,
<specimenCode box="[561,685,563,587]" pageId="18" pageNumber="1557">PDC 5748</specimenCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1457261569" collectingDate="2000-01-15" collectionCode="THNM" collectorName="P. Dankittipakul" country="Thailand" elevation="1750" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCode="PDC 5749" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province">
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97ACACD26AFCB09A42" box="[705,867,563,587]" country="Thailand" name="Doi Inthanon" pageId="18" pageNumber="1557" stateProvince="Doi Inthanon">Doi Inthanon</location>
, evergreen hill forest,
<elevation box="[151,241,594,618]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="1557" unit="m" value="1750.0">
<quantity box="[151,241,594,618]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="1557" unit="m" value="1750.0">1750 m</quantity>
</elevation>
, litter sample,
<collectingDate box="[419,614,594,619]" pageId="18" pageNumber="1557" value="2000-01-15">
<date box="[419,614,594,619]" pageId="18" pageNumber="1557" value="2000-01-15">15 January 2000</date>
</collectingDate>
,
<collectorName box="[630,822,594,618]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount box="[892,926,595,619]" pageId="18" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode box="[946,1037,595,619]" collectionName="THNM" pageId="18" pageNumber="1557">THNM</collectionCode>
,
<specimenCode pageId="18" pageNumber="1557">PDC 5749</specimenCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1457261588" collectingDate="1999-10-09" collectionCode="THNM" collectorName="P. Dankittipakul" country="Thailand" elevation="1500" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCode="PDC 5750" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Chiang Mai Province">
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97AE8AD22BFE589A83" box="[231,395,626,650]" country="Thailand" name="Doi Inthanon" pageId="18" pageNumber="1557" stateProvince="Doi Inthanon">Doi Inthanon</location>
, evergreen hill forest,
<elevation box="[657,746,626,650]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="1557" unit="m" value="1500.0">
<quantity box="[657,746,626,650]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="1557" unit="m" value="1500.0">1500 m</quantity>
</elevation>
, litter sample,
<collectingDate box="[923,1106,626,651]" pageId="18" pageNumber="1557" value="1999-10-09">
<date box="[923,1106,626,651]" pageId="18" pageNumber="1557" value="1999-10-09">9 October 1999</date>
</collectingDate>
,
<collectorName box="[151,343,658,682]" pageId="18" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount box="[414,449,659,683]" pageId="18" pageNumber="1557" type="male">1♂</specimenCount>
,
<specimenCount box="[468,496,659,683]" pageId="18" pageNumber="1557" type="female">2♀</specimenCount>
(
<collectionCode box="[516,607,659,683]" collectionName="THNM" pageId="18" pageNumber="1557">THNM</collectionCode>
,
<specimenCode box="[624,751,658,683]" pageId="18" pageNumber="1557">PDC 5750</specimenCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1457261590" collectingDate="1985-11-06" collectionCode="MHNG" collectorName="P. J. Schwendinger" country="Thailand" elevation="1250" location="Doi Inthanon" municipality="Chomthong District" pageId="18" pageNumber="1557" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province">
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97AD6AD2CBFC7F9AA3" box="[775,940,658,682]" country="Thailand" name="Doi Inthanon" pageId="18" pageNumber="1557" stateProvince="Doi Inthanon">Doi Inthanon</location>
, evergreen hill forest,
<elevation box="[234,324,690,714]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="1557" unit="m" value="1250.0">
<quantity box="[234,324,690,714]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="1557" unit="m" value="1250.0">1250 m</quantity>
</elevation>
,
<collectingDate box="[341,554,690,714]" pageId="18" pageNumber="1557" value="1985-11-06">
<date box="[341,554,690,714]" pageId="18" pageNumber="1557" value="1985-11-06">6 November 1985</date>
</collectingDate>
,
<collectorName box="[571,777,690,714]" pageId="18" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount box="[848,882,690,715]" pageId="18" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode box="[903,997,691,715]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="18" pageNumber="1557">MHNG</collectionCode>
, misidentified as
<taxonomicName box="[256,368,723,746]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="erawan">
<emphasis box="[256,368,723,746]" italics="true" pageId="18" pageNumber="1557">S. erawan</emphasis>
</taxonomicName>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1457261571" collectingDate="1987-06-18" collectionCode="MHNG" collectorName="P. J. Schwendinger" country="Thailand" elevation="1200" location="Doi Suthep" municipality="Doi Suthep-Pui NP" pageId="18" pageNumber="1557" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Province">
<collectingRegion box="[396,647,722,746]" pageId="18" pageNumber="1557">Chiang Mai Province</collectingRegion>
and District,
<collectingMunicipality box="[819,1051,722,746]" pageId="18" pageNumber="1557">Doi Suthep-Pui NP</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97AA46D28BFF3B9B03" country="Thailand" municipality="Doi Suthep-Pui NP" name="Doi Suthep" pageId="18" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Suthep</location>
,
<quantity box="[246,333,755,779]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="1557" unit="m" value="1200.0">1200 m</quantity>
,
<collectingDate box="[346,499,755,779]" pageId="18" pageNumber="1557" value="1987-06-18">
<date box="[346,499,755,779]" pageId="18" pageNumber="1557" value="1987-06-18">18 June 1987</date>
</collectingDate>
,
<collectorName box="[513,716,754,778]" pageId="18" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount box="[780,808,754,779]" pageId="18" pageNumber="1557" type="female">1♀</specimenCount>
(
<collectionCode box="[826,917,755,779]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="18" pageNumber="1557">MHNG</collectionCode>
, misidentified as
<taxonomicName box="[151,332,786,810]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis box="[151,332,786,810]" italics="true" pageId="18" pageNumber="1557">S. schwendingeri</emphasis>
</taxonomicName>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1457261576" collectingDate="1986-11-31" collectingDateMax="1987-01-02" collectingDateMin="1986-11-31" collectionCode="RMNH" collectorName="P. J. Schwendinger" country="Thailand" elevation="960" httpUri="https://data.biodiversitydata.nl/naturalis/specimen/RMNH.ARA.15205" location="Doi Suthep" municipality="Doi Suthep" pageId="18" pageNumber="1557" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province">
<location LSID="urn:lsid:plazi:treatment:EC2A175AFFA4FF91AFC4D614FDBF9D43:615CF097FFA7FF97AF0ED34BFE359B23" box="[355,486,786,810]" country="Thailand" municipality="Doi Suthep" name="Doi Suthep" pageId="18" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Suthep</location>
,
<elevation box="[497,568,786,810]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="18" pageNumber="1557" unit="m" value="960.0">
<quantity box="[497,568,786,810]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="18" pageNumber="1557" unit="m" value="960.0">960 m</quantity>
</elevation>
, pitfall trap,
<collectingDate box="[711,922,786,810]" pageId="18" pageNumber="1557" value="1986-11-31">
<date box="[711,922,786,810]" pageId="18" pageNumber="1557" value="1986-11-31">31 November 1986</date>
</collectingDate>
<collectingDate box="[936,1106,786,811]" pageId="18" pageNumber="1557" value="1987-01-02">
<date box="[936,1106,786,811]" pageId="18" pageNumber="1557" value="1987-01-02">2 January 1987</date>
</collectingDate>
,
<collectorName box="[151,351,818,842]" pageId="18" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount box="[410,444,818,842]" pageId="18" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode box="[459,553,818,842]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="18" pageNumber="1557">RMNH</collectionCode>
, misidentified as
<taxonomicName box="[746,927,818,842]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis box="[746,927,818,842]" italics="true" pageId="18" pageNumber="1557">S. schwendingeri</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1557" type="diagnosis">
<paragraph blockId="18.[151,1111,893,1212]" box="[151,259,893,916]" pageId="18" pageNumber="1557">
<emphasis box="[151,259,893,916]" italics="true" pageId="18" pageNumber="1557">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="18.[151,1111,893,1212]" pageId="18" pageNumber="1557">
Males of
<taxonomicName box="[262,379,932,956]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis box="[262,379,932,956]" italics="true" pageId="18" pageNumber="1557">J. lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[388,476,933,956]" pageId="18" pageNumber="1557" rank="species">sp. nov.</taxonomicNameLabel>
are similar to those of
<taxonomicName box="[750,930,932,956]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis box="[750,930,932,956]" italics="true" pageId="18" pageNumber="1557">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[940,1064,932,956]" pageId="18" pageNumber="1557">comb. nov.</taxonomicNameLabel>
but differ by having a crescent-shaped conductor that is distinctly broader and strongly curved backward (
<figureCitation box="[372,485,996,1020]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="18" pageNumber="1557">Figures 6</figureCitation>
<collectionCode box="[485,507,996,1020]" pageId="18" pageNumber="1557">A</collectionCode>
,
<figureCitation box="[525,540,996,1020]" captionStart="Figure 9" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." pageId="18" pageNumber="1557">9</figureCitation>
<collectionCode box="[540,563,996,1020]" pageId="18" pageNumber="1557">D</collectionCode>
,
<collectionCode box="[581,601,996,1020]" pageId="18" pageNumber="1557">E</collectionCode>
), a tegulum that is greatly expanded, and the distal RTA of the male palp that is digitiform (
<figureCitation box="[758,858,1027,1051]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="18" pageNumber="1557">Figure 6</figureCitation>
<collectionCode box="[858,879,1027,1051]" pageId="18" pageNumber="1557">C</collectionCode>
). In comparison, in
<taxonomicName box="[151,328,1060,1084]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis box="[151,328,1060,1084]" italics="true" pageId="18" pageNumber="1557">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[336,459,1059,1083]" pageId="18" pageNumber="1557">comb. nov.</taxonomicNameLabel>
the conductor is narrower and its sharply pointed apex is abruptly bent (
<figureCitation box="[315,412,1091,1116]" captionStart="Figure 4" captionStartId="6.[151,218,1390,1412]" captionTargetBox="[152,1109,131,1356]" captionTargetId="figure@6.[152,1109,131,1357]" captionTargetPageId="6" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (D F); J. mihun, holotype (G I)." httpUri="https://zenodo.org/record/556131/files/figure.png" pageId="18" pageNumber="1557">Figure 4</figureCitation>
<collectionCode box="[411,433,1092,1116]" pageId="18" pageNumber="1557">A</collectionCode>
,
<collectionCode box="[444,465,1092,1116]" pageId="18" pageNumber="1557">C</collectionCode>
), the tegulum and the distal RTA are significantly smaller. Females can be distinguished by the rather simple internal genitalia, the ID of which consists of circular proximal ducts, the more convoluted ID, and the poorly defined spermathecae (
<figureCitation box="[320,414,1187,1212]" captionStart="Figure 8" captionStartId="12.[151,218,913,935]" captionTargetBox="[152,1108,131,877]" captionTargetId="figure@12.[152,1109,131,879]" captionTargetPageId="12" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." httpUri="https://zenodo.org/record/556135/files/figure.png" pageId="18" pageNumber="1557">Figure 8</figureCitation>
<collectionCode box="[414,436,1188,1212]" pageId="18" pageNumber="1557">G</collectionCode>
cf.
<figureCitation box="[475,489,1188,1212]" captionStart="Figure 8" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." pageId="18" pageNumber="1557">8</figureCitation>
<collectionCode box="[490,509,1188,1212]" pageId="18" pageNumber="1557">F</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1557" type="etymology">
<paragraph blockId="18.[151,1111,1272,1368]" box="[151,271,1272,1296]" pageId="18" pageNumber="1557">
<emphasis box="[151,271,1272,1296]" italics="true" pageId="18" pageNumber="1557">Etymology</emphasis>
</paragraph>
<paragraph blockId="18.[151,1111,1272,1368]" pageId="18" pageNumber="1557">
The specific epithet is derived from Latin adjective (
<emphasis box="[773,873,1312,1336]" italics="true" pageId="18" pageNumber="1557">lunulatus</emphasis>
= crescent-shaped), referring to the lamina-shaped conductor on the male palp.
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="1559" pageId="18" pageNumber="1557" type="description">
<paragraph blockId="18.[151,1111,1430,1653]" box="[151,278,1430,1453]" pageId="18" pageNumber="1557">
<emphasis box="[151,278,1430,1453]" italics="true" pageId="18" pageNumber="1557">Description</emphasis>
</paragraph>
<paragraph blockId="18.[151,1111,1430,1653]" lastBlockId="19.[83,1043,129,505]" lastPageId="19" lastPageNumber="1558" pageId="18" pageNumber="1557">
<emphasis box="[151,209,1469,1493]" italics="true" pageId="18" pageNumber="1557">Male</emphasis>
(
<typeStatus box="[223,328,1468,1492]" pageId="18" pageNumber="1557">holotype</typeStatus>
). Total length 6.6. Carapace 3.2 long, 2.5 wide. Opisthosoma 3.4 long, 2.0 wide. Eye sizes and interdistances:
<collectionCode box="[629,693,1501,1525]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="18" pageNumber="1557">AME</collectionCode>
0.15; ALE 0.13; PME 0.10; PLE 0.13;
<collectionCode box="[215,279,1533,1557]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="18" pageNumber="1557">AME</collectionCode>
<collectionCode box="[293,357,1533,1557]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="18" pageNumber="1557">AME</collectionCode>
0.06;
<collectionCode box="[431,494,1533,1557]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="18" pageNumber="1557">AME</collectionCode>
ALE 0.05; PMEPME 0.15; PMEPLE 0.16. MOQ 0.33 long; front width 0.33; back width 0.35. Leg formula 4123. Spination. Leg I: femur 11pl; tibia 8pl 7rl; metatarsus 4pl 4rl. Leg II: tibia 7pl 6rl; metatarsus 4pl 4rl. Leg III: tibia 1
<date box="[237,263,1629,1653]" pageId="18" pageNumber="1557">1v</date>
1pl; metatarsus 1
<date box="[479,505,1629,1653]" pageId="18" pageNumber="1557">1v</date>
1pl. Leg IV: tibia 12
<date box="[767,794,1629,1653]" pageId="18" pageNumber="1557">2v</date>
11rl; metatarsus 111
<date box="[1085,1111,1629,1653]" pageId="18" pageNumber="1557">1v</date>
1pl 1rl. Leg measurements: Leg I 8.1 (2.3, 0.9, 2.2, 1.8, 0.9); II 7.5 (2.1, 0.9, 1.9, 1.7, 0.9); III 6.7 (1.9, 0.9, 1.4, 1.6, 0.9); IV 9.4 (2.6, 1.0, 2.1, 2.5, 1.2).
</paragraph>
<paragraph blockId="19.[83,1043,129,505]" pageId="19" pageNumber="1558">
Colouration and pattern (
<figureCitation box="[416,510,193,218]" captionStart="Figure 1" captionStartId="3.[83,150,1062,1084]" captionTargetBox="[84,1041,131,1029]" captionTargetId="figure@3.[84,1041,131,1029]" captionTargetPageId="3" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." httpUri="https://zenodo.org/record/556128/files/figure.png" pageId="19" pageNumber="1558">Figure 1</figureCitation>
<collectionCode box="[510,532,194,218]" pageId="19" pageNumber="1558">B</collectionCode>
): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Coxae, trochanters and femora dark brown; patellae and tibiae brown, distal part of tibiae orange; metatarsi III brown, IIIIV orange; tarsi orange. Opisthosoma ovoid; dorsum of opisthosoma dark grey, with pattern consisting of one pair of pale patches, followed by four transverse chevrons and pale area above spinnerets. Dorsal scutum occupying approximately one-third of opisthosoma length, posterior margin rounded, with oval extensions fusing with dorsal muscle apodemes.
</paragraph>
<paragraph blockId="19.[83,1043,129,505]" lastBlockId="20.[151,1111,129,282]" lastPageId="20" lastPageNumber="1559" pageId="19" pageNumber="1558">
Palp (
<figureCitation box="[194,304,448,472]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="19" pageNumber="1558">Figures 6</figureCitation>
<collectionCode box="[304,325,448,472]" pageId="19" pageNumber="1558">A</collectionCode>
<collectionCode box="[341,360,448,472]" pageId="19" pageNumber="1558">C</collectionCode>
,
<figureCitation box="[375,389,449,473]" captionStart="Figure 9" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." pageId="19" pageNumber="1558">9</figureCitation>
<collectionCode box="[389,411,449,473]" pageId="19" pageNumber="1558">D</collectionCode>
,
<collectionCode box="[427,447,449,472]" pageId="19" pageNumber="1558">E</collectionCode>
,
<figureCitation box="[462,489,449,473]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="19" pageNumber="1558">12</figureCitation>
<collectionCode box="[489,511,449,473]" pageId="19" pageNumber="1558">A</collectionCode>
<collectionCode box="[527,546,449,473]" pageId="19" pageNumber="1558">C</collectionCode>
): Proximal RTA triangular in ventral view (
<figureCitation box="[90,199,480,505]" captionStart="Figure 12" captionStartId="19.[83,150,1508,1530]" captionTargetBox="[84,1039,572,1475]" captionTargetId="figure@19.[84,1041,567,1475]" captionTargetPageId="19" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." httpUri="https://zenodo.org/record/556139/files/figure.png" pageId="19" pageNumber="1558">Figure 12</figureCitation>
<collectionCode box="[199,221,481,505]" pageId="19" pageNumber="1558">A</collectionCode>
); apical surface partially membranous, slightly excavated, forming shallow concavity (
<figureCitation box="[277,376,129,154]" captionStart="Figure 9" captionStartId="15.[83,150,886,908]" captionTargetBox="[86,1041,133,850]" captionTargetId="figure@15.[84,1041,131,853]" captionTargetPageId="15" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." httpUri="https://zenodo.org/record/556136/files/figure.png" pageId="20" pageNumber="1559">Figure 9</figureCitation>
<collectionCode box="[376,399,130,154]" pageId="20" pageNumber="1559">D</collectionCode>
,
<collectionCode box="[414,435,130,154]" pageId="20" pageNumber="1559">E</collectionCode>
). Distal RTA digitiform, its apex blunt (
<figureCitation box="[910,1022,129,153]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="20" pageNumber="1559">Figures 6</figureCitation>
<collectionCode box="[1021,1041,129,153]" pageId="20" pageNumber="1559">C</collectionCode>
,
<figureCitation box="[1057,1085,130,154]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[1084,1106,130,154]" pageId="20" pageNumber="1559">A</collectionCode>
,
<collectionCode box="[151,173,162,186]" pageId="20" pageNumber="1559">C</collectionCode>
). Conductor (
<figureCitation box="[341,451,161,185]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="20" pageNumber="1559">Figures 6</figureCitation>
<collectionCode box="[451,473,161,185]" pageId="20" pageNumber="1559">A</collectionCode>
, 9
<collectionCode box="[504,527,162,186]" pageId="20" pageNumber="1559">D</collectionCode>
,
<collectionCode box="[543,563,162,185]" pageId="20" pageNumber="1559">E</collectionCode>
,
<figureCitation box="[580,606,162,186]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[606,627,162,186]" pageId="20" pageNumber="1559">A</collectionCode>
<collectionCode box="[642,660,162,186]" pageId="20" pageNumber="1559">C</collectionCode>
) crescent-shaped, enlarged and broad, semi-transparent, with heavily pigmented outer margin, sharply pointed apex curving backwards. Tegular apophysis hyaline, lanceolate, originating behind base of conductor, aligning with tip of embolus (
<figureCitation box="[608,715,257,281]" captionStart="Figure 6" captionStartId="10.[151,218,1105,1127]" captionTargetBox="[152,1109,134,1071]" captionTargetId="figure@10.[152,1109,129,1071]" captionTargetPageId="10" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." httpUri="https://zenodo.org/record/556133/files/figure.png" pageId="20" pageNumber="1559">Figures 6</figureCitation>
<collectionCode box="[715,737,257,281]" pageId="20" pageNumber="1559">A</collectionCode>
,
<figureCitation box="[749,764,258,282]" captionStart="Figure 9" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." pageId="20" pageNumber="1559">9</figureCitation>
<collectionCode box="[764,787,258,282]" pageId="20" pageNumber="1559">D</collectionCode>
).
</paragraph>
<caption httpUri="https://zenodo.org/record/556139/files/figure.png" pageId="19" pageNumber="1558" targetBox="[84,1039,572,1475]" targetPageId="19">
<paragraph blockId="19.[83,1044,1508,1647]" pageId="19" pageNumber="1558">
Figure 12.
<taxonomicName box="[199,367,1508,1530]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="1558" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis box="[199,367,1508,1530]" italics="true" pageId="19" pageNumber="1558">Jacaena lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[374,449,1509,1530]" pageId="19" pageNumber="1558" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca.
</paragraph>
</caption>
<paragraph blockId="20.[151,1112,321,1047]" pageId="20" pageNumber="1559">
<emphasis box="[151,230,321,345]" italics="true" pageId="20" pageNumber="1559">Female</emphasis>
(
<typeStatus box="[245,347,321,345]" pageId="20" pageNumber="1559">paratype</typeStatus>
,
<collectionCode box="[358,449,322,345]" collectionName="Libya, Tripoli, Natural History Museum" pageId="20" pageNumber="1559">NHML</collectionCode>
,
<collectionCode box="[460,517,321,345]" pageId="20" pageNumber="1559">PDC</collectionCode>
5747). Total length 8.1. Carapace 3.3 long, 2.5 wide. Opisthosoma 4.5 long, 2.7 wide. Eye sizes and interdistances:
<collectionCode box="[857,921,353,377]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="20" pageNumber="1559">AME</collectionCode>
0.11; ALE 0.13; PME 0.08; PLE 0.13;
<collectionCode box="[409,473,385,409]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="20" pageNumber="1559">AME</collectionCode>
<collectionCode box="[487,551,385,409]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="20" pageNumber="1559">AME</collectionCode>
0.08;
<collectionCode box="[624,687,385,409]" collectionName="USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum" pageId="20" pageNumber="1559">AME</collectionCode>
ALE 0.05; PMEPME 0.15; PME PLE 0.16. MOQ 0.38 long; front width 0.31; back width 0.35. Leg formula 1423. Spination. Leg I: femur 11pl; tibia 8pl 7rl; metatarsus 4pl 4rl. Leg II: tibia 7pl 6rl; metatarsus 4pl 4rl. Leg III: tibia 2
<date box="[555,578,481,505]" pageId="20" pageNumber="1559">1v</date>
1pl; metatarsus 1
<date box="[795,821,481,505]" pageId="20" pageNumber="1559">1v</date>
1pl. Leg IV: tibia 11
<date box="[1082,1111,481,505]" pageId="20" pageNumber="1559">2v</date>
1rl; metatarsus 111
<date box="[406,432,513,537]" pageId="20" pageNumber="1559">1v</date>
1pl 1rl. Leg measurements: Leg I 8.6 (2.3, 1.0, 2.3, 2.0, 1.0); II 7.9 (2.2, 1.0, 1.9, 1.8, 1.0); III 6.6 (1.8, 2.8, 1.5, 1.5, 1.0); IV 8.3 (2.3, 0.8, 2.0, 2.2, 1.0).
</paragraph>
<paragraph blockId="20.[151,1112,321,1047]" pageId="20" pageNumber="1559">
Colouration and pattern (
<figureCitation box="[484,582,608,632]" captionStart="Figure 2" captionStartId="4.[151,218,1094,1116]" captionTargetBox="[152,1109,130,1060]" captionTargetId="figure@4.[152,1109,130,1061]" captionTargetPageId="4" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E, F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." httpUri="https://zenodo.org/record/556129/files/figure.png" pageId="20" pageNumber="1559">Figure 2</figureCitation>
<collectionCode box="[582,601,608,632]" pageId="20" pageNumber="1559">B</collectionCode>
): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Legs III brown, femora slightly darker than other segments, tibiae orangebrown distally; legs IIIIV orange-brown except femora dark brown. Opisthosoma elongate-ovoid; dorsum grey, with pair of pale patches connected anteriorly, followed by four chevrons and pale area above spinnerets (first pair very faint and narrowed, disconnected medially).
</paragraph>
<paragraph blockId="20.[151,1112,321,1047]" pageId="20" pageNumber="1559">
Copulatory organ (
<figureCitation box="[410,517,831,856]" captionStart="Figure 7" captionStartId="11.[83,150,753,775]" captionTargetBox="[84,1041,131,718]" captionTargetId="figure@11.[84,1041,131,720]" captionTargetPageId="11" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." httpUri="https://zenodo.org/record/556134/files/figure.png" pageId="20" pageNumber="1559">Figures 7</figureCitation>
<collectionCode box="[517,537,832,856]" pageId="20" pageNumber="1559">C</collectionCode>
,
<figureCitation box="[549,563,832,856]" captionStart="Figure 8" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." pageId="20" pageNumber="1559">8</figureCitation>
<collectionCode box="[563,582,832,856]" pageId="20" pageNumber="1559">F</collectionCode>
,
<figureCitation box="[594,622,832,856]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[622,644,832,856]" pageId="20" pageNumber="1559">D</collectionCode>
,
<collectionCode box="[655,676,832,856]" pageId="20" pageNumber="1559">E</collectionCode>
): Copulatory orifices situated medially on heavily sclerotized epigynal region (
<figureCitation box="[592,700,863,887]" captionStart="Figure 7" captionStartId="11.[83,150,753,775]" captionTargetBox="[84,1041,131,718]" captionTargetId="figure@11.[84,1041,131,720]" captionTargetPageId="11" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." httpUri="https://zenodo.org/record/556134/files/figure.png" pageId="20" pageNumber="1559">Figures 7</figureCitation>
<collectionCode box="[701,721,863,887]" pageId="20" pageNumber="1559">C</collectionCode>
,
<figureCitation box="[734,763,863,887]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[764,787,863,887]" pageId="20" pageNumber="1559">D</collectionCode>
). Proximal half of insemination ducts forming two loops encircling copulatory ducts (
<figureCitation box="[832,943,895,919]" captionStart="Figure 8" captionStartId="12.[151,218,913,935]" captionTargetBox="[152,1108,131,877]" captionTargetId="figure@12.[152,1109,131,879]" captionTargetPageId="12" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." httpUri="https://zenodo.org/record/556135/files/figure.png" pageId="20" pageNumber="1559">Figures 8</figureCitation>
<collectionCode box="[942,966,895,919]" pageId="20" pageNumber="1559">G</collectionCode>
,
<figureCitation box="[982,1010,895,919]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[1010,1029,895,919]" pageId="20" pageNumber="1559">E</collectionCode>
); inner loop distinctly enlarged, filled with dark substance (
<figureCitation box="[750,847,927,951]" captionStart="Figure 8" captionStartId="12.[151,218,913,935]" captionTargetBox="[152,1108,131,877]" captionTargetId="figure@12.[152,1109,131,879]" captionTargetPageId="12" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." httpUri="https://zenodo.org/record/556135/files/figure.png" pageId="20" pageNumber="1559">Figure 8</figureCitation>
<collectionCode box="[847,871,927,951]" pageId="20" pageNumber="1559">G</collectionCode>
); distal half of ducts ascending anteriorly then traversing and descending to connect with poorly defined spermathecae (
<figureCitation box="[327,439,990,1015]" captionStart="Figure 8" captionStartId="12.[151,218,913,935]" captionTargetBox="[152,1108,131,877]" captionTargetId="figure@12.[152,1109,131,879]" captionTargetPageId="12" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A G). Jacaena mihun (A D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." httpUri="https://zenodo.org/record/556135/files/figure.png" pageId="20" pageNumber="1559">Figures 8</figureCitation>
<collectionCode box="[439,463,991,1015]" pageId="20" pageNumber="1559">G</collectionCode>
,
<figureCitation box="[481,509,991,1015]" captionStart="Figure 12" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." pageId="20" pageNumber="1559">12</figureCitation>
<collectionCode box="[509,528,991,1015]" pageId="20" pageNumber="1559">E</collectionCode>
); glandular pores situated at anterior margin of spermathecae (
<figureCitation box="[320,429,1022,1047]" captionStart="Figure 12" captionStartId="19.[83,150,1508,1530]" captionTargetBox="[84,1039,572,1475]" captionTargetId="figure@19.[84,1041,567,1475]" captionTargetPageId="19" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." httpUri="https://zenodo.org/record/556139/files/figure.png" pageId="20" pageNumber="1559">Figure 12</figureCitation>
<collectionCode box="[429,448,1023,1047]" pageId="20" pageNumber="1559">E</collectionCode>
,
<collectionCode box="[459,502,1023,1047]" pageId="20" pageNumber="1559">GF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="1559" type="biology_ecology">
<paragraph blockId="20.[151,1111,1121,1216]" box="[151,319,1121,1145]" pageId="20" pageNumber="1559">
<emphasis box="[151,319,1121,1145]" italics="true" pageId="20" pageNumber="1559">Natural history</emphasis>
</paragraph>
<paragraph blockId="20.[151,1111,1121,1216]" pageId="20" pageNumber="1559">
<typeStatus box="[151,217,1161,1185]" pageId="20" pageNumber="1559">Types</typeStatus>
of this new species were collected in evergreen hill forests of Doi Suthep and Doi Inthanon between
<date box="[362,524,1192,1216]" pageId="20" pageNumber="1559" value="1250" valueMax="1750">
1250 and
<quantity box="[471,555,1192,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="1559" unit="m" value="1750.0">1750 m</quantity>
</date>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,620,1291,1354]" pageId="20" pageNumber="1559" type="distribution">
<paragraph blockId="20.[151,620,1291,1354]" box="[151,284,1291,1315]" pageId="20" pageNumber="1559">
<emphasis box="[151,284,1291,1315]" italics="true" pageId="20" pageNumber="1559">Distribution</emphasis>
</paragraph>
<paragraph blockId="20.[151,620,1291,1354]" box="[151,620,1330,1354]" pageId="20" pageNumber="1559">Chiang Mai Province, northern Thailand.</paragraph>
</subSubSection>
</treatment>
</document>