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<document id="4B515FE00D96848CA59D76E8215CD05A" ID-CLB-Dataset="63563" ID-DOI="10.5281/zenodo.6608102" ID-GBIF-Dataset="3629b48a-c3e7-4f82-846f-187418823ef3" ID-ISBN="978-84-96553-99-6" ID-Zenodo-Dep="6608102" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654085738184" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2015" docId="EA7087C1FF82246FFF07FC2B07BA07C3" docLanguage="en" docName="hbmw_5_Dasyuridae_0232.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 5 Monotremes and Marsupials, Barcelona: Lynx Edicions" docTitle="Planigale ingrami" docType="treatment" docVersion="10" lastPageNumber="328" masterDocId="1649FFB9FFA92444FFA3FFF10F480164" masterDocTitle="Dasyuridae" masterLastPageNumber="348" masterPageNumber="232" pageNumber="328" updateTime="1699338574489" updateUser="ExternalLinkService">
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<mods:title id="A279A0874FB4687770F07DEE2BA9B14B">Dasyuridae</mods:title>
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<mods:namePart id="5EB7C2C5A61E39793DCF788445A14D0B">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="584D06C1C1B47F939278D71A35014ECC">Don E. Wilson</mods:namePart>
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<mods:title id="5F2F0A00AFA1E0BF75A6031189C0C45D">Handbook of the Mammals of the World Volume 5 Monotremes and Marsupials</mods:title>
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<treatment id="EA7087C1FF82246FFF07FC2B07BA07C3" ID-DOI="http://doi.org/10.5281/zenodo.6602843" ID-GBIF-Taxon="195729070" ID-Zenodo-Dep="6602843" LSID="urn:lsid:plazi:treatment:EA7087C1FF82246FFF07FC2B07BA07C3" httpUri="http://treatment.plazi.org/id/EA7087C1FF82246FFF07FC2B07BA07C3" lastPageNumber="328" pageId="43" pageNumber="328">
<subSubSection id="2AC3655CFF82246FFF07FC2B0F95056C" box="[164,221,986,1032]" pageId="43" pageNumber="328" type="multiple">
<paragraph id="626636D7FF82246FFF07FC2B0F95056C" blockId="43.[161,1251,986,1115]" box="[164,221,986,1032]" pageId="43" pageNumber="328">
<heading id="392E81BBFF82246FFF07FC2B0F95056C" box="[164,221,986,1032]" pageId="43" pageNumber="328">
<figureCitation id="FAE22A52FF82246FFF07FC2B0F95056C" box="[164,221,986,1032]" captionStart="On" captionStartId="36.[104,134,3331,3356]" captionTargetBox="[11,2730,13,3642]" captionTargetPageId="35" captionText="On following pages: 41. Short-furred Dasyure (Murexia longicaudata); 42. Broad-striped Dasyure (Murexia rothschildi); 43. Long-nosed Dasyure (Murexia naso); 44. Red-tailed Phascogale (Phascogale calura); 45. Northern Brush-tailed Phascogale (Phascogale pirata); 46. Common Brush-tailed Phascogale (Phascogale tapoatafa); 47. Giles's Planigale (Planigale giles); 48. Long-tailed Planigale (Planigale ingrami); 49. Common Planigale (Planigale maculata); 50. Papuan Planigale (Planigale novaeguineae); 51. Narrow-nosed Planigale (Planigale tenuirostris); 52. Wongai Ningaui (Ningauiride); 53. Pilbara Ningaui (Ningaui timealeyi); 54. Southern Ningaui (Ningaui yvonneae); 55. Kultarr (Antechinomys laniger)." figureDoi="http://doi.org/10.5281/zenodo.6608326" httpUri="https://zenodo.org/record/6608326/files/figure.png" pageId="43" pageNumber="328">48.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="2AC3655CFF82246FFF4DFC2B0DEF056C" box="[238,679,986,1032]" pageId="43" pageNumber="328" type="vernacular_names">
<paragraph id="626636D7FF82246FFF4DFC2B0DEF056C" blockId="43.[161,1251,986,1115]" box="[238,679,986,1032]" pageId="43" pageNumber="328">
<heading id="392E81BBFF82246FFF4DFC2B0DEF056C" box="[238,679,986,1032]" pageId="43" pageNumber="328">
<vernacularName id="ECDA46F9FF82246FFF4DFC2B0DEF056C" box="[238,679,986,1032]" pageId="43" pageNumber="328">Long-tailed Planigale</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFD4EFC2B0B79056C" box="[749,1073,986,1032]" pageId="43" pageNumber="328" type="nomenclature">
<paragraph id="626636D7FF82246FFD4EFC2B0B79056C" blockId="43.[161,1251,986,1115]" box="[749,1073,986,1032]" pageId="43" pageNumber="328">
<heading id="392E81BBFF82246FFD4EFC2B0B79056C" box="[749,1073,986,1032]" pageId="43" pageNumber="328">
<taxonomicName id="A5D94D54FF82246FFD4EFC2B0B79056C" ID-CoL="4JJYY" baseAuthorityName="Thomas" baseAuthorityYear="1906" box="[749,1073,986,1032]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">
<emphasis id="50ADEAC5FF82246FFD4EFC2B0B79056C" box="[749,1073,986,1032]" italics="true" pageId="43" pageNumber="328">Planigale ingrami</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFF00FBEB0C540532" pageId="43" pageNumber="328" type="vernacular_names">
<paragraph id="626636D7FF82246FFF00FBEB0BAA054B" blockId="43.[161,1251,986,1115]" box="[163,1250,1050,1071]" pageId="43" pageNumber="328">
<heading id="392E81BBFF82246FFF00FBEB0BAA054B" box="[163,1250,1050,1071]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF00FBEB0FA7054B" bold="true" box="[163,239,1050,1071]" pageId="43" pageNumber="328">French:</emphasis>
<vernacularName id="ECDA46F9FF82246FFF5BFBEB0EE7054B" box="[248,431,1050,1071]" pageId="43" pageNumber="328">Planigale d'Ingram</vernacularName>
/
<emphasis id="50ADEAC5FF82246FFE67FBEB0D57054B" bold="true" box="[452,543,1050,1071]" pageId="43" pageNumber="328">German:</emphasis>
<vernacularName id="ECDA46F9FF82246FFD8AFBEB0CC4054B" box="[553,908,1050,1071]" pageId="43" pageNumber="328">Langschwanz-Flachkopfbeutelmaus</vernacularName>
/
<emphasis id="50ADEAC5FF82246FFC02FBEB0CB4054B" bold="true" box="[929,1020,1050,1071]" pageId="43" pageNumber="328">Spanish:</emphasis>
<vernacularName id="ECDA46F9FF82246FFBA5FBEB0BAA054B" box="[1030,1250,1050,1071]" pageId="43" pageNumber="328">Planigalo de cola larga</vernacularName>
</heading>
</paragraph>
<paragraph id="626636D7FF82246FFF01FBB00C540532" blockId="43.[161,1251,986,1115]" box="[162,796,1089,1110]" pageId="43" pageNumber="328">
<heading id="392E81BBFF82246FFF01FBB00C540532" box="[162,796,1089,1110]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF01FBB00ED10532" bold="true" box="[162,409,1089,1110]" pageId="43" pageNumber="328">Other common names:</emphasis>
<vernacularName id="ECDA46F9FF82246FFE00FBB00D120532" box="[419,602,1089,1110]" pageId="43" pageNumber="328">Blacksoil Planigale</vernacularName>
,
<vernacularName id="ECDA46F9FF82246FFDCBFBB00C540532" box="[616,796,1089,1110]" pageId="43" pageNumber="328">Ingrams Planigale</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFCA5FB750C1805A9" pageId="43" pageNumber="328" type="reference_group">
<paragraph id="626636D7FF82246FFCA5FB750C1805A9" blockId="43.[773,1366,1156,1583]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFCA5FB750CEA05C1" bold="true" box="[774,930,1156,1189]" pageId="43" pageNumber="328">Taxonomy.</emphasis>
<taxonomicName id="A5D94D54FF82246FFC1EFB750C1805A9" authority="Thomas" authorityName="Thomas" authorityYear="1906" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">Phascogale ingrami Thomas, 1906,</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFCCDFB410CC00590" pageId="43" pageNumber="328" type="materials_examined">
<paragraph id="626636D7FF82246FFCCDFB410CC00590" blockId="43.[773,1366,1156,1583]" pageId="43" pageNumber="328">
<materialsCitation id="D2B13C8AFF82246FFCCDFB410CC00590" ID-GBIF-Occurrence="3802852326" country="Australia" location="Alexandria" municipality="Alexandria" pageId="43" pageNumber="328" specimenCount="1" stateProvince="Northern Territory">
<collectingMunicipality id="8202ACADFF82246FFCCDFB410B4405A9" box="[878,1036,1200,1229]" pageId="43" pageNumber="328">Alexandria</collectingMunicipality>
,
<collectingRegion id="A01DF835FF82246FFB92FB410A1805A9" box="[1073,1360,1200,1229]" country="Australia" name="Northern Territory" pageId="43" pageNumber="328">Northern Territory</collectingRegion>
,
<collectingCountry id="1ACE7647FF82246FFCA6FB260CCC0590" box="[773,900,1239,1268]" name="Australia" pageId="43" pageNumber="328">Australia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFCA4FB0A0BF408F6" pageId="43" pageNumber="328" type="discussion">
<paragraph id="626636D7FF82246FFCA4FB0A0BF408F6" blockId="43.[773,1366,1156,1583]" lastBlockId="43.[160,1368,1589,3474]" pageId="43" pageNumber="328">
E. Le G. Troughton erected the genus
<taxonomicName id="A5D94D54FF82246FFCA4FAD30CC90427" box="[775,897,1314,1347]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Planigale</taxonomicName>
in 1928 to accommodate three species of small, flatheaded dasyurids known from Australia at that time. Much later, in 1976, M. Archer conducted a comprehensive revision of the genus
<taxonomicName id="A5D94D54FF82246FFB74FA310A180485" box="[1239,1360,1472,1505]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Planigale</taxonomicName>
. He recognized three species groups based on morphology: the
<taxonomicName id="A5D94D54FF82246FFBE5F9FF0BA0074B" authorityName="Kerr" authorityYear="1792" box="[1094,1256,1550,1583]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="maculata">P. maculata</taxonomicName>
group, containing
<taxonomicName id="A5D94D54FF82246FFEE7F9C40E950732" authorityName="Kerr" authorityYear="1792" box="[324,477,1589,1622]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="maculata">P. maculata</taxonomicName>
and
<taxonomicName id="A5D94D54FF82246FFD87F9C40DBA0732" box="[548,754,1589,1622]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="novaeguineae">P. novaeguineae</taxonomicName>
, the
<taxonomicName id="A5D94D54FF82246FFC99F9C40C880732" authorityName="Thomas" authorityYear="1906" box="[826,960,1589,1622]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
group, containing
<taxonomicName id="A5D94D54FF82246FFB72F9C40A100732" authorityName="Thomas" authorityYear="1906" box="[1233,1368,1589,1622]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
and P. tenwirostris, both extremely similar morphologically; and a monotypic group containing FP.
<taxonomicName id="A5D94D54FF82246FFECFF9750EE407C1" authorityName="Aitken" authorityYear="1972" box="[364,428,1668,1701]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="gilesi">gilesi</taxonomicName>
, which is morphologically distinct from other planigales in possessing only two upper and lower premolar teeth. Various genetic studies have been conducted on the genus
<taxonomicName id="A5D94D54FF82246FFDA1F9220D330790" box="[514,635,1747,1780]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Planigale</taxonomicName>
since the early 1980s. Initially, the genus was placed in its own subfamily, because it shared no morphological features allying it clearly with other dasyurids.
<taxonomicName id="A5D94D54FF82246FFE2FF8D30D4E0627" box="[396,518,1826,1859]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Planigale</taxonomicName>
was later incorporated in a distinct tribe, Planigalini, within the subfamily
<taxonomicName id="A5D94D54FF82246FFEC6F8B80D72060E" authorityName="Archer" authorityYear="1982" box="[357,570,1865,1898]" class="Mammalia" family="Dasyuridae" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" rank="subFamily" subFamily="Sminthopsinae">Sminthopsinae</taxonomicName>
, along with three other dasyurid genera (
<taxonomicName id="A5D94D54FF82246FFBDDF8B80A51060E" box="[1150,1305,1865,1898]" class="Mammalia" family="Dasyuridae" genus="Sminthopsis" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Sminthopsis</taxonomicName>
,
<taxonomicName id="A5D94D54FF82246FFA8BF8B80E6206F6" authorityName="Krefft" authorityYear="1867" class="Mammalia" family="Dasyuridae" genus="Antechinomys" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Antechinomys</taxonomicName>
, and
<taxonomicName id="A5D94D54FF82246FFEDEF8800EA006F6" box="[381,488,1905,1938]" class="Mammalia" family="Dasyuridae" genus="Ningaui" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Ningaui</taxonomicName>
) based on genetics and serology. More recent DNA sequencing has corroborated the close genetic relationship of these four genera. Monophyly of
<taxonomicName id="A5D94D54FF82246FFF6AF84E0E0A0684" box="[201,322,1983,2016]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="genus">Planigale</taxonomicName>
has been well established genetically, which corroborates morphological uniqueness of the group. Using mtDNA, one study resolved
<taxonomicName id="A5D94D54FF82246FFC49F8160B27096C" authorityName="Thomas" authorityYear="1906" box="[1002,1135,2023,2056]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
as clearly monophyletic and positioned it as a poorly resolved sister to a clade containing
<taxonomicName id="A5D94D54FF82246FFB46F7FF0F9C0933" authorityName="Kerr" authorityYear="1792" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="maculata">P. maculata</taxonomicName>
and P. tenwirostris. Interestingly,
<taxonomicName id="A5D94D54FF82246FFD38F7C70C6A0933" authorityName="Thomas" authorityYear="1906" box="[667,802,2102,2135]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
and
<taxonomicName id="A5D94D54FF82246FFCC9F7C70B500933" authorityName="Troughton" authorityYear="1928" box="[874,1048,2102,2135]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="tenuirostris">P. tenuirostris</taxonomicName>
in South Australia are broadly sympatric with
<taxonomicName id="A5D94D54FF82246FFE44F7AC0D0E091A" authorityName="Aitken" authorityYear="1972" box="[487,582,2141,2174]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="gilesi">P. gilesi</taxonomicName>
but not with each other. There appears to be ecological habitat separation between all three species, with
<taxonomicName id="A5D94D54FF82246FFCC0F7740CA209C2" authorityName="Thomas" authorityYear="1906" box="[867,1002,2181,2214]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
being the most restricted in distribution and habitat requirements. Three forms of P. ingram: have been recognized in the past but are no longer formally considered subspecies:
<taxonomicName id="A5D94D54FF82246FFBCCF7220E140878" authorityName="Troughton" authorityYear="1928" class="Mammalia" family="Dasyuridae" form="brunnea" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="form" species="ingrami">Planigale ingrami form brunnea</taxonomicName>
from the Richmond area;
<taxonomicName id="A5D94D54FF82246FFD75F70A0B520878" authority="form ingrami" box="[726,1050,2299,2332]" class="Mammalia" family="Dasyuridae" form="ingrami" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="form" species="ingrami">P. ingrami form ingrami</taxonomicName>
found from Northern Territory to Townsville;
<taxonomicName id="A5D94D54FF82246FFE55F6D30C270827" baseAuthorityName="Lonnberg" baseAuthorityYear="1913" box="[502,879,2338,2371]" class="Mammalia" family="Dasyuridae" form="subtilissima" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="form" species="ingrami">P. ingrami form subtilissima</taxonomicName>
found in Kimberley. It is possible that
<taxonomicName id="A5D94D54FF82246FFF42F6BB0E2E080F" baseAuthorityName="Thomas" baseAuthorityYear="1906" box="[225,358,2378,2411]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="ingrami">P. ingrami</taxonomicName>
may also be genetically heterogeneous across this part ofits distribution or these forms may represent a south-east to a north-west cline. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFF02F6690C5A0885" pageId="43" pageNumber="328" type="distribution">
<caption id="36A6665FFF82246FFF02F6690C5A0885" ID-DOI="http://doi.org/10.5281/zenodo.6608250" ID-Zenodo-Dep="6608250" httpUri="https://zenodo.org/record/6608250/files/figure.png" inLine="true" pageId="43" pageNumber="328" targetBox="[159,751,1165,1580]" targetPageId="43">
<paragraph id="626636D7FF82246FFF02F6690C5A0885" blockId="43.[160,1368,1589,3474]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF02F6690E1908DD" bold="true" box="[161,337,2456,2489]" pageId="43" pageNumber="328">Distribution.</emphasis>
Mainly across N Australia, in NE Western Australia, Northern Territory, and Queensland, also in NE South Australia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFF02F61D0C2F0A78" pageId="43" pageNumber="328" type="description">
<paragraph id="626636D7FF82246FFF02F61D0C2F0A78" blockId="43.[160,1368,1589,3474]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF02F61D0ED40B6D" bold="true" box="[161,412,2540,2569]" pageId="43" pageNumber="328">Descriptive notes.</emphasis>
Head—body 5.7-1 cm (males) and 4.9-6.4 cm (females), tail 5-3— 6-8 cm (males) and 5.1-6.5 cm (females); weight 2:8-6-6 g (males) and 2:6-5-8 g (females). There is mild sexual dimorphism for size. The Long-tailed Planigale is the worlds smallest marsupial and one of the smallest mammals. Extremely flattened, wedge-shaped head and laterally inclined hindlimbs allow it to squeeze through very small spaces. Long-tailed Planigales have a permanently thin tail, usually slightly longer than head-body length (although proportionately shorter with increasing body size). It has three upper and lower premolar teeth.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFF02F4D30EA50C26" pageId="43" pageNumber="328" type="biology_ecology">
<paragraph id="626636D7FF82246FFF02F4D30EA50C26" blockId="43.[160,1368,1589,3474]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF02F4D30E590A27" bold="true" box="[161,273,2850,2883]" pageId="43" pageNumber="328">Habitat.</emphasis>
Narrow crevices in cracking clays of black-soil plains. These plains, which support tussock grasslands or sparse woodlands, are found throughout the drier interior of tropical northern Australia and occur as floodplains associated with major river systems nearer the coast. Surveys incorporating pitfall traps have shown Longtailed Planigales to be the most frequently encountered small mammalin its favored habitat. One study focusing on South Australian populations found that the Longtailed Planigale is restricted to the far north-east of the state, having most affinity with plains and flooded areas. In South Australia, vegetation communities in which Long-tailed Planigale has been recorded are typical of arid floodplain environments: dwarf chenopod shrubs, herbs, and grasses with taller lignum (
<taxonomicName id="A5D94D54FF82246FFBE1F3740A4B0DC2" box="[1090,1283,3205,3238]" class="Magnoliopsida" family="Polygonaceae" genus="Muehlenbeckia" kingdom="Plantae" order="Caryophyllales" pageId="43" pageNumber="328" phylum="Tracheophyta" rank="genus">Muehlenbeckia</taxonomicName>
florulenta,
<taxonomicName id="A5D94D54FF82246FFF50F35D0EFD0DA9" box="[243,437,3244,3277]" class="Magnoliopsida" family="Polygonaceae" kingdom="Plantae" order="Caryophyllales" pageId="43" pageNumber="328" phylum="Tracheophyta" rank="family">Polygonaceae</taxonomicName>
) or Queensland bluebush (
<taxonomicName id="A5D94D54FF82246FFCEBF35D0CB40DA9" box="[840,1020,3244,3277]" class="Magnoliopsida" family="Amaranthaceae" genus="Chenopodium" kingdom="Plantae" order="Caryophyllales" pageId="43" pageNumber="328" phylum="Tracheophyta" rank="genus">Chenopodium</taxonomicName>
auricomum, Chenopodioideae). Shrubs characterize the more regularly flooded areas, and these support an open overstory of coolibah (
<taxonomicName id="A5D94D54FF82246FFDE3F30A0D870C78" box="[576,719,3323,3356]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="43" pageNumber="328" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
coolabahs,
<taxonomicName id="A5D94D54FF82246FFCC9F30A0CB30C78" box="[874,1019,3323,3356]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="43" pageNumber="328" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) woodland in the most regularly flooded areas.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFF03F2BB09C400D3" pageId="43" pageNumber="328" type="food_feeding">
<paragraph id="626636D7FF82246FFF03F2BB09C400D3" blockId="43.[160,1368,1589,3474]" lastBlockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFF03F2BB0EE30C0F" bold="true" box="[160,427,3402,3435]" pageId="43" pageNumber="328">Food and Feeding.</emphasis>
Despite its tiny size, the Long-tailed Planigale is a ferocious nocturnal predator, seeking prey through soil cracks and hunting on the surface. Large centipedes, often several times the length of the Long-tailed Planigale, are a favorite prey item. Any other creature small enough to be subdued is also viciously attacked, including spiders, grasshoppers, other insects and their larvae, small lizards, and even young mammals.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFA3DFE4F085A0563" pageId="43" pageNumber="328" type="breeding">
<paragraph id="626636D7FF82246FFA3DFE4F085A0563" blockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFA3DFE4F096D00BB" bold="true" box="[1438,1573,446,479]" pageId="43" pageNumber="328">Breeding.</emphasis>
Female Long-tailed Planigales have a well-developed, rear-opening pouch with 8-12 teats and have litters of 4-8 young. Pouch morphology of Long-tailed Planigales from Kimberley, with nipples located in two anterior pockets, is unique among carnivorous marsupials. Females with pouch young have been seen mostly in September-December, although births can occur throughout the year. Young become detached from teats at c.6 weeks old and are left in a grassy nest underlitter or within tussock grass clumps while the mother forages. Independent, immature Long-tailed Planigales generally appear in the population in December—April. In a recent study on reproduction in the Long-tailed Planigale based on individuals extracted from stomachs of feral cats shot in north-western Queensland, the extended breeding season commenced in August and most likely ended in December. Evidently, males and females live to breed in more than one season. Females examined had twelve nipples in the pouch. Pouch morphology was noted to differ from that of other species of planigales (and may be a useful diagnostic if pouch condition is closely monitored in field-captured specimens).
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFA3FFBFF08370519" pageId="43" pageNumber="328" type="activity">
<paragraph id="626636D7FF82246FFA3FFBFF08370519" blockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFA3FFBFF09C3054B" bold="true" box="[1436,1675,1038,1071]" pageId="43" pageNumber="328">Activity patterns.</emphasis>
Although mostly nocturnal, Long-tailed Planigales may also be active in the early morning, particularly after cold nights. They may become completely torpid under very cold conditions.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFA3DFB75093A05A8" pageId="43" pageNumber="328" type="biology_ecology">
<paragraph id="626636D7FF82246FFA3DFB75093A05A8" blockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFA3DFB75072D05C1" bold="true" box="[1438,2149,1156,1189]" pageId="43" pageNumber="328">Movements, Home range and Social organization.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFA3DFB22057A074B" pageId="43" pageNumber="328" type="conservation">
<paragraph id="626636D7FF82246FFA3DFB22057A074B" blockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFA3DFB2209B10590" bold="true" box="[1438,1785,1235,1268]" pageId="43" pageNumber="328">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Longtailed Planigale was originally known only from northern Australia. Many (genetically confirmed) specimens have since been collected farther south into South Australia, and the distribution of the Long-tailed Planigale now appears to be almost as extensive as that of the Common Planigale (
<taxonomicName id="A5D94D54FF82246FF822FA80075704F6" authorityName="Kerr" authorityYear="1792" box="[1921,2079,1393,1426]" class="Mammalia" family="Dasyuridae" genus="Planigale" kingdom="Animalia" order="Dasyuromorphia" pageId="43" pageNumber="328" phylum="Chordata" rank="species" species="maculata">P. maculata</taxonomicName>
). Habitat of the Long-tailed Planigale in the south-eastern part of its distribution has been substantially impacted by conversion of native grasslands and woodlands to agriculture. The Long-tailed Planigale appears to persist under the dominant pastoral land use throughout the remainder ofits distribution, although densities notably decline in areas subjected to heavy grazing pressure.
</paragraph>
</subSubSection>
<subSubSection id="2AC3655CFF82246FFA3DF9CE07BA07C3" pageId="43" pageNumber="328" type="bibRefCitation_list">
<paragraph id="626636D7FF82246FFA3DF9CE07BA07C3" blockId="43.[1436,2646,289,1708]" pageId="43" pageNumber="328">
<emphasis id="50ADEAC5FF82246FFA3DF9CE097F073C" bold="true" box="[1438,1591,1599,1624]" pageId="43" pageNumber="328">Bibliography.</emphasis>
Archer (1976a, 1982c), Baverstock et al. (1982), Blacket, Adams et al. (2000), Blacket, Kemper &amp; Brandle (2008), Burnett (2008b), Fisher (2008), Krajewski, Blacket et al. (1997), Krajewski, Young et al. (1997), Painter et al. (1995), Thomas (1906), Troughton (1928), Woolley &amp; Elliott (2013).
</paragraph>
</subSubSection>
</treatment>
</document>