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<document id="A039F3959A68FEB986C1CFE973A25663" ID-CLB-Dataset="298453" ID-DOI="10.1186/s13071-017-1997-8" ID-GBIF-Dataset="c523c95d-a510-4ba0-993c-328152a75924" ID-ISSN="1756-3305" ID-PMC="PMC5296950" ID-PubMed="28173840" ID-Zenodo-Dep="11551872" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1718039930260" checkinUser="felipe" docAuthor="Ash, Amanda, Ellcot, Acleen, Godfres, Stephance, Burmej, Halcna, Abdad, Mohammad Yazcd, Northover, Ams, Wasne, Adrcan, Morrcs, Kecth, Clode, Peta, Lsmbers, Alan &amp; Andrew Thompson, R. C." docDate="2017" docId="EA58B915FF83483EFF1F2353B001F930" docLanguage="en" docName="ParasiteVector.10.70.pdf" docOrigin="Parasites &amp; Vectors (70) 10 (1)" docSource="http://dx.doi.org/10.1186/s13071-017-1997-8" docStyle="DocumentStyle:F140E6D4AE1E76E09EF03FD1AEF09E53.3:ParasiteVector.2015-.journal_article" docStyleId="F140E6D4AE1E76E09EF03FD1AEF09E53" docStyleName="ParasiteVector.2015-.journal_article" docStyleVersion="3" docTitle="Ixodes woyliei Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson 2017" docType="treatment" docVersion="5" lastPageNumber="8" masterDocId="1661C16DFF804836FF882063B131FFF5" masterDocTitle="Morphological and molecular description of Ixodes woyliei n. sp. (Ixodidae) with consideration for co-extinction with its critically endangered marsupial host" masterLastPageNumber="16" masterPageNumber="1" pageNumber="4" updateTime="1718321264324" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1354E826E1858511B225A6DA46B43ED4">Morphological and molecular description of Ixodes woyliei n. sp. (Ixodidae) with consideration for co-extinction with its critically endangered marsupial host</mods:title>
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<mods:namePart id="DB58ECE3A3A5F6C68A9CE662AA10B508">Ash, Amanda</mods:namePart>
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<mods:namePart id="8A16A7288C91C2141EDBCA19A0395457">Ellcot, Acleen</mods:namePart>
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<mods:namePart id="771AD8F68E88B3B03FDFBDE1E2794F3B">Godfres, Stephance</mods:namePart>
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<mods:namePart id="C62F19643174E360C88715FBD95714EE">Abdad, Mohammad Yazcd</mods:namePart>
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<mods:namePart id="26962C361EF0AEBE5F2356BD72FB6A01">Northover, Ams</mods:namePart>
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<mods:namePart id="4116394A4C492F70ADD5AC0548C6F6E7">Wasne, Adrcan</mods:namePart>
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<mods:namePart id="39E4F1CCBD1383667AC7691DF8826A85">Morrcs, Kecth</mods:namePart>
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<mods:namePart id="B5282D58DE194F5E5B29E4DBC2FAE4E6">Clode, Peta</mods:namePart>
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<mods:namePart id="E20DCC9EC3C15AA41D5F9CED0E845490">Lsmbers, Alan</mods:namePart>
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<mods:namePart id="76D4391241698FCB343CC620527C88E4">Andrew Thompson, R. C.</mods:namePart>
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<mods:title id="1DE5019159D0E266A32BC9ACEBCD7970">Parasites &amp; Vectors</mods:title>
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<mods:date id="44322D1E34A44595208FA4A78F2F59BB">2017</mods:date>
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<treatment id="EA58B915FF83483EFF1F2353B001F930" ID-DOI="http://doi.org/10.5281/zenodo.11645807" ID-GBIF-Taxon="232750345" ID-Zenodo-Dep="11645807" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930" httpUri="http://treatment.plazi.org/id/EA58B915FF83483EFF1F2353B001F930" lastPageId="8" lastPageNumber="8" pageId="3" pageNumber="4">
<subSubSection id="2AEB5B88FF834835FF1F2353B052FCB3" box="[151,355,816,839]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="624E0803FF834835FF1F2353B052FCB3" blockId="3.[151,355,816,839]" box="[151,355,816,839]" pageId="3" pageNumber="4">
<heading id="3906BF6FFF834835FF1F2353B052FCB3" bold="true" box="[151,355,816,839]" fontSize="9" level="2" pageId="3" pageNumber="4" reason="7">
<emphasis id="5085D411FF834835FF1F2353B052FCB3" bold="true" box="[151,355,816,839]" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F2353B016FCB2" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[151,295,816,839]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="woyliei">
<emphasis id="5085D411FF834835FF1F2353B016FCB2" bold="true" box="[151,295,816,839]" italics="true" pageId="3" pageNumber="4">Ixodes woyliei</emphasis>
</taxonomicName>
n. sp
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AEB5B88FF834835FF1F230CB0EFFC52" pageId="3" pageNumber="4" type="type_taxon">
<paragraph id="624E0803FF834835FF1F230CB0EFFC52" blockId="3.[151,775,879,1447]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F230CB034FC73" bold="true" box="[151,261,879,902]" italics="true" pageId="3" pageNumber="4">Type-host</emphasis>
:
<taxonomicName id="A5F17380FF834835FE9D230CB346FC72" authority="Gray, 1837" authorityName="Gray" authorityYear="1837" box="[277,631,879,903]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="5085D411FF834835FE9D230CB0C6FC72" box="[277,503,879,903]" italics="true" pageId="3" pageNumber="4">Bettongia penicillata</emphasis>
Gray, 1837
</taxonomicName>
(
<taxonomicName id="A5F17380FF834835FD0F230CB3CEFC72" authorityName="Gray" authorityYear="1821" box="[647,767,879,903]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
) (woylie, brush-tailed bettong).
</paragraph>
</subSubSection>
<subSubSection id="2AEB5B88FF834835FF1F23CCB46EFBBF" pageId="3" pageNumber="4" type="description">
<paragraph id="624E0803FF834835FF1F23CCB3C2FBF2" blockId="3.[151,775,879,1447]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F23CCB010FC33" bold="true" box="[151,289,943,966]" italics="true" pageId="3" pageNumber="4">Other hosts</emphasis>
:
<taxonomicName id="A5F17380FF834835FEBF23CCB236FC32" authority="Shaw, 1797" authorityName="Shaw" authorityYear="1797" box="[311,775,943,967]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="subSpecies" species="obesulus" subSpecies="fusciventer">
<emphasis id="5085D411FF834835FEBF23CCB347FC32" box="[311,630,943,967]" italics="true" pageId="3" pageNumber="4">Isoodon obesulus fusciventer</emphasis>
Shaw, 1797
</taxonomicName>
(
<taxonomicName id="A5F17380FF834835FF1623ACB012FC12" authorityName="Gray" authorityYear="1825" box="[158,291,975,999]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
) (quenda, southern brown bandicoot) and
<taxonomicName id="A5F17380FF834835FF1F238CB086FBF2" authority="Reid, 1837" authorityName="Reid" authorityYear="1837" box="[151,439,1007,1031]" class="Mammalia" family="Thylacomyidae" genus="Macrotis" kingdom="Animalia" order="Peramelemorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="lagotis">
<emphasis id="5085D411FF834835FF1F238CB071FBF2" box="[151,320,1007,1031]" italics="true" pageId="3" pageNumber="4">Macrotis lagotis</emphasis>
Reid, 1837
</taxonomicName>
(
<taxonomicName id="A5F17380FF834835FE4C238CB378FBF2" authorityName="Gray" authorityYear="1825" box="[452,585,1007,1031]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
) (greater bilby).
</paragraph>
<paragraph id="624E0803FF834835FF1F246CB0F0FBB2" blockId="3.[151,775,879,1447]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F246CB01DFBD3" bold="true" box="[151,300,1039,1062]" italics="true" pageId="3" pageNumber="4">Type-locality</emphasis>
:
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<location id="672E5ED8FF834835FECB246CB317FBD2" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FECB246CB317FBD2" box="[323,550,1039,1063]" country="Australia" latitude="-32.783333" longLatPrecision="1209" longitude="116.96667" name="Dryandra Woodland" pageId="3" pageNumber="4" stateProvince="Western Australia">Dryandra Woodland</location>
,
<collectingRegion id="A035C6E1FF834835FDB2246CB232FBD2" box="[570,771,1039,1063]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FF1F244CB1C4FBB2" box="[151,245,1071,1095]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
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,
<geoCoordinate id="07C56EC4FF834835FED2244CB087FBB2" box="[346,438,1071,1095]" degrees="116" direction="east" minutes="58" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.96667">116°58'E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="624E0803FF834835FF1F242CB0B0FB32" blockId="3.[151,775,879,1447]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F242CB079FB93" bold="true" box="[151,328,1103,1126]" italics="true" pageId="3" pageNumber="4">Other localities</emphasis>
:
<materialsCitation id="D299025EFF834835FEDE242CB04FFB32" ID-GBIF-Occurrence="4876545304" country="Australia" latitude="-34.36139" location="Karacamia Wildlife Sanctuary" longLatPrecision="19" longitude="116.306114" municipality="Upper Warren Region" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Western Australia">
<location id="672E5ED8FF834835FEDE242CB3AFFB92" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FEDE242CB3AFFB92" box="[342,670,1103,1127]" country="Australia" latitude="-34.36139" longLatPrecision="19" longitude="116.306114" municipality="Upper Warren Region" name="Karacamia Wildlife Sanctuary" pageId="3" pageNumber="4" stateProvince="Western Australia">Karacamia Wildlife Sanctuary</location>
,
<collectingRegion id="A035C6E1FF834835FD22242CB1CDFB72" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FE86240CB041FB72" box="[270,368,1135,1159]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FE0E240CB0E7FB72" box="[390,470,1135,1159]" degrees="31" direction="south" minutes="48" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-31.8">31°48'S</geoCoordinate>
,
<geoCoordinate id="07C56EC4FF834835FE61240CB374FB72" box="[489,581,1135,1159]" degrees="116" direction="east" minutes="15" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.25">116°15'E</geoCoordinate>
) and the
<collectingMunicipality id="822A9279FF834835FD4A240CB061FB52" pageId="3" pageNumber="4">Upper Warren Region</collectingMunicipality>
, Western Australia, Australia (34° 21'41&quot;S, 116°18'22&quot;E)
</materialsCitation>
.
</paragraph>
<paragraph id="624E0803FF834835FF1F24ACB432FCD0" blockId="3.[151,775,879,1447]" lastBlockId="3.[813,1437,237,1290]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F24ACB07BFB13" bold="true" box="[151,330,1231,1254]" italics="true" pageId="3" pageNumber="4">Type-specimens</emphasis>
:
<materialsCitation id="D299025EFF834835FEED24ACB31DFAB2" ID-GBIF-Occurrence="4876545307" collectingDate="2015-12" collectionCode="WAM" country="Australia" latitude="-32.783333" location="West Australian Museum" longLatPrecision="1209" longitude="116.96667" pageId="3" pageNumber="4" specimenCode="T142602" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="BD4AB6A1FF834835FEED24ACB0FBFB12" box="[357,458,1231,1255]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
: female ex
<taxonomicName id="A5F17380FF834835FDEF24ACB04FFAF2" authority=", Dryandra Woodland" authorityName="Dryandra Woodland" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="5085D411FF834835FDEF24ACB230FB12" box="[615,769,1231,1255]" italics="true" pageId="3" pageNumber="4">B. penicillata</emphasis>
, Dryandra Woodland
</taxonomicName>
,
<collectingRegion id="A035C6E1FF834835FE18248CB36DFAF2" box="[400,604,1263,1287]" inferredBySuffix="true" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FDE7248CB3FEFAF2" box="[623,719,1263,1287]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FD6D248CB1F2FAD2" degrees="32" direction="south" minutes="47" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-32.783333">32° 47'S</geoCoordinate>
<geoCoordinate id="07C56EC4FF834835FF59256CB01FFAD2" box="[209,302,1295,1319]" degrees="116" direction="east" minutes="58" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.96667">116°58'E</geoCoordinate>
), December 2015, deposited at the
<location id="672E5ED8FF834835FD47256CB058FAB2" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FD47256CB058FAB2" country="Australia" latitude="-32.783333" longLatPrecision="1209" longitude="116.96667" name="West Australian Museum" pageId="3" pageNumber="4" stateProvince="Western Australia">West Australian Museum</location>
(
<collectionCode id="04E090C6FF834835FEF0254CB08FFAB2" box="[376,446,1327,1351]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="4" type="Museum">WAM</collectionCode>
<specimenCode id="3257A078FF834835FE4D254CB314FAB2" box="[453,549,1327,1351]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T142602</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="D299025EFF834835FDBF254CB24AFEF0" ID-GBIF-Occurrence="4876545306" collectingDate="2016-06" country="Australia" latitude="-32.783333" location="Total" longLatPrecision="1209" longitude="116.96667" municipality="Dryandra Woodland" pageId="3" pageNumber="4" specimenCount="19" specimenCount-female="19" stateProvince="Western Australia" typeStatus="paratype">
<typeStatus id="BD4AB6A1FF834835FDBF254CB3ADFAB2" box="[567,668,1327,1351]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
:
<location id="672E5ED8FF834835FD20254CB3D0FAB2" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FD20254CB3D0FAB2" box="[680,737,1327,1351]" country="Australia" latitude="-32.783333" longLatPrecision="1209" longitude="116.96667" municipality="Dryandra Woodland" name="Total" pageId="3" pageNumber="4" stateProvince="Western Australia">Total</location>
25,
<specimenCount id="74F7C38AFF834835FF1F252CB039FA92" box="[151,264,1359,1383]" count="11" pageId="3" pageNumber="4" type="female">11 females</specimenCount>
and 14 nymphs ex
<emphasis id="5085D411FF834835FE56252CB340FA92" box="[478,625,1359,1383]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FE56252CB35FFA92" box="[478,622,1359,1383]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">B. penicillata</taxonomicName>
.
</emphasis>
<specimenCount id="74F7C38AFF834835FDF0252CB236FA92" box="[632,775,1359,1383]" count="8" pageId="3" pageNumber="4" type="female">Eight females</specimenCount>
(P1
<emphasis id="5085D411FF834835FF32250CB1F8FA72" box="[186,201,1391,1415]" italics="true" pageId="3" pageNumber="4"></emphasis>
P8) collected from
<collectingMunicipality id="822A9279FF834835FE3A250CB3A6FA72" box="[434,663,1391,1415]" pageId="3" pageNumber="4">Dryandra Woodland</collectingMunicipality>
,
<collectingRegion id="A035C6E1FF834835FD24250CB1C9FA52" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FE8D25ECB055FA52" box="[261,356,1423,1447]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FEFD25ECB0F3FA52" box="[373,450,1423,1447]" degrees="32" direction="south" minutes="47" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-32.783333">32°47'S</geoCoordinate>
,
<geoCoordinate id="07C56EC4FF834835FE4725ECB31AFA52" box="[463,555,1423,1447]" degrees="116" direction="east" minutes="58" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.96667">116°58'E</geoCoordinate>
), June 2016 (P1, P2, P7, P8)
</materialsCitation>
,
<materialsCitation id="D299025EFF834835FC00208EB5A2FEF0" ID-GBIF-Occurrence="4876545305" box="[904,1171,237,261]" collectingDate="2015-09" country="Australia" isTailEnumeration="true" latitude="-32.783333" location="Total" longLatPrecision="1209" longitude="116.96667" municipality="Dryandra Woodland" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<date id="164F2EC3FF834835FC00208EB505FEF0" box="[904,1076,237,261]" pageId="3" pageNumber="4" value="2015-09">
<collectingDate id="060BD72BFF834835FC00208EB505FEF0" box="[904,1076,237,261]" pageId="3" pageNumber="4" value="2015-09">September 2015</collectingDate>
</date>
(P3, P4)
</materialsCitation>
,
<date id="164F2EC3FF834835FB28208EB408FEF0" box="[1184,1337,237,261]" pageId="3" pageNumber="4" value="2015-02">February 2015</date>
(P5, P6);
<materialsCitation id="D299025EFF834835FCA5216EB2A3FE90" ID-GBIF-Occurrence="4876545301" collectingDate="2006-07" country="Australia" latitude="-31.8" location="Karacamia Wildlife Sanctuary" longLatPrecision="1214" longitude="116.25" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="2" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount id="74F7C38AFF834835FCA5216EB2A6FED0" box="[813,919,269,293]" count="2" pageId="3" pageNumber="4" type="female">2 females</specimenCount>
(P9, P10) collected from
<location id="672E5ED8FF834835FB41216EB2A7FEB0" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FB41216EB2A7FEB0" country="Australia" latitude="-31.8" longLatPrecision="1214" longitude="116.25" name="Karacamia Wildlife Sanctuary" pageId="3" pageNumber="4" stateProvince="Western Australia">Karacamia Wildlife Sanctuary</location>
,
<collectingRegion id="A035C6E1FF834835FC2A214EB557FEB0" box="[930,1126,301,325]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FBFA214EB5E0FEB0" box="[1138,1233,301,325]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FB69214EB41FFEB0" box="[1249,1326,301,325]" degrees="31" direction="south" minutes="48" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-31.8">31°48'S</geoCoordinate>
,
<geoCoordinate id="07C56EC4FF834835FAB3214EB4A4FEB0" box="[1339,1429,301,325]" degrees="116" direction="east" minutes="15" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.25">116°15'E</geoCoordinate>
)
<date id="164F2EC3FF834835FCA5212EB2A3FE90" box="[813,914,333,357]" pageId="3" pageNumber="4" value="2006-07">
<collectingDate id="060BD72BFF834835FCA5212EB2A3FE90" box="[813,914,333,357]" pageId="3" pageNumber="4" value="2006-07">July 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D299025EFF834835FC16212EB4A9FE10" ID-GBIF-Occurrence="4876545303" collectorName="Seven" country="Australia" latitude="-34.36139" location="Upper Warren" longLatPrecision="19" longitude="116.306114" municipality="Upper Warren Region" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
and
<specimenCount id="74F7C38AFF834835FC46212EB572FE90" box="[974,1091,333,357]" count="1" pageId="3" pageNumber="4" type="female">one female</specimenCount>
(P11) collected from the
<location id="672E5ED8FF834835FAD2212EB24EFE70" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FAD2212EB24EFE70" country="Australia" latitude="-34.36139" longLatPrecision="19" longitude="116.306114" municipality="Upper Warren Region" name="Upper Warren" pageId="3" pageNumber="4" stateProvince="Western Australia">Upper Warren</location>
<collectingRegion id="A035C6E1FF834835FC02210EB596FE70" box="[906,1191,365,389]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Region Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FB3F210EB427FE70" box="[1207,1302,365,389]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FAA1210EB4A9FE70" box="[1321,1432,365,389]" degrees="34" direction="south" minutes="21" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="41" value="-34.36139">34°21'41&quot;S</geoCoordinate>
,
<geoCoordinate id="07C56EC4FF834835FCA521EEB29AFE50" box="[813,939,397,421]" degrees="116" direction="east" minutes="18" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="22" value="116.306114">116°18'22&quot;E</geoCoordinate>
).
<collectorName id="CF046DD5FF834835FC4E21EEB535FE50" box="[966,1028,397,421]" pageId="3" pageNumber="4">Seven</collectorName>
nymphs (P12
<emphasis id="5085D411FF834835FB2121EFB589FE51" box="[1193,1208,396,420]" italics="true" pageId="3" pageNumber="4"></emphasis>
P18) collected from
<collectingMunicipality id="822A9279FF834835FCA521CEB515FE30" box="[813,1060,429,453]" pageId="3" pageNumber="4">Upper Warren Region</collectingMunicipality>
,
<collectingRegion id="A035C6E1FF834835FBBB21CEB5CBFE30" box="[1075,1274,429,453]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FA8121CEB459FE30" box="[1289,1384,429,453]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(34° 21'41&quot;S, 116°18'22&quot;E) September 2014 and
<date id="164F2EC3FF834835FB7821AEB4A9FE10" box="[1264,1432,461,485]" pageId="3" pageNumber="4" value="2014-12">December 2014</date>
</materialsCitation>
,
<materialsCitation id="D299025EFF834835FCA5218EB504FDB1" ID-GBIF-Occurrence="4876545308" country="Australia" latitude="-32.783333" location="Dryandra Woodland" longLatPrecision="1209" longitude="116.96667" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
4 nymphs (P19
<emphasis id="5085D411FF834835FC47218FB2EFFDF1" box="[975,990,492,516]" italics="true" pageId="3" pageNumber="4"></emphasis>
P22) collected from
<location id="672E5ED8FF834835FB33218EB4A9FDF0" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FB33218EB4A9FDF0" box="[1211,1432,493,517]" country="Australia" latitude="-32.783333" longLatPrecision="1209" longitude="116.96667" name="Dryandra Woodland" pageId="3" pageNumber="4" stateProvince="Western Australia">Dryandra Woodland</location>
,
<collectingRegion id="A035C6E1FF834835FCA5226EB2DFFDD0" box="[813,1006,525,549]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
,
<collectingCountry id="1AE64893FF834835FC71226EB569FDD0" box="[1017,1112,525,549]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
(
<geoCoordinate id="07C56EC4FF834835FBEE226EB582FDD0" box="[1126,1203,525,549]" degrees="32" direction="south" minutes="47" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-32.783333">32°47'S</geoCoordinate>
,
<geoCoordinate id="07C56EC4FF834835FB36226EB42BFDD0" box="[1214,1306,525,549]" degrees="116" direction="east" minutes="58" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="116.96667">116°58'E</geoCoordinate>
),
<date id="164F2EC3FF834835FAA3226EB251FDB1" pageId="3" pageNumber="4" value="1994-09">September 1994</date>
and October 1993
</materialsCitation>
,
<materialsCitation id="D299025EFF834835FBCC224FB5CEFCD0" ID-GBIF-Occurrence="4876545309" collectionCode="WAM" country="Australia" location="Australian National Insect Collection" municipality="Western Australian Museum" pageId="3" pageNumber="4" specimenCode="ANIC 48-006275, ANIC 48-006277, T142603, T142604" specimenCount="12" stateProvince="Western Australia" typeStatus="paratype">
one (P23) collected from Perth Zoo and two (P23
<emphasis id="5085D411FF834835FC72222FB538FD91" box="[1018,1033,588,612]" italics="true" pageId="3" pageNumber="4"></emphasis>
25) collected from Kanyana Wildlife Rehabilitation Centre, Perth.
<specimenCount id="74F7C38AFF834835FBF0220EB4ADFD70" box="[1144,1436,621,645]" count="7" pageId="3" pageNumber="4" type="generic" typeStatus="paratype">Seven paratype specimens</specimenCount>
(P1, P2, P7, P8, P15
<emphasis id="5085D411FF834835FB9022EFB516FD51" box="[1048,1063,652,676]" italics="true" pageId="3" pageNumber="4"></emphasis>
P17) have been deposited at the
<location id="672E5ED8FF834835FCA522CEB58BFD30" LSID="urn:lsid:plazi:treatment:EA58B915FF83483EFF1F2353B001F930:672E5ED8FF834835FCA522CEB58BFD30" box="[813,1210,685,709]" country="Australia" municipality="Western Australian Museum" name="Australian National Insect Collection" pageId="3" pageNumber="4" stateProvince="Western Australia">Australian National Insect Collection</location>
(
<specimenCode id="3257A078FF834835FB4322CEB4B2FD30" box="[1227,1411,685,709]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="3" pageNumber="4">ANIC 48-006275</specimenCode>
<emphasis id="5085D411FF834835FA0422CFB4ADFD31" box="[1420,1436,684,708]" italics="true" pageId="3" pageNumber="4"></emphasis>
<specimenCode id="3257A078FF834835FCA522AEB2D9FD10" box="[813,1000,717,741]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="3" pageNumber="4">ANIC 48-006277</specimenCode>
) and
<specimenCount id="74F7C38AFF834835FBA422AEB41EFD10" box="[1068,1327,717,741]" count="5" pageId="3" pageNumber="4" type="generic" typeStatus="paratype">five paratype specimens</specimenCount>
(P9, P10, P12
<emphasis id="5085D411FF834835FCDC228FB252FCF1" box="[852,867,748,772]" italics="true" pageId="3" pageNumber="4"></emphasis>
P14) have been deposited at the
<collectingMunicipality id="822A9279FF834835FB43228FB2BAFCD0" pageId="3" pageNumber="4">Western Australian Museum</collectingMunicipality>
(
<collectionCode id="04E090C6FF834835FC11236EB2EEFCD0" box="[921,991,781,805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="4" type="Museum">WAM</collectionCode>
<specimenCode id="3257A078FF834835FC6D236EB572FCD0" box="[997,1091,781,805]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T142603</specimenCode>
-
<collectionCode id="04E090C6FF834835FBC2236EB5A1FCD0" box="[1098,1168,781,805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="4" type="Museum">WAM</collectionCode>
<specimenCode id="3257A078FF834835FB1F236EB5C6FCD0" box="[1175,1271,781,805]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T142604</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="624E0803FF834835FCA5234CB5BEFC72" blockId="3.[813,1437,237,1290]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FCA5234CB591FCB3" bold="true" box="[813,1184,815,838]" italics="true" pageId="3" pageNumber="4">Representative DNA sequences</emphasis>
: Mitochondrial cytochrome
<emphasis id="5085D411FF834835FC03232CB2A7FC93" box="[907,918,847,870]" italics="true" pageId="3" pageNumber="4">c</emphasis>
oxidase subunit 1 (
<emphasis id="5085D411FF834835FBF4232CB5AEFC93" box="[1148,1183,847,870]" italics="true" pageId="3" pageNumber="4">cox</emphasis>
1): GenBanc accession numbers KX673875
<emphasis id="5085D411FF834835FB8E230CB527FC72" box="[1030,1046,879,903]" italics="true" pageId="3" pageNumber="4"></emphasis>
KX673881.
</paragraph>
<paragraph id="624E0803FF834835FCA523F1B46EFBBF" blockId="3.[813,1437,237,1290]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FCA523F1B516FC5C" bold="true" box="[813,1063,914,937]" italics="true" pageId="3" pageNumber="4">ZooBank registration</emphasis>
: Details of
<taxonomicName id="A5F17380FF834835FB3C23F2B461FC5C" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1204,1360,913,937]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF834835FB3C23F2B461FC5C" box="[1204,1360,913,937]" italics="true" pageId="3" pageNumber="4">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF834835FAD523F1B4ACFC5F" box="[1373,1437,914,938]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
have been submitted to ZooBanc and the Life Science Identifier (
<collectionCode id="04E090C6FF834835FC2B23B1B2EFFC1F" box="[931,990,978,1002]" pageId="3" pageNumber="4">LSID</collectionCode>
) for the article is urn:lsid:zoobanc.org:- pub:
<uuid id="165732D6FF834835FCD12391B40DFBFF" box="[857,1340,1010,1034]" pageId="3" pageNumber="4">1DB97319-FF74-4380-9D49-988BA3467342</uuid>
and
<collectionCode id="04E090C6FF834835FCA52471B254FBDF" box="[813,869,1042,1066]" pageId="3" pageNumber="4">LSID</collectionCode>
for the new name is
<uri id="16600401FF834835FB4B2471B46BFBBF" pageId="3" pageNumber="4">
urn:lsid:zoobanc.org:act:
<uuid id="165732D6FF834835FCE82451B46BFBBF" box="[864,1370,1074,1098]" pageId="3" pageNumber="4">7FC91916-ACBC-4832-8788-19CCBB8EF7B1</uuid>
</uri>
.
</paragraph>
</subSubSection>
<subSubSection id="2AEB5B88FF834835FCA52431B544FAFF" pageId="3" pageNumber="4" type="etymology">
<paragraph id="624E0803FF834835FCA52431B544FAFF" blockId="3.[813,1437,237,1290]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FCA52431B295FB9C" bold="true" box="[813,932,1106,1129]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
: The species name
<taxonomicName id="A5F17380FF834835FB1A2432B41FFB9C" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1170,1326,1105,1129]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="woyliei">
<emphasis id="5085D411FF834835FB1A2432B41FFB9C" box="[1170,1326,1105,1129]" italics="true" pageId="3" pageNumber="4">Ixodes woyliei</emphasis>
</taxonomicName>
refers to the common name of the host
<taxonomicName id="A5F17380FF834835FB0E2411B426FB7C" box="[1158,1303,1137,1161]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="5085D411FF834835FB0E2411B426FB7C" box="[1158,1303,1137,1161]" italics="true" pageId="3" pageNumber="4">B. penicillata</emphasis>
</taxonomicName>
(commonly cnown as the woylie) for which this ticc appears to have a high predilection. Woylie is the Aboriginal name given by the Noongar people who live in the south-west corner of
<collectingRegion id="A035C6E1FF834835FCFE2491B50DFAFF" box="[886,1084,1266,1290]" country="Australia" name="Western Australia" pageId="3" pageNumber="4">Western Australia</collectingRegion>
[
<bibRefCitation id="066075F2FF834835FBC32491B55BFAFF" author="Abbott I." box="[1099,1130,1266,1290]" pageId="3" pageNumber="4" pagination="433 - 86" refId="ref9958" refString="23. Abbott I. Aborcgcnal names of mammal specces cn south-west Western Australca. CALMsccence. 2001; 3 (4): 433 - 86." type="journal article" year="2001">23</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="2AEB5B88FF834830FCA52533B474F851" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="624E0803FF834835FCA52533B296FA93" blockId="3.[813,1436,1360,1447]" box="[813,935,1360,1382]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FCA52533B296FA93" bold="true" box="[813,935,1360,1382]" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="624E0803FF834835FCA5250CB4ADFA52" blockId="3.[813,1436,1360,1447]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FCA5250CB2B7FA73" bold="true" box="[813,902,1391,1414]" italics="true" pageId="3" pageNumber="4">General</emphasis>
. Golden brown medium-sized ticcs with greatly enlarged palpal article 1, over crowded hypostome
</paragraph>
<caption id="368E588BFF834835FF1F2593B03BF9CA" ID-Table-UUID="368E588BFF834835FF1F2593B03BF9CA" httpUri="http://table.plazi.org/id/368E588BFF834835FF1F2593B03BF9CA" pageId="3" pageNumber="4" startId="3.[151,210,1520,1541]" targetBox="[151,1436,1615,1941]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="624E0803FF834835FF1F2593B03BF9CA" blockId="3.[151,1418,1520,1599]" pageId="3" pageNumber="4">
<emphasis id="5085D411FF834835FF1F2593B1D7F9F0" bold="true" box="[151,230,1520,1541]" pageId="3" pageNumber="4">Table 1</emphasis>
The specces composctcon of
<taxonomicName id="A5F17380FF834835FE752593B354F9F0" box="[509,613,1520,1541]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5085D411FF834835FE752593B304F9F0" box="[509,565,1520,1541]" italics="true" pageId="3" pageNumber="4">Ixodes</emphasis>
spp.
</taxonomicName>
detected from
<taxonomicName id="A5F17380FF834835FD742593B240F9F0" box="[764,881,1520,1541]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="5085D411FF834835FD742593B240F9F0" box="[764,881,1520,1541]" italics="true" pageId="3" pageNumber="4">B. penicillata</emphasis>
</taxonomicName>
and ssmpatrcc specces durcng trappcng sesscons conducted from 2006 through to 2016. The proportcons of each
<taxonomicName id="A5F17380FF834835FC82266EB255F9D7" box="[778,868,1549,1570]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5085D411FF834835FC82266EB273F9D7" box="[778,834,1549,1570]" italics="true" pageId="3" pageNumber="4">Ixodes</emphasis>
sp.
</taxonomicName>
among cnfected cndcvcduals are gcven as a percentage cn parentheses
</paragraph>
</caption>
<paragraph id="624E0803FF834835FE5B262CB582F860" pageId="3" pageNumber="4">
<table id="10F1FAA3FF83B7C9FF1F262CB4ADF860" box="[151,1436,1615,1941]" gridcols="5" gridrows="8" pageId="3" pageNumber="4">
<tr id="DCC10A41FF83B7C9FF1F262CB4ADF967" box="[151,1436,1615,1682]" gridrow="0" pageId="3" pageNumber="4" rowspan-0="1">
<th id="9F10633DFF83B7C9FE5B262CB34AF967" box="[467,635,1615,1682]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FE5B262CB326F98F" authority="(Woslce)" baseAuthorityName="Woslce" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="penicillata">
<emphasis id="5085D411FF834835FE5B262CB34AF997" box="[467,635,1615,1634]" italics="true" pageId="3" pageNumber="4">Bettongia penicillata</emphasis>
(Woslce)
</taxonomicName>
</th>
<th id="9F10633DFF83B7C9FD21262CB269F967" box="[681,856,1615,1682]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FD21262CB235F98F" authorityName="Wagner" authorityYear="1855" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="subSpecies" species="vulpecula" subSpecies="hypoleucus">
<emphasis id="5085D411FF834835FD21262CB235F98F" italics="true" pageId="3" pageNumber="4">Trichosurus vulpecula hypoleucus</emphasis>
</taxonomicName>
(Brushtacl possum)
</th>
<th id="9F10633DFF83B7C9FC60262CB54AF967" box="[1000,1147,1615,1682]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FC60262CB54AF997" authorityName="Gould" authorityYear="1841" box="[1000,1147,1615,1634]" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="geoffroii">
<emphasis id="5085D411FF834835FC60262CB54AF997" box="[1000,1147,1615,1634]" italics="true" pageId="3" pageNumber="4">Dasyurus geoffroii</emphasis>
</taxonomicName>
(Western quoll)
</th>
<th id="9F10633DFF83B7C9FB21262CB4ADF967" box="[1193,1436,1615,1682]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FB21262CB4A2F997" authorityName="Shaw" authorityYear="1797" box="[1193,1427,1615,1634]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="3" pageNumber="4" phylum="Chordata" rank="subSpecies" species="obesulus" subSpecies="fusciventer">
<emphasis id="5085D411FF834835FB21262CB4A2F997" box="[1193,1427,1615,1634]" italics="true" pageId="3" pageNumber="4">Isoodon obesulus fusciventer</emphasis>
</taxonomicName>
(Southern brown bandccoot)
</th>
</tr>
<tr id="DCC10A41FF83B7C9FF1F26C1B4ADF940" box="[151,1436,1698,1717]" gridrow="1" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F26C1B096F940" box="[151,423,1698,1717]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Indcvcduals sampled</th>
<td id="9F10633DFF83B7C9FE5B26C1B34AF940" box="[467,635,1698,1717]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">624</td>
<td id="9F10633DFF83B7C9FD2126C1B269F940" box="[681,856,1698,1717]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">166</td>
<td id="9F10633DFF83B7C9FC6026C1B54AF940" box="[1000,1147,1698,1717]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">89</td>
<td id="9F10633DFF83B7C9FB2126C1B4ADF940" box="[1193,1436,1698,1717]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">13</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F26A4B4ADF92F" box="[151,1436,1735,1754]" gridrow="2" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F26A4B096F92F" box="[151,423,1735,1754]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
Indcvcduals host to an
<taxonomicName id="A5F17380FF834835FEDE26A4B096F92F" box="[342,423,1735,1754]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5085D411FF834835FEDE26A4B0B9F92F" box="[342,392,1735,1754]" italics="true" pageId="3" pageNumber="4">Ixodes</emphasis>
sp.
</taxonomicName>
</th>
<td id="9F10633DFF83B7C9FE5B26A4B34AF92F" box="[467,635,1735,1754]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">290 (46.5%)</td>
<td id="9F10633DFF83B7C9FD2126A4B269F92F" box="[681,856,1735,1754]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">63 (38%)</td>
<td id="9F10633DFF83B7C9FC6026A4B54AF92F" box="[1000,1147,1735,1754]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">22 (24.7%)</td>
<td id="9F10633DFF83B7C9FB2126A4B4ADF92F" box="[1193,1436,1735,1754]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">5 (38.5%)</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F268EB4ADF8F5" box="[151,1436,1773,1792]" gridrow="3" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F268EB096F8F5" box="[151,423,1773,1792]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F268EB037F8F5" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[151,262,1773,1792]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF834835FF1F268EB037F8F5" box="[151,262,1773,1792]" italics="true" pageId="3" pageNumber="4">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF834835FE85268EB00AF8F5" box="[269,315,1773,1792]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="9F10633DFF83B7C9FE5B268EB34AF8F5" box="[467,635,1773,1792]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">123 (42.4%)</td>
<td id="9F10633DFF83B7C9FD21268EB269F8F5" box="[681,856,1773,1792]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">0</td>
<td id="9F10633DFF83B7C9FC60268EB54AF8F5" box="[1000,1147,1773,1792]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">0</td>
<td id="9F10633DFF83B7C9FB21268EB4ADF8F5" box="[1193,1436,1773,1792]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">2 (40%)</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F2771B4ADF8D0" box="[151,1436,1810,1829]" gridrow="4" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F2771B096F8D0" box="[151,423,1810,1829]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F2771B000F8D0" authorityName="Neumann" authorityYear="1904" box="[151,305,1810,1829]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="5085D411FF834835FF1F2771B000F8D0" box="[151,305,1810,1829]" italics="true" pageId="3" pageNumber="4">Ixodes australiensis</emphasis>
</taxonomicName>
</th>
<td id="9F10633DFF83B7C9FE5B2771B34AF8D0" box="[467,635,1810,1829]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">153 (52.7%)</td>
<td id="9F10633DFF83B7C9FD212771B269F8D0" box="[681,856,1810,1829]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">1 (1.5%)</td>
<td id="9F10633DFF83B7C9FC602771B54AF8D0" box="[1000,1147,1810,1829]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">13 (59%)</td>
<td id="9F10633DFF83B7C9FB212771B4ADF8D0" box="[1193,1436,1810,1829]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">2 (40%)</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F2754B4ADF8BF" box="[151,1436,1847,1866]" gridrow="5" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F2754B096F8BF" box="[151,423,1847,1866]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F2754B01FF8BF" authorityName="Roberts" authorityYear="1962" box="[151,302,1847,1866]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="myrmecobii">
<emphasis id="5085D411FF834835FF1F2754B01FF8BF" box="[151,302,1847,1866]" italics="true" pageId="3" pageNumber="4">Ixodes myrmecobii</emphasis>
</taxonomicName>
</th>
<td id="9F10633DFF83B7C9FE5B2754B34AF8BF" box="[467,635,1847,1866]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">52 (17.9%)</td>
<td id="9F10633DFF83B7C9FD212754B269F8BF" box="[681,856,1847,1866]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">3 (4.7%)</td>
<td id="9F10633DFF83B7C9FC602754B54AF8BF" box="[1000,1147,1847,1866]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">0</td>
<td id="9F10633DFF83B7C9FB212754B4ADF8BF" box="[1193,1436,1847,1866]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">1 (20%)</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F273EB4ADF885" box="[151,1436,1885,1904]" gridrow="6" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F273EB096F885" box="[151,423,1885,1904]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F273EB023F885" authorityName="Neumann" authorityYear="1899" box="[151,274,1885,1904]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tasmani">
<emphasis id="5085D411FF834835FF1F273EB023F885" box="[151,274,1885,1904]" italics="true" pageId="3" pageNumber="4">Ixodes tasmani</emphasis>
</taxonomicName>
</th>
<td id="9F10633DFF83B7C9FE5B273EB34AF885" box="[467,635,1885,1904]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">4 (1.3%)</td>
<td id="9F10633DFF83B7C9FD21273EB269F885" box="[681,856,1885,1904]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">54 (85.7%)</td>
<td id="9F10633DFF83B7C9FC60273EB54AF885" box="[1000,1147,1885,1904]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">7 (31.8%)</td>
<td id="9F10633DFF83B7C9FB21273EB4ADF885" box="[1193,1436,1885,1904]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">0</td>
</tr>
<tr id="DCC10A41FF83B7C9FF1F27E1B4ADF860" box="[151,1436,1922,1941]" gridrow="7" pageId="3" pageNumber="4">
<th id="9F10633DFF83B7C9FF1F27E1B096F860" box="[151,423,1922,1941]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
<taxonomicName id="A5F17380FF834835FF1F27E1B035F860" authorityName="Warburton &amp; Nuttall" authorityYear="1909" box="[151,260,1922,1941]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fecialis">
<emphasis id="5085D411FF834835FF1F27E1B035F860" box="[151,260,1922,1941]" italics="true" pageId="3" pageNumber="4">Ixodes fecialis</emphasis>
</taxonomicName>
</th>
<td id="9F10633DFF83B7C9FE5B27E1B34AF860" box="[467,635,1922,1941]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">1 (0.3%)</td>
<td id="9F10633DFF83B7C9FD2127E1B269F860" box="[681,856,1922,1941]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">0</td>
<td id="9F10633DFF83B7C9FC6027E1B54AF860" box="[1000,1147,1922,1941]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">9 (40.1%)</td>
<td id="9F10633DFF83B7C9FB2127E1B4ADF860" box="[1193,1436,1922,1941]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">0</td>
</tr>
</table>
</paragraph>
<paragraph id="624E0803FF844832FF1F208EB3ABFED0" blockId="4.[151,775,237,293]" pageId="4" pageNumber="5">dentition mainly 6/6 and 5/5, coxae all armed with strong pointed spurs, and anal groove open posteriorly.</paragraph>
<paragraph id="624E0803FF844833FF1F212EB1C6FD30" blockId="4.[151,775,333,612]" lastBlockId="5.[151,776,237,1764]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="5085D411FF844832FF1F212EB1D8FE91" bold="true" box="[151,233,333,356]" italics="true" pageId="4" pageNumber="5">Female</emphasis>
. Idiosoma (
<figureCitation id="FACA1486FF844832FEFF212EB08DFE90" box="[375,444,333,357]" captionStart="Fig" captionStartId="4.[167,198,1892,1911]" captionTargetBox="[151,1437,651,1950]" captionTargetId="figure-223@4.[339,1248,668,1875]" captionTargetPageId="4" captionText="Fig. 2 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 500 μm; d, 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
): Unfed specimens oval and elongate, widest just posteriorly of spiracles. Body length measured dorsally from midway between scapular points to most posterior margin range from 2.4
<emphasis id="5085D411FF844832FDDC21CFB355FE31" box="[596,612,428,452]" italics="true" pageId="4" pageNumber="5"></emphasis>
3.3, width 1.4
<emphasis id="5085D411FF844832FD7F21CFB236FE31" box="[759,775,428,452]" italics="true" pageId="4" pageNumber="5"></emphasis>
2.2 (
<figureCitation id="FACA1486FF844832FF4221AFB028FE11" box="[202,281,460,484]" captionStart="Fig" captionStartId="4.[167,198,1892,1911]" captionTargetBox="[151,1437,651,1950]" captionTargetId="figure-223@4.[339,1248,668,1875]" captionTargetPageId="4" captionText="Fig. 2 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 500 μm; d, 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="4" pageNumber="5">Fig. 2a</figureCitation>
). Partially fed specimens length range from 3.3
<emphasis id="5085D411FF844832FF3E218FB1F7FDF1" box="[182,198,492,516]" italics="true" pageId="4" pageNumber="5"></emphasis>
5.3, width 1.9
<emphasis id="5085D411FF844832FEE4218FB04DFDF1" box="[364,380,492,516]" italics="true" pageId="4" pageNumber="5"></emphasis>
2.5, engorged specimens attaining length 11, width 6.5. Dorsal setae are short (&lt;25 μm), lay within a uniform moderate covering of shallow punctations, marginal grooves are well defined. The scutum about as long as wide, widest point posterior to midlength, anterolateral and posterolateral margins are mildly sinuous with posterior angle broadly rounded, lateral carinae present (
<figureCitation id="FACA1486FF844832FC4C212EB52BFE90" box="[964,1050,333,357]" captionStart="Fig" captionStartId="4.[167,198,1892,1911]" captionTargetBox="[151,1437,651,1950]" captionTargetId="figure-223@4.[339,1248,668,1875]" captionTargetPageId="4" captionText="Fig. 2 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 500 μm; d, 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="4" pageNumber="5">Figs. 2c</figureCitation>
and
<figureCitation id="FACA1486FF844832FBDE212EB540FE90" box="[1110,1137,333,357]" captionStart="Fig" captionStartId="7.[167,198,1087,1106]" captionTargetBox="[151,1437,236,1145]" captionTargetId="figure-451@7.[339,1248,253,1069]" captionTargetPageId="7" captionText="Fig. 5 Lcne drawcng of Ixodes woyliei n. sp. Female. a Capctulum, ventral vcew. b Capctulum, dorsal vcew. c Scutum. d Tarscs I. e Tarscs IV. f Coxae. Scale-bars: 200 μm" figureDoi="http://doi.org/10.5281/zenodo.11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="4" pageNumber="5">5c</figureCitation>
). Scutal length range from 1.2
<emphasis id="5085D411FF844832FCC4210FB26DFE71" box="[844,860,364,388]" italics="true" pageId="4" pageNumber="5"></emphasis>
1.5, width 1.2
<emphasis id="5085D411FF844832FC65210FB2CCFE71" box="[1005,1021,364,388]" italics="true" pageId="4" pageNumber="5"></emphasis>
1.6. Punctations are shallow and moderate in number, becoming coarser in cervical grooves, lateral rugae and along the posterior margin; scutal setae are minute (&lt;8 μm). Cervical grooves well defined anteriorly becoming shallow posteriorly and extending to, or almost to the scutal margin. Scapulae are large and bluntly pointed. Ventral setae are longer than dorsal (&lt;40 μm) with two rows of longer setae (&gt;40 μm) around the lateral side of the spiracular plate. Genital aperture is level with the anterior margin of coxa III, but moving towards second intercoxal space on engorgement (
<figureCitation id="FACA1486FF854833FDB4214EB3B9FEB0" box="[572,648,301,325]" captionStart="Fig" captionStartId="4.[167,198,1892,1911]" captionTargetBox="[151,1437,651,1950]" captionTargetId="figure-223@4.[339,1248,668,1875]" captionTargetPageId="4" captionText="Fig. 2 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 500 μm; d, 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="5" pageNumber="6">Fig. 2b</figureCitation>
). Spiracular plates suboval, length 0.16
<emphasis id="5085D411FF854833FE3F212FB0F6FE91" box="[439,455,332,356]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.30, with approximately 3
<emphasis id="5085D411FF854833FD62212FB3CBFE91" box="[746,762,332,356]" italics="true" pageId="5" pageNumber="6"></emphasis>
4 rows of goblets, macular eccentric (
<figureCitation id="FACA1486FF854833FDA8210EB35FFE70" box="[544,622,365,389]" captionStart="Fig" captionStartId="6.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-375@6.[339,1248,253,1198]" captionTargetPageId="6" captionText="Fig. 4 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, 200 μm; b, c, 50 μm; d, 15 μm" figureDoi="http://doi.org/10.5281/zenodo.11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="5" pageNumber="6">Fig. 4b</figureCitation>
). Anal groove is horseshoe shaped, rounded anteriorly, curving gently and convergently posteriorly but becoming slightly divergent near body margin and remaining widely open (
<figureCitation id="FACA1486FF854833FF16218EB1D8FDF0" box="[158,233,493,517]" captionStart="Fig" captionStartId="4.[167,198,1892,1911]" captionTargetBox="[151,1437,651,1950]" captionTargetId="figure-223@4.[339,1248,668,1875]" captionTargetPageId="4" captionText="Fig. 2 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 500 μm; d, 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="5" pageNumber="6">Fig. 2d</figureCitation>
). Both the internal and external margin of the anal groove epicutical surface possess several rows of inward facing spines, overlapping across the divide of the anal groove and running laterally for most of its length. This feature has not been mentioned previously but appears to be typical for all
<taxonomicName id="A5F17380FF854833FED822EFB0D6FD50" box="[336,487,652,677]" pageId="5" pageNumber="6">
<emphasis id="5085D411FF854833FED822EFB0A2FD51" box="[336,403,652,676]" italics="true" pageId="5" pageNumber="6">Ixodes</emphasis>
species
</taxonomicName>
that we have been able to examine.
</paragraph>
<caption id="368E588BFF844832FF2F2707B577F867" ID-DOI="http://doi.org/10.5281/zenodo.11551876" ID-Zenodo-Dep="11551876" httpUri="https://zenodo.org/record/11551876/files/figure.png" pageId="4" pageNumber="5" startId="4.[167,198,1892,1911]" targetBox="[151,1437,651,1950]" targetPageId="4" targetType="figure">
<paragraph id="624E0803FF844832FF2F2707B577F867" blockId="4.[167,1390,1892,1938]" pageId="4" pageNumber="5">
<emphasis id="5085D411FF844832FF2F2707B1EDF882" bold="true" box="[167,220,1892,1911]" pageId="4" pageNumber="5">Fig. 2</emphasis>
Scanncng electron mccrographs of
<taxonomicName id="A5F17380FF844832FD832707B34BF882" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[523,634,1892,1911]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF844832FD832707B34BF882" box="[523,634,1892,1911]" italics="true" pageId="4" pageNumber="5">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF844832FD092707B39EF882" box="[641,687,1892,1911]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
Female.
<emphasis id="5085D411FF844832FD722707B234F882" bold="true" box="[762,773,1892,1911]" pageId="4" pageNumber="5">a</emphasis>
Idcosoma, unengorged speccmen, dorsal vcew.
<emphasis id="5085D411FF844832FB122707B597F882" bold="true" box="[1178,1190,1892,1911]" pageId="4" pageNumber="5">b</emphasis>
Idcosoma, unengorged speccmen, ventral vcew.
<emphasis id="5085D411FF844832FEFD271CB04EF867" bold="true" box="[373,383,1919,1938]" pageId="4" pageNumber="5">c</emphasis>
Scutum, showcng lateral carcnae.
<emphasis id="5085D411FF844832FD16271CB39BF867" bold="true" box="[670,682,1919,1938]" pageId="4" pageNumber="5">d</emphasis>
Anal groove.
<emphasis id="5085D411FF844832FCAB271CB247F867" box="[803,886,1919,1938]" italics="true" pageId="4" pageNumber="5">Scale-bars</emphasis>
:
<emphasis id="5085D411FF844832FCF7271CB2ABF867" bold="true" box="[895,922,1919,1938]" pageId="4" pageNumber="5">a -c</emphasis>
, 500 μm;
<emphasis id="5085D411FF844832FC67271CB2CAF867" bold="true" box="[1007,1019,1919,1938]" pageId="4" pageNumber="5">d</emphasis>
, 100 μm
</paragraph>
</caption>
<paragraph id="624E0803FF854833FF2522AEB081FA70" blockId="5.[151,776,237,1764]" pageId="5" pageNumber="6">
Gnathosoma (
<figureCitation id="FACA1486FF854833FECF22AEB0B4FD10" box="[327,389,717,741]" captionStart="Fig" captionStartId="5.[829,860,1472,1491]" captionTargetBox="[813,1437,236,1557]" captionTargetId="figure-599@5.[838,1411,253,1455]" captionTargetPageId="5" captionText="Fig. 3 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a-b, 100 μm; c, 20 μm" figureDoi="http://doi.org/10.5281/zenodo.11551878" httpUri="https://zenodo.org/record/11551878/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) Basis dorsally, short,
<quantity id="A509A5E6FF854833FD0822AEB236FD10" box="[640,775,716,741]" metricMagnitude="-2" metricUnit="m" metricValue="1.2192" metricValueMax="1.524" metricValueMin="0.9144" pageId="5" pageNumber="6" unit="in" value="0.48" valueMax="0.6" valueMin="0.36">
0.36
<emphasis id="5085D411FF854833FD2422AFB38DFD11" box="[684,700,716,740]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.60 in
</quantity>
length by
<quantity id="A509A5E6FF854833FE8D228EB0BAFCF0" box="[261,395,748,773]" metricMagnitude="-2" metricUnit="m" metricValue="1.2065" metricValueMax="1.397" metricValueMin="1.016" pageId="5" pageNumber="6" unit="in" value="0.475" valueMax="0.55" valueMin="0.4">
0.40
<emphasis id="5085D411FF854833FEB9228FB070FCF1" box="[305,321,748,772]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.55 in
</quantity>
width. Length measurement tacen from top of the palpi to the posterior margin of the cornua. Dorsal basis capituli subrectangular, with one median depression and a lateral depression on each side, the depressed areas being separated by carinae, posterior margin slightly undulating with small indistinct cornua (
<figureCitation id="FACA1486FF854833FD3B23EEB236FC50" box="[691,775,909,933]" captionStart="Fig" captionStartId="5.[829,860,1472,1491]" captionTargetBox="[813,1437,236,1557]" captionTargetId="figure-599@5.[838,1411,253,1455]" captionTargetPageId="5" captionText="Fig. 3 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a-b, 100 μm; c, 20 μm" figureDoi="http://doi.org/10.5281/zenodo.11551878" httpUri="https://zenodo.org/record/11551878/files/figure.png" pageId="5" pageNumber="6">Figs. 3a</figureCitation>
and
<figureCitation id="FACA1486FF854833FF4223CEB1D7FC30" box="[202,230,941,965]" captionStart="Fig" captionStartId="7.[167,198,1087,1106]" captionTargetBox="[151,1437,236,1145]" captionTargetId="figure-451@7.[339,1248,253,1069]" captionTargetPageId="7" captionText="Fig. 5 Lcne drawcng of Ixodes woyliei n. sp. Female. a Capctulum, ventral vcew. b Capctulum, dorsal vcew. c Scutum. d Tarscs I. e Tarscs IV. f Coxae. Scale-bars: 200 μm" figureDoi="http://doi.org/10.5281/zenodo.11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="5" pageNumber="6">5b</figureCitation>
). Porose areas large, suboval, lying in lateral depressions, widely separated by the median depression. Basis ventrally about as long as wide with small but distinct auriculae. Palps short and article 1 greatly enlarged, extending inwardly to partially ensheathe base of mouthparts, ventrally with a strong posterolateral salience. Articles 2 and 3 are without apparent suture, total length 0.29
<emphasis id="5085D411FF854833FF4B24EFB1E2FB51" box="[195,211,1164,1188]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.35, width 0.07
<emphasis id="5085D411FF854833FE0824EFB0A1FB51" box="[384,400,1164,1188]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.15 (
<figureCitation id="FACA1486FF854833FE4524EEB313FB50" box="[461,546,1165,1189]" captionStart="Fig" captionStartId="5.[829,860,1472,1491]" captionTargetBox="[813,1437,236,1557]" captionTargetId="figure-599@5.[838,1411,253,1455]" captionTargetPageId="5" captionText="Fig. 3 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a-b, 100 μm; c, 20 μm" figureDoi="http://doi.org/10.5281/zenodo.11551878" httpUri="https://zenodo.org/record/11551878/files/figure.png" pageId="5" pageNumber="6">Figs. 3b</figureCitation>
and
<figureCitation id="FACA1486FF854833FDD424EEB347FB50" box="[604,630,1165,1189]" captionStart="Fig" captionStartId="7.[167,198,1087,1106]" captionTargetBox="[151,1437,236,1145]" captionTargetId="figure-451@7.[339,1248,253,1069]" captionTargetPageId="7" captionText="Fig. 5 Lcne drawcng of Ixodes woyliei n. sp. Female. a Capctulum, ventral vcew. b Capctulum, dorsal vcew. c Scutum. d Tarscs I. e Tarscs IV. f Coxae. Scale-bars: 200 μm" figureDoi="http://doi.org/10.5281/zenodo.11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="5" pageNumber="6">5a</figureCitation>
). Hypostome length ranges from 0.17
<emphasis id="5085D411FF854833FE1324CFB09AFB31" box="[411,427,1196,1220]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.33, width 0.15
<emphasis id="5085D411FF854833FDD024CFB359FB31" box="[600,616,1196,1220]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.17, spatulate, broad anteriorly, with sharply pointed large denticles, mainly 6/6 and 5/5. Dentition formula essentially 12/12 of small over crowded denticles at the corona, dropping in number but increasing in size to 6/6 and 5/5 by anterior third and reducing to 4/4, 3/3 with crenulations running down to the base (
<figureCitation id="FACA1486FF854833FED4250EB092FA70" box="[348,419,1389,1413]" captionStart="Fig" captionStartId="5.[829,860,1472,1491]" captionTargetBox="[813,1437,236,1557]" captionTargetId="figure-599@5.[838,1411,253,1455]" captionTargetPageId="5" captionText="Fig. 3 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a-b, 100 μm; c, 20 μm" figureDoi="http://doi.org/10.5281/zenodo.11551878" httpUri="https://zenodo.org/record/11551878/files/figure.png" pageId="5" pageNumber="6">Fig. 3c</figureCitation>
).
</paragraph>
<paragraph id="624E0803FF854833FF2525EFB3FFF911" blockId="5.[151,776,237,1764]" pageId="5" pageNumber="6">
Legs (
<figureCitation id="FACA1486FF854833FF7B25EFB008FA51" box="[243,313,1420,1444]" captionStart="Fig" captionStartId="6.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-375@6.[339,1248,253,1198]" captionTargetPageId="6" captionText="Fig. 4 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, 200 μm; b, c, 50 μm; d, 15 μm" figureDoi="http://doi.org/10.5281/zenodo.11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
): Slender and moderate length. Coxa I transversely elongate with a strong pointed external spur. Coxae II, III, and IV somewhat square with progressively smaller pointed external spurs, all coxae with few setae, syncoxae absent (
<figureCitation id="FACA1486FF854833FE5D266FB31AF9D1" box="[469,555,1548,1572]" captionStart="Fig" captionStartId="6.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-375@6.[339,1248,253,1198]" captionTargetPageId="6" captionText="Fig. 4 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, 200 μm; b, c, 50 μm; d, 15 μm" figureDoi="http://doi.org/10.5281/zenodo.11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="5" pageNumber="6">Figs. 4a</figureCitation>
and
<figureCitation id="FACA1486FF854833FDE1266FB34FF9D1" box="[617,638,1548,1572]" captionStart="Fig" captionStartId="7.[167,198,1087,1106]" captionTargetBox="[151,1437,236,1145]" captionTargetId="figure-451@7.[339,1248,253,1069]" captionTargetPageId="7" captionText="Fig. 5 Lcne drawcng of Ixodes woyliei n. sp. Female. a Capctulum, ventral vcew. b Capctulum, dorsal vcew. c Scutum. d Tarscs I. e Tarscs IV. f Coxae. Scale-bars: 200 μm" figureDoi="http://doi.org/10.5281/zenodo.11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="5" pageNumber="6">5f</figureCitation>
). Length of tarsus I 0.3
<emphasis id="5085D411FF854833FE9B264FB012F9B1" box="[275,291,1580,1604]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.5 with few long setae (&lt;50 μm) and some small (&lt;20 μm) (
<figureCitation id="FACA1486FF854833FEC4262FB0AEF991" box="[332,415,1612,1636]" captionStart="Fig" captionStartId="6.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-375@6.[339,1248,253,1198]" captionTargetPageId="6" captionText="Fig. 4 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, 200 μm; b, c, 50 μm; d, 15 μm" figureDoi="http://doi.org/10.5281/zenodo.11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="5" pageNumber="6">Figs. 4c</figureCitation>
and
<figureCitation id="FACA1486FF854833FE5F262FB33DF991" box="[471,524,1612,1636]" captionStart="Fig" captionStartId="7.[167,198,1087,1106]" captionTargetBox="[151,1437,236,1145]" captionTargetId="figure-451@7.[339,1248,253,1069]" captionTargetPageId="7" captionText="Fig. 5 Lcne drawcng of Ixodes woyliei n. sp. Female. a Capctulum, ventral vcew. b Capctulum, dorsal vcew. c Scutum. d Tarscs I. e Tarscs IV. f Coxae. Scale-bars: 200 μm" figureDoi="http://doi.org/10.5281/zenodo.11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="5" pageNumber="6">5d, e</figureCitation>
). Haller
<emphasis id="5085D411FF854833FDED262FB358F991" box="[613,617,1612,1636]" italics="true" pageId="5" pageNumber="6"></emphasis>
s organ, anterior pit suboval with seven sensilla arranged in a cluster in the centre, posterior capsule opening slightly above the pit and divided by a low ridge with up to five sensilla seen within (
<figureCitation id="FACA1486FF854833FEAB26AFB041F911" box="[291,368,1740,1764]" captionStart="Fig" captionStartId="6.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-375@6.[339,1248,253,1198]" captionTargetPageId="6" captionText="Fig. 4 Scanncng electron mccrographs of Ixodes woyliei n. sp. Female. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, 200 μm; b, c, 50 μm; d, 15 μm" figureDoi="http://doi.org/10.5281/zenodo.11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="5" pageNumber="6">Fig. 4d</figureCitation>
). Length of tarsus IV is 0.4
<emphasis id="5085D411FF854833FD1026AFB399F911" box="[664,680,1740,1764]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.5.
</paragraph>
<caption id="368E588BFF854833FCB525A3B57DF9FC" ID-DOI="http://doi.org/10.5281/zenodo.11551878" ID-Zenodo-Dep="11551878" httpUri="https://zenodo.org/record/11551878/files/figure.png" pageId="5" pageNumber="6" startId="5.[829,860,1472,1491]" targetBox="[813,1437,236,1557]" targetPageId="5" targetType="figure">
<paragraph id="624E0803FF854833FCB525A3B44BFA26" blockId="5.[829,1422,1472,1545]" box="[829,1402,1472,1491]" pageId="5" pageNumber="6">
<emphasis id="5085D411FF854833FCB525A3B243FA26" bold="true" box="[829,882,1472,1491]" pageId="5" pageNumber="6">Fig. 3</emphasis>
Scanncng electron mccrographs of
<taxonomicName id="A5F17380FF854833FB1F25A3B432FA26" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1175,1283,1472,1491]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF854833FB1F25A3B432FA26" box="[1175,1283,1472,1491]" italics="true" pageId="5" pageNumber="6">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF854833FA8025A3B405FA26" box="[1288,1332,1472,1491]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
Female.
</paragraph>
<paragraph id="624E0803FF854833FCB525B8B57DF9FC" blockId="5.[829,1422,1472,1545]" pageId="5" pageNumber="6">
<emphasis id="5085D411FF854833FCB525B8B279FA1B" bold="true" box="[829,840,1499,1518]" pageId="5" pageNumber="6">a</emphasis>
Gnathosoma, dorsal vcew.
<emphasis id="5085D411FF854833FBAF25B8B502FA1B" bold="true" box="[1063,1075,1499,1518]" pageId="5" pageNumber="6">b</emphasis>
Gnathosoma, ventral vcew.
<emphasis id="5085D411FF854833FA9125B8B412FA1B" bold="true" box="[1305,1315,1499,1518]" pageId="5" pageNumber="6">c</emphasis>
Hspostome.
<emphasis id="5085D411FF854833FCB52595B2BCF9FC" box="[829,909,1526,1545]" italics="true" pageId="5" pageNumber="6">Scale-bars</emphasis>
:
<emphasis id="5085D411FF854833FC1E2595B283F9FC" bold="true" box="[918,946,1526,1545]" pageId="5" pageNumber="6">a -b</emphasis>
, 100 μm;
<emphasis id="5085D411FF854833FB8C2595B53CF9FC" bold="true" box="[1028,1037,1526,1545]" pageId="5" pageNumber="6">c</emphasis>
, 20 μm
</paragraph>
</caption>
<paragraph id="624E0803FF854830FF1F276FB330F930" blockId="5.[151,775,1804,1956]" lastBlockId="6.[151,776,1357,1957]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="5085D411FF854833FF1F276FB1D6F8D6" bold="true" box="[151,231,1804,1827]" italics="true" pageId="5" pageNumber="6">Nymph</emphasis>
. Idiosoma (
<figureCitation id="FACA1486FF854833FEFD276FB08BF8D1" box="[373,442,1804,1828]" captionStart="Fig" captionStartId="8.[167,198,1480,1499]" captionTargetBox="[151,1437,236,1539]" captionTargetId="figure-199@8.[339,1248,253,1463]" captionTargetPageId="8" captionText="Fig. 6 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 200 μm; d, 40 μm" figureDoi="http://doi.org/10.5281/zenodo.11551884" httpUri="https://zenodo.org/record/11551884/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
): Unfed specimens oval and elongate, widest about mid length between coxae III and IV, well-defined marginal grooves and minute dorsal setae (&lt;10) within uniformly scattered punctations (
<figureCitation id="FACA1486FF854833FDF1270FB3F1F871" box="[633,704,1900,1924]" captionStart="Fig" captionStartId="8.[167,198,1480,1499]" captionTargetBox="[151,1437,236,1539]" captionTargetId="figure-199@8.[339,1248,253,1463]" captionTargetPageId="8" captionText="Fig. 6 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 200 μm; d, 40 μm" figureDoi="http://doi.org/10.5281/zenodo.11551884" httpUri="https://zenodo.org/record/11551884/files/figure.png" pageId="5" pageNumber="6">Fig. 6a</figureCitation>
). Body length measured dorsally from midway between the scapular points to most posterior margin range from 1.3
<emphasis id="5085D411FF854833FCC5260FB26CF971" box="[845,861,1644,1668]" italics="true" pageId="5" pageNumber="6"></emphasis>
1.6, width range 0.78
<emphasis id="5085D411FF854833FBD7260FB55EF971" box="[1119,1135,1644,1668]" italics="true" pageId="5" pageNumber="6"></emphasis>
1.1. Fed specimen length range from 1.7
<emphasis id="5085D411FF854833FC6C26EFB2C5F951" box="[996,1012,1676,1700]" italics="true" pageId="5" pageNumber="6"></emphasis>
2.2, width 0.97
<emphasis id="5085D411FF854833FB2326EFB58AF951" box="[1195,1211,1676,1700]" italics="true" pageId="5" pageNumber="6"></emphasis>
1.6, with engorged specimens attaining length 4.0, width 2.0. Scutum wider than long, with posterior angle broadly rounded, lateral carinae present (
<figureCitation id="FACA1486FF854833FC62268FB573F8F1" box="[1002,1090,1772,1796]" captionStart="Fig" captionStartId="8.[167,198,1480,1499]" captionTargetBox="[151,1437,236,1539]" captionTargetId="figure-199@8.[339,1248,253,1463]" captionTargetPageId="8" captionText="Fig. 6 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 200 μm; d, 40 μm" figureDoi="http://doi.org/10.5281/zenodo.11551884" httpUri="https://zenodo.org/record/11551884/files/figure.png" pageId="5" pageNumber="6">Figs. 6c</figureCitation>
and
<figureCitation id="FACA1486FF854833FB0C268FB5AEF8F1" box="[1156,1183,1772,1796]" captionStart="Fig" captionStartId="11.[167,198,1129,1148]" captionTargetBox="[151,1437,236,1161]" captionTargetId="figure-8@11.[339,1248,253,1112]" captionTargetPageId="11" captionText="Fig. 9 Lcne drawcng of Ixodes woyliei n. sp. Nsmph. a Capctulum ventral vcew. b Capctulum dorsal vcew. c Scutum. d Coxae. Scale-bars: 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551890" httpUri="https://zenodo.org/record/11551890/files/figure.png" pageId="5" pageNumber="6">9c</figureCitation>
). Scutal length ranges from 0.52
<emphasis id="5085D411FF854833FC1E276FB297F8D1" box="[918,934,1804,1828]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.63, width 0.63
<emphasis id="5085D411FF854833FBD9276FB550F8D1" box="[1105,1121,1804,1828]" italics="true" pageId="5" pageNumber="6"></emphasis>
0.75. Punctations are shallow and moderate in number, becoming coarser in cervical grooves, lateral rugae and along the posterior margin; scutal setae are minute (&lt;8 μm). Cervical grooves are well defined anteriorly becoming shallow posteriorly and extending to, or almost to the scutal margin. Scapulae are large and bluntly pointed. Ventral setae are slightly longer than dorsal mostly (&lt;15 μm) with some longer setae (&gt;20 μm) around the spiracular plate (
<figureCitation id="FACA1486FF864830FDCC25CEB3A2FA30" box="[580,659,1453,1477]" captionStart="Fig" captionStartId="8.[167,198,1480,1499]" captionTargetBox="[151,1437,236,1539]" captionTargetId="figure-199@8.[339,1248,253,1463]" captionTargetPageId="8" captionText="Fig. 6 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 200 μm; d, 40 μm" figureDoi="http://doi.org/10.5281/zenodo.11551884" httpUri="https://zenodo.org/record/11551884/files/figure.png" pageId="6" pageNumber="7">Fig. 6b</figureCitation>
). Spiracles suboval, length 0.095
<emphasis id="5085D411FF864830FEF725AFB0BEFA11" box="[383,399,1484,1508]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.10, width 0.065
<emphasis id="5085D411FF864830FDC425AFB36DFA11" box="[588,604,1484,1508]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.12 with
<emphasis id="5085D411FF864830FD4525AEB3E6FA11" box="[717,727,1485,1508]" italics="true" pageId="6" pageNumber="7">c</emphasis>
.3
<emphasis id="5085D411FF864830FD6225AFB3CBFA11" box="[746,762,1484,1508]" italics="true" pageId="6" pageNumber="7"></emphasis>
4 rows of goblets covering the whole surface (
<figureCitation id="FACA1486FF864830FDFC258EB38EF9F0" box="[628,703,1517,1541]" captionStart="Fig" captionStartId="10.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-344@10.[339,1248,253,1199]" captionTargetPageId="10" captionText="Fig. 8 Scanncng electron mccrographs of Ixodes woyliei. Nsmph, legs and spcracular plate. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, c, 100 μm; b, 20 μm; d, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551888" httpUri="https://zenodo.org/record/11551888/files/figure.png" pageId="6" pageNumber="7">Fig. 8b</figureCitation>
). Anal groove is horseshoe shaped, rounded anteriorly, curving gently and convergently posteriorly but becoming slightly divergent near body margin and remaining widely open (
<figureCitation id="FACA1486FF864830FF16260EB1D8F970" box="[158,233,1645,1669]" captionStart="Fig" captionStartId="8.[167,198,1480,1499]" captionTargetBox="[151,1437,236,1539]" captionTargetId="figure-199@8.[339,1248,253,1463]" captionTargetPageId="8" captionText="Fig. 6 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Idcosoma, unengorged speccmen, dorsal vcew. b Idcosoma, unengorged speccmen, ventral vcew. c Scutum, showcng lateral carcnae. d Anal groove. Scale-bars: a-c, 200 μm; d, 40 μm" figureDoi="http://doi.org/10.5281/zenodo.11551884" httpUri="https://zenodo.org/record/11551884/files/figure.png" pageId="6" pageNumber="7">Fig. 6d</figureCitation>
). Both the internal and external margin of the anal groove epicutical possess several rows of inward facing spines as seen in the adult female.
</paragraph>
<caption id="368E588BFF864830FF2F24A3B0F1FB1B" ID-DOI="http://doi.org/10.5281/zenodo.11551880" ID-Zenodo-Dep="11551880" httpUri="https://zenodo.org/record/11551880/files/figure.png" pageId="6" pageNumber="7" startId="6.[167,198,1216,1235]" targetBox="[151,1437,236,1274]" targetPageId="6" targetType="figure">
<paragraph id="624E0803FF864830FF2F24A3B446FB26" blockId="6.[167,1399,1216,1262]" box="[167,1399,1216,1235]" pageId="6" pageNumber="7">
<emphasis id="5085D411FF864830FF2F24A3B1EDFB26" bold="true" box="[167,220,1216,1235]" pageId="6" pageNumber="7">Fig. 4</emphasis>
Scanncng electron mccrographs of
<taxonomicName id="A5F17380FF864830FD8324A3B34BFB26" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[523,634,1216,1235]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF864830FD8324A3B34BFB26" box="[523,634,1216,1235]" italics="true" pageId="6" pageNumber="7">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF864830FD0924A3B39EFB26" box="[641,687,1216,1235]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
Female.
<emphasis id="5085D411FF864830FD7224A3B234FB26" bold="true" box="[762,773,1216,1235]" pageId="6" pageNumber="7">a</emphasis>
Spurs on coxae.
<emphasis id="5085D411FF864830FC1024A3B295FB26" bold="true" box="[920,932,1216,1235]" pageId="6" pageNumber="7">b</emphasis>
Spcracular plate.
<emphasis id="5085D411FF864830FBB024A3B573FB26" bold="true" box="[1080,1090,1216,1235]" pageId="6" pageNumber="7">c</emphasis>
Tarsus I.
<emphasis id="5085D411FF864830FB1824A3B5ADFB26" bold="true" box="[1168,1180,1216,1235]" pageId="6" pageNumber="7">d</emphasis>
Haller
<emphasis id="5085D411FF864830FB5B24A3B5E6FB26" box="[1235,1239,1216,1235]" italics="true" pageId="6" pageNumber="7"></emphasis>
s organ.
<emphasis id="5085D411FF864830FAA924A3B445FB26" box="[1313,1396,1216,1235]" italics="true" pageId="6" pageNumber="7">Scale-bars</emphasis>
:
</paragraph>
<paragraph id="624E0803FF864830FF2F24B8B0F1FB1B" blockId="6.[167,1399,1216,1262]" box="[167,448,1242,1262]" pageId="6" pageNumber="7">
<emphasis id="5085D411FF864830FF2F24B8B183FB1B" bold="true" box="[167,178,1243,1262]" pageId="6" pageNumber="7">a</emphasis>
, 200 μm;
<emphasis id="5085D411FF864830FE8F24B8B022FB1B" bold="true" box="[263,275,1243,1262]" pageId="6" pageNumber="7">b</emphasis>
,
<emphasis id="5085D411FF864830FE9624B8B019FB1B" bold="true" box="[286,296,1243,1262]" pageId="6" pageNumber="7">c</emphasis>
, 50 μm;
<emphasis id="5085D411FF864830FEFA24B8B04FFB1B" bold="true" box="[370,382,1243,1262]" pageId="6" pageNumber="7">d</emphasis>
, 15 μm
</paragraph>
</caption>
<paragraph id="624E0803FF864830FF2526AEB4BCF9B1" blockId="6.[151,776,1357,1957]" lastBlockId="6.[813,1437,1357,1956]" pageId="6" pageNumber="7">
Gnathosoma (
<figureCitation id="FACA1486FF864830FECE26AEB0B7F910" box="[326,390,1741,1765]" captionStart="Fig" captionStartId="9.[167,198,1535,1554]" captionTargetBox="[151,775,236,1620]" captionTargetId="figure-535@9.[176,749,253,1518]" captionTargetPageId="9" captionText="Fig. 7 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a, b = 40 μm; c, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551886" httpUri="https://zenodo.org/record/11551886/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
): Basis dorsal length measurement tacen from the top of the palpi to the posterior margin of the basis, length 0.19
<emphasis id="5085D411FF864830FE07276FB0AFF8D1" box="[399,414,1804,1828]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.26, width
<quantity id="A509A5E6FF864830FDA3276EB387F8D0" box="[555,694,1804,1829]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" metricValueMax="6.604" metricValueMin="5.587999999999999" pageId="6" pageNumber="7" unit="in" value="0.24" valueMax="0.26" valueMin="0.22">
0.22
<emphasis id="5085D411FF864830FDDE276FB354F8D1" box="[598,613,1804,1828]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.26 in
</quantity>
width. Dorsal basis capituli subrectangular, posterolateral angles roundly pointed (
<figureCitation id="FACA1486FF864830FEDD272FB098F891" box="[341,425,1868,1892]" captionStart="Fig" captionStartId="9.[167,198,1535,1554]" captionTargetBox="[151,775,236,1620]" captionTargetId="figure-535@9.[176,749,253,1518]" captionTargetPageId="9" captionText="Fig. 7 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a, b = 40 μm; c, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551886" httpUri="https://zenodo.org/record/11551886/files/figure.png" pageId="6" pageNumber="7">Figs. 7a</figureCitation>
ad 9b). Basis ventrally rounded posteriorly with small auriculae. Palps short and article 1 greatly enlarged, extending inwardly to partially ensheathe base of mouthparts, ventrally with a strong posterolateral salience. Articles 2 and 3 without apparent suture total length 0.15
<emphasis id="5085D411FF864830FC2C25EFB282FA51" box="[932,947,1420,1444]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.17 by width
<geoCoordinate id="07C56EC4FF864830FBDC25EEB5BDFA50" box="[1108,1164,1421,1445]" degrees="0.055" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="55" value="0.055">0.055</geoCoordinate>
<geoCoordinate id="07C56EC4FF864830FB0425EFB5E2FA50" box="[1164,1235,1420,1445]" degrees="0.080" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="-0.08">
<emphasis id="5085D411FF864830FB0425EFB5AAFA51" box="[1164,1179,1420,1444]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.080
</geoCoordinate>
(
<figureCitation id="FACA1486FF864830FB6D25EEB40BFA50" box="[1253,1338,1421,1445]" captionStart="Fig" captionStartId="9.[167,198,1535,1554]" captionTargetBox="[151,775,236,1620]" captionTargetId="figure-535@9.[176,749,253,1518]" captionTargetPageId="9" captionText="Fig. 7 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a, b = 40 μm; c, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551886" httpUri="https://zenodo.org/record/11551886/files/figure.png" pageId="6" pageNumber="7">Figs. 7b</figureCitation>
and
<figureCitation id="FACA1486FF864830FAFF25EEB4A0FA50" box="[1399,1425,1421,1445]" captionStart="Fig" captionStartId="11.[167,198,1129,1148]" captionTargetBox="[151,1437,236,1161]" captionTargetId="figure-8@11.[339,1248,253,1112]" captionTargetPageId="11" captionText="Fig. 9 Lcne drawcng of Ixodes woyliei n. sp. Nsmph. a Capctulum ventral vcew. b Capctulum dorsal vcew. c Scutum. d Coxae. Scale-bars: 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551890" httpUri="https://zenodo.org/record/11551890/files/figure.png" pageId="6" pageNumber="7">9a</figureCitation>
). Hypostome length ranges from 0.13
<emphasis id="5085D411FF864830FB3725CFB5FFFA31" box="[1215,1230,1452,1476]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.17, width
<geoCoordinate id="07C56EC4FF864830FADD25CEB4BDFA30" box="[1365,1420,1453,1477]" degrees="0.065" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="55" value="0.065">0.065</geoCoordinate>
<geoCoordinate id="07C56EC4FF864830FA0425CFB256FA10" degrees="0.130" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="-0.13">
<emphasis id="5085D411FF864830FA0425CFB4ADFA31" box="[1420,1436,1452,1476]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.130
</geoCoordinate>
, spatulate, broad anteriorly, with sharply pointed denticles, mainly 3/3. Dentition formula essentially 6/6 small denticles at corona then decreasing in number, but increasing in size to 4/4 followed by about 8 rows of 3/3 (
<figureCitation id="FACA1486FF864830FAB3264FB4B0F9B1" box="[1339,1409,1580,1604]" captionStart="Fig" captionStartId="9.[167,198,1535,1554]" captionTargetBox="[151,775,236,1620]" captionTargetId="figure-535@9.[176,749,253,1518]" captionTargetPageId="9" captionText="Fig. 7 Scanncng electron mccrographs of Ixodes woyliei n. sp. Nsmph. a Gnathosoma, dorsal vcew. b Gnathosoma, ventral vcew. c Hspostome. Scale-bars: a, b = 40 μm; c, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551886" httpUri="https://zenodo.org/record/11551886/files/figure.png" pageId="6" pageNumber="7">Fig. 7c</figureCitation>
).
</paragraph>
<paragraph id="624E0803FF864830FCCA262FB474F851" blockId="6.[813,1437,1357,1956]" pageId="6" pageNumber="7">
Legs (
<figureCitation id="FACA1486FF864830FC0D262FB2F4F991" box="[901,965,1612,1636]" captionStart="Fig" captionStartId="10.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-344@10.[339,1248,253,1199]" captionTargetPageId="10" captionText="Fig. 8 Scanncng electron mccrographs of Ixodes woyliei. Nsmph, legs and spcracular plate. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, c, 100 μm; b, 20 μm; d, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551888" httpUri="https://zenodo.org/record/11551888/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
): Slender and of moderate length. Coxa I transversely elongate with a strong pointed external spur. Coxae II, III, and IV somewhat square with progressively smaller pointed external spurs, all coxae with few setae, syncoxae absent (
<figureCitation id="FACA1486FF864830FBE026AEB58FF910" box="[1128,1214,1741,1765]" captionStart="Fig" captionStartId="10.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-344@10.[339,1248,253,1199]" captionTargetPageId="10" captionText="Fig. 8 Scanncng electron mccrographs of Ixodes woyliei. Nsmph, legs and spcracular plate. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, c, 100 μm; b, 20 μm; d, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551888" httpUri="https://zenodo.org/record/11551888/files/figure.png" pageId="6" pageNumber="7">Figs. 8a</figureCitation>
and
<figureCitation id="FACA1486FF864830FB7426AEB428F910" box="[1276,1305,1741,1765]" captionStart="Fig" captionStartId="11.[167,198,1129,1148]" captionTargetBox="[151,1437,236,1161]" captionTargetId="figure-8@11.[339,1248,253,1112]" captionTargetPageId="11" captionText="Fig. 9 Lcne drawcng of Ixodes woyliei n. sp. Nsmph. a Capctulum ventral vcew. b Capctulum dorsal vcew. c Scutum. d Coxae. Scale-bars: 100 μm" figureDoi="http://doi.org/10.5281/zenodo.11551890" httpUri="https://zenodo.org/record/11551890/files/figure.png" pageId="6" pageNumber="7">9d</figureCitation>
). Length of tarsus I 0.22
<emphasis id="5085D411FF864830FC36268FB2FFF8F1" box="[958,974,1772,1796]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.30 with few long setae (&lt;40 μm) and some minute (&lt;10 μm) (
<figureCitation id="FACA1486FF864830FBB5276FB5B9F8D1" box="[1085,1160,1804,1828]" captionStart="Fig" captionStartId="10.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-344@10.[339,1248,253,1199]" captionTargetPageId="10" captionText="Fig. 8 Scanncng electron mccrographs of Ixodes woyliei. Nsmph, legs and spcracular plate. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, c, 100 μm; b, 20 μm; d, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551888" httpUri="https://zenodo.org/record/11551888/files/figure.png" pageId="6" pageNumber="7">Fig. 8c</figureCitation>
). Haller
<emphasis id="5085D411FF864830FB6A276FB5D7F8D1" box="[1250,1254,1804,1828]" italics="true" pageId="6" pageNumber="7"></emphasis>
s organ, anterior pit suboval with seven sensilla arranged in a cluster in the centre, posterior capsule opening slightly above the pit and divided by a low ridge with at least four sensilla visible (
<figureCitation id="FACA1486FF864830FC0827EFB2FDF851" box="[896,972,1932,1956]" captionStart="Fig" captionStartId="10.[167,198,1216,1235]" captionTargetBox="[151,1437,236,1274]" captionTargetId="figure-344@10.[339,1248,253,1199]" captionTargetPageId="10" captionText="Fig. 8 Scanncng electron mccrographs of Ixodes woyliei. Nsmph, legs and spcracular plate. a Spurs on coxae. b Spcracular plate. c Tarsus I. d Hallers organ. Scale-bars: a, c, 100 μm; b, 20 μm; d, 10 μm" figureDoi="http://doi.org/10.5281/zenodo.11551888" httpUri="https://zenodo.org/record/11551888/files/figure.png" pageId="6" pageNumber="7">Fig. 8b</figureCitation>
). Length of tarsus IV is 0.20
<emphasis id="5085D411FF864830FA8A27EFB423F851" box="[1282,1298,1932,1956]" italics="true" pageId="6" pageNumber="7"></emphasis>
0.30.
</paragraph>
</subSubSection>
<caption id="368E588BFF874831FF2F245CB074FB99" ID-DOI="http://doi.org/10.5281/zenodo.11551882" ID-Zenodo-Dep="11551882" httpUri="https://zenodo.org/record/11551882/files/figure.png" pageId="7" pageNumber="8" startId="7.[167,198,1087,1106]" targetBox="[151,1437,236,1145]" targetPageId="7" targetType="figure">
<paragraph id="624E0803FF874831FF2F245CB074FB99" blockId="7.[167,1399,1087,1132]" pageId="7" pageNumber="8">
<emphasis id="5085D411FF874831FF2F245CB1EDFBA7" bold="true" box="[167,220,1087,1106]" pageId="7" pageNumber="8">Fig. 5</emphasis>
Lcne drawcng of
<taxonomicName id="A5F17380FF874831FEF8245CB0EEFBA7" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[368,479,1087,1106]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF874831FEF8245CB0EEFBA7" box="[368,479,1087,1106]" italics="true" pageId="7" pageNumber="8">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FE6D245CB322FBA7" box="[485,531,1087,1106]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
Female.
<emphasis id="5085D411FF874831FDD7245CB35BFBA7" bold="true" box="[607,618,1087,1106]" pageId="7" pageNumber="8">a</emphasis>
Capctulum, ventral vcew.
<emphasis id="5085D411FF874831FCCB245CB27EFBA7" bold="true" box="[835,847,1087,1106]" pageId="7" pageNumber="8">b</emphasis>
Capctulum, dorsal vcew.
<emphasis id="5085D411FF874831FBAA245CB51DFBA7" bold="true" box="[1058,1068,1087,1106]" pageId="7" pageNumber="8">c</emphasis>
Scutum.
<emphasis id="5085D411FF874831FBF5245CB5B8FBA7" bold="true" box="[1149,1161,1087,1106]" pageId="7" pageNumber="8">d</emphasis>
Tarscs I.
<emphasis id="5085D411FF874831FB5A245CB5ECFBA7" bold="true" box="[1234,1245,1087,1106]" pageId="7" pageNumber="8">e</emphasis>
Tarscs IV.
<emphasis id="5085D411FF874831FAB8245CB406FBA7" bold="true" box="[1328,1335,1087,1106]" pageId="7" pageNumber="8">f</emphasis>
Coxae.
<emphasis id="5085D411FF874831FF2F243AB1CBFB99" box="[167,250,1113,1132]" italics="true" pageId="7" pageNumber="8">Scale-bars</emphasis>
: 200 μm
</paragraph>
</caption>
<subSubSection id="2AEB5B88FF87483EFF1F24CDB001F930" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="624E0803FF874831FF1F24CDB04BFB31" blockId="7.[151,775,1198,1957]" box="[151,378,1198,1220]" pageId="7" pageNumber="8">
<emphasis id="5085D411FF874831FF1F24CDB04BFB31" bold="true" box="[151,378,1198,1220]" pageId="7" pageNumber="8">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="624E0803FF874831FF1F24AEB5E2F891" blockId="7.[151,775,1198,1957]" lastBlockId="7.[813,1437,1196,1956]" pageId="7" pageNumber="8">
Morphologically,
<taxonomicName id="A5F17380FF874831FEEA24AEB0FDFB11" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[354,460,1228,1252]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei">
<emphasis id="5085D411FF874831FEEA24AEB0FDFB11" box="[354,460,1228,1252]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
logically conforms to the subgenus
<taxonomicName id="A5F17380FF874831FE85248FB0AEFAF1" authorityName="Schulze" authorityYear="1935" box="[269,415,1260,1284]" class="Arachnida" family="Ixodidae" genus="Endopalpiger" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="5085D411FF874831FE85248FB0AEFAF1" box="[269,415,1260,1284]" italics="true" pageId="7" pageNumber="8">Endopalpiger</emphasis>
</taxonomicName>
as described by Roberts [
<bibRefCitation id="066075F2FF874831FD6A248EB3CEFAF0" author="Roberts FSH" box="[738,767,1261,1285]" pageId="7" pageNumber="8" refId="ref9502" refString="12. Roberts FSH. Australcan Tccks. Melbourne: Commonwealth Sccentcfcc and Industrcal Research Organcsatcon; 1970." type="book" year="1970">12</bibRefCitation>
] due to the enlarged palpal article 1 that extends inwardly to partially ensheathe the base of the mouthparts; common to all females within this subgenus. However it is also pertinent to consider the species of
<taxonomicName id="A5F17380FF874831FF1F25EFB0F5FA50" authority="Schulze, 1935" authorityName="Schulze" authorityYear="1935" box="[151,452,1420,1445]" class="Arachnida" family="Ixodidae" genus="Exopalpiger" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="5085D411FF874831FF1F25EFB028FA51" box="[151,281,1420,1444]" italics="true" pageId="7" pageNumber="8">Exopalpiger</emphasis>
Schulze, 1935
</taxonomicName>
(
<taxonomicName id="A5F17380FF874831FE5125EEB371FA51" authorityName="Warburton &amp; Nuttall" authorityYear="1909" box="[473,576,1420,1444]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fecialis">
<emphasis id="5085D411FF874831FE5125EEB371FA51" box="[473,576,1420,1444]" italics="true" pageId="7" pageNumber="8">I. fecialis</emphasis>
</taxonomicName>
,
<taxonomicName id="A5F17380FF874831FDDC25EEB02EFA30" authority="Neumann, 1908" authorityName="Neumann" authorityYear="1908" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="5085D411FF874831FDDC25EEB3F3FA51" box="[596,706,1421,1444]" italics="true" pageId="7" pageNumber="8">I. vestitus</emphasis>
Neumann, 1908
</taxonomicName>
and
<taxonomicName id="A5F17380FF874831FEEB25CEB3A2FA30" authority="Roberts, 1960" authorityName="Roberts" authorityYear="1960" box="[355,659,1452,1477]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="antechini">
<emphasis id="5085D411FF874831FEEB25CEB0D7FA31" box="[355,486,1452,1476]" italics="true" pageId="7" pageNumber="8">I. antechini</emphasis>
Roberts, 1960
</taxonomicName>
) as more recently the species of these two subgenera have been amalgamated into
<taxonomicName id="A5F17380FF874831FEF2258FB0CDF9F1" authorityName="Schulze" authorityYear="1935" box="[378,508,1516,1540]" class="Arachnida" family="Ixodidae" genus="Exopalpiger" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="5085D411FF874831FEF2258FB0CDF9F1" box="[378,508,1516,1540]" italics="true" pageId="7" pageNumber="8">Exopalpiger</emphasis>
</taxonomicName>
[
<bibRefCitation id="066075F2FF874831FD9E258EB307F9F0" author="Camccas JL &amp; Hervs JP &amp; Adam F &amp; Morel PC" box="[534,566,1517,1541]" pageId="7" pageNumber="8" refId="ref9986" refString="24. Camccas JL, Hervs JP, Adam F, Morel PC. [The tccks of the world. Nomenclature, descrcbed stages, hosts, dcstrcbutcon (Acarcda, Ixodcdca)]. Parcs: Orstom Edctcons Publcsher; 1998. (In French)." type="book" year="1998">24</bibRefCitation>
]. The species of these two subgenera however can be quite easily differentiated by the palpal article 1 which in
<taxonomicName id="A5F17380FF874831FD14264FB023F990" pageId="7" pageNumber="8">
<emphasis id="5085D411FF874831FD14264FB186F991" italics="true" pageId="7" pageNumber="8">Exopalpiger</emphasis>
species
</taxonomicName>
is enlarged but does not extend inwardly, as seen in
<taxonomicName id="A5F17380FF874831FE9D260EB04BF971" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[277,378,1644,1668]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FE9D260EB04BF971" box="[277,378,1644,1668]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FE0D260EB0F2F970" box="[389,451,1645,1669]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
and the other
<taxonomicName id="A5F17380FF874831FDFD260FB1DDF951" class="Arachnida" family="Ixodidae" genus="Endopalpiger" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5085D411FF874831FDFD260FB236F971" box="[629,775,1644,1668]" italics="true" pageId="7" pageNumber="8">Endopalpiger</emphasis>
species.
</taxonomicName>
Further morphological differences are outlined in
<tableCitation id="2F733DB8FF874831FF7226CEB061F930" box="[250,336,1709,1733]" captionStart="Table 2" captionStartId="11.[151,210,1438,1459]" captionTargetBox="[151,1437,1534,1897]" captionTargetPageId="11" captionText="Table 2 Macn morphologccal dcfferences between adult female (F) and nsmph (N) lcfe stages of Ixodes woyliei and descrcbed specces of Exopalpiger and Endopalpiger. Morphologccal dcfferences between lcfe stages are noted wcth (F) or (N); otherwcse tracts are the same for both" httpUri="http://table.plazi.org/id/368E588BFF8B483DFF1F25FDB011FA1B" pageId="7" pageNumber="8" tableUuid="368E588BFF8B483DFF1F25FDB011FA1B">Table 2</tableCitation>
. Differentiation of adult females from the other four
<taxonomicName id="A5F17380FF874831FEC626AFB30DF910" box="[334,572,1740,1765]" pageId="7" pageNumber="8">
<emphasis id="5085D411FF874831FEC626AFB0EEF911" box="[334,479,1740,1764]" italics="true" pageId="7" pageNumber="8">Endopalpiger</emphasis>
species
</taxonomicName>
can be achieved by the presence of an open anal groove, the large pointed spurs on each coxa, presence of syncoxae, the large number of denticles on the hypostome, and the shape of the scutum (
<tableCitation id="2F733DB8FF874831FE14272FB0C7F891" box="[412,502,1868,1892]" captionStart="Table 2" captionStartId="11.[151,210,1438,1459]" captionTargetBox="[151,1437,1534,1897]" captionTargetPageId="11" captionText="Table 2 Macn morphologccal dcfferences between adult female (F) and nsmph (N) lcfe stages of Ixodes woyliei and descrcbed specces of Exopalpiger and Endopalpiger. Morphologccal dcfferences between lcfe stages are noted wcth (F) or (N); otherwcse tracts are the same for both" httpUri="http://table.plazi.org/id/368E588BFF8B483DFF1F25FDB011FA1B" pageId="7" pageNumber="8" tableUuid="368E588BFF8B483DFF1F25FDB011FA1B">Table 2</tableCitation>
). Specifically, the presence of an open anal groove, the lacc of syncoxae and greater dentition differentiates
<taxonomicName id="A5F17380FF874831FDCE27EEB383F851" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[582,690,1932,1956]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FDCE27EEB383F851" box="[582,690,1932,1956]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FD4B27EEB236F850" box="[707,775,1933,1957]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
from
<taxonomicName id="A5F17380FF874831FCF824CEB526FB31" authorityName="Neumann" authorityYear="1904" box="[880,1047,1196,1220]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="5085D411FF874831FCF824CEB526FB31" box="[880,1047,1196,1220]" italics="true" pageId="7" pageNumber="8">I. australiensis</emphasis>
</taxonomicName>
, while the presence of spurs on the coxae (armed) differentiates
<taxonomicName id="A5F17380FF874831FB2024AEB43EFB11" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1192,1295,1228,1252]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FB2024AEB43EFB11" box="[1192,1295,1228,1252]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FA9324AEB46AFB10" box="[1307,1371,1229,1253]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
from
<taxonomicName id="A5F17380FF874831FCA5248EB297FAF1" authorityName="Neumann" authorityYear="1899" box="[813,934,1261,1284]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tasmani">
<emphasis id="5085D411FF874831FCA5248EB297FAF1" box="[813,934,1261,1284]" italics="true" pageId="7" pageNumber="8">I. tasmani</emphasis>
</taxonomicName>
and
<taxonomicName id="A5F17380FF874831FC7A248EB5A8FAF1" authorityName="Swan" authorityYear="1931" box="[1010,1177,1260,1284]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hydromyidis">
<emphasis id="5085D411FF874831FC7A248EB5A8FAF1" box="[1010,1177,1260,1284]" italics="true" pageId="7" pageNumber="8">I. hydromyidis</emphasis>
</taxonomicName>
, both of which lacc spurs. The most morphologically similar species to
<taxonomicName id="A5F17380FF874831FA06256EB248FAB1" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FA06256EB248FAB1" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FC0C254EB2F3FAB0" box="[900,962,1325,1349]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
is
<taxonomicName id="A5F17380FF874831FC61254EB54CFAB1" authorityName="Nuttall" authorityYear="1916" box="[1001,1149,1325,1348]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="victoriensis">
<emphasis id="5085D411FF874831FC61254EB54CFAB1" box="[1001,1149,1325,1348]" italics="true" pageId="7" pageNumber="8">I. victoriensis</emphasis>
</taxonomicName>
; however these two species can be readily differentiated by dentition and the shape of the scutum, spurs on the coxae, and palpal article 1.
<taxonomicName id="A5F17380FF874831FC2825EFB571FA51" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[928,1088,1420,1444]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="sp. nov.">
<emphasis id="5085D411FF874831FC2825EFB571FA51" box="[928,1088,1420,1444]" italics="true" pageId="7" pageNumber="8">Ixodes woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FBC625EFB5BEFA51" box="[1102,1167,1420,1444]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
has a remarcable and complex dentition with small overcrowded denticles at the corona (12/12), dropping in number but increasing in size to mainly a 6/6 and 5/5 dentition whereas
<taxonomicName id="A5F17380FF874831FCA5266EB288F9D1" authorityName="Nuttall" authorityYear="1916" box="[813,953,1549,1572]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="victoriensis">
<emphasis id="5085D411FF874831FCA5266EB288F9D1" box="[813,953,1549,1572]" italics="true" pageId="7" pageNumber="8">I. victoriensis</emphasis>
</taxonomicName>
dentition is mostly 5/5, with rows of 4/4 at both anterior and posterior ends [
<bibRefCitation id="066075F2FF874831FB22264FB5F6F9B1" author="Weaver H." box="[1194,1223,1580,1604]" pageId="7" pageNumber="8" pagination="820 - 9" refId="ref9734" refString="17. Weaver H. Redescrcptcon of Ixodes victoriensis Nuttall, 1916 (Ixodcda: Ixodcdae) from marsupcals cn Vcctorca, Australca. Ssst Appl Acarol. 2016; 21 (6): 820 - 9." type="journal article" year="2016">17</bibRefCitation>
]. The shape of the scutum in
<taxonomicName id="A5F17380FF874831FC2B262EB532F991" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[931,1027,1612,1636]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FC2B262EB532F991" box="[931,1027,1612,1636]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FB84262FB576F991" box="[1036,1095,1612,1636]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
is longer than that of
<taxonomicName id="A5F17380FF874831FAC9262EB257F971" authorityName="Nuttall" authorityYear="1916" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="victoriensis">
<emphasis id="5085D411FF874831FAC9262EB257F971" italics="true" pageId="7" pageNumber="8">I. victoriensis</emphasis>
</taxonomicName>
(about as long as wide
<emphasis id="5085D411FF874831FBEB260EB549F971" box="[1123,1144,1645,1668]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
wider than long for
<taxonomicName id="A5F17380FF874831FADD260EB27CF951" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei">
<emphasis id="5085D411FF874831FADD260EB27CF951" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
), and appears more angular. The coxae of
<taxonomicName id="A5F17380FF874831FAA926EEB4B1F951" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1313,1408,1676,1700]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FAA926EEB4B1F951" box="[1313,1408,1676,1700]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FA0026EFB27BF931" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
are all armed with large, pointed spurs and lacc syncoxae, while
<taxonomicName id="A5F17380FF874831FC3526AEB57BF911" authorityName="Nuttall" authorityYear="1916" box="[957,1098,1741,1764]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="victoriensis">
<emphasis id="5085D411FF874831FC3526AEB57BF911" box="[957,1098,1741,1764]" italics="true" pageId="7" pageNumber="8">I. victoriensis</emphasis>
</taxonomicName>
coxae are armed with smaller spurs that are not as pointed and possess syncoxae. The enlarged palpal article 1 described for
<taxonomicName id="A5F17380FF874831FB46276EB41CF8D1" authority="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson, 2017" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[1230,1325,1804,1828]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="woyliei" status="n. sp.">
<emphasis id="5085D411FF874831FB46276EB41CF8D1" box="[1230,1325,1804,1828]" italics="true" pageId="7" pageNumber="8">I. woyliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BB6696AFF874831FABE276FB441F8D1" box="[1334,1392,1804,1828]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
has a posterolateral prominence macing it more widely rectangular than that seen on
<taxonomicName id="A5F17380FF874831FBCC272EB5FCF891" authorityName="Nuttall" authorityYear="1916" box="[1092,1229,1869,1892]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="victoriensis">
<emphasis id="5085D411FF874831FBCC272EB5FCF891" box="[1092,1229,1869,1892]" italics="true" pageId="7" pageNumber="8">I. victoriensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="624E0803FF87483EFCCA270FB001F930" blockId="7.[813,1437,1196,1956]" lastBlockId="8.[151,775,1580,1733]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Differential diagnosis of the nymphal stage can be largely achieved as for the adult female (
<tableCitation id="2F733DB8FF874831FB4A27EFB42AF851" box="[1218,1307,1932,1956]" captionStart="Table 2" captionStartId="11.[151,210,1438,1459]" captionTargetBox="[151,1437,1534,1897]" captionTargetPageId="11" captionText="Table 2 Macn morphologccal dcfferences between adult female (F) and nsmph (N) lcfe stages of Ixodes woyliei and descrcbed specces of Exopalpiger and Endopalpiger. Morphologccal dcfferences between lcfe stages are noted wcth (F) or (N); otherwcse tracts are the same for both" httpUri="http://table.plazi.org/id/368E588BFF8B483DFF1F25FDB011FA1B" pageId="7" pageNumber="8" tableUuid="368E588BFF8B483DFF1F25FDB011FA1B">Table 2</tableCitation>
).
<taxonomicName id="A5F17380FF874831FAB027EFB4ACF851" box="[1336,1437,1932,1956]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="minor">A minor</taxonomicName>
exception involves
<taxonomicName id="A5F17380FF88483EFEEB264EB0CAF9B1" authorityName="Neumann" authorityYear="1904" box="[355,507,1580,1604]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="5085D411FF88483EFEEB264EB0CAF9B1" box="[355,507,1580,1604]" italics="true" pageId="8" pageNumber="9">I. australiensis</emphasis>
</taxonomicName>
whereby the nymph has an anal groove which remains open, but not as widely as seen in
<taxonomicName id="A5F17380FF88483EFF7B260EB065F971" authorityName="Ash &amp; Ellcot &amp; Godfres &amp; Burmej &amp; Abdad &amp; Northover &amp; Wasne &amp; Morrcs &amp; Clode &amp; Lsmbers &amp; Andrew Thompson" authorityYear="2017" box="[243,340,1644,1668]" class="Arachnida" family="Ixodidae" genus="Ixodes" kingdom="Animalia" order="Ixodida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="woyliei">
<emphasis id="5085D411FF88483EFF7B260EB065F971" box="[243,340,1644,1668]" italics="true" pageId="8" pageNumber="9">I. woyliei</emphasis>
</taxonomicName>
nymphs. However the shape of palpal article one, spurs and presence of syncoxae allow differentiation.
</paragraph>
</subSubSection>
</treatment>
</document>