152 lines
17 KiB
XML
152 lines
17 KiB
XML
<document id="B053729F1C88F41620433AD9BD580DF3" ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZooBank="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas & Leschen, Richard A. B." docDate="2017" docId="DA39A95003381D8C9A8563E27AE9D513" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Hypocaccus (Baeckmanniolus) varians subsp. varians Schmidt 1890" docType="treatment" docVersion="6" lastPageNumber="41" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="40" updateTime="1701316739810" updateUser="plazi">
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<mods:title id="EEA3B944230794F0FD5A06BB086CDE5D">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="32EA997D669604271915692E1E577C21">Lackner, Tomas</mods:namePart>
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<mods:namePart id="7C77989E94ADD03254A42283D1BA88ED">Leschen, Richard A. B.</mods:namePart>
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<mods:typeOfResource id="E75BB9BD098DBA7BE245A11C7EED4F6A">text</mods:typeOfResource>
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<mods:date id="7F9CD1F121A3FD905137A2BE783BCDCA">2017</mods:date>
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<mods:number id="FF1B7AE8D7D62E0758D29B62C002D21B">689</mods:number>
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<mods:start id="EB5D84CBF0D305E71243E2A4AEA3A0B5">1</mods:start>
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<mods:classification id="CA96C029A45A918FB573810EADB955D6">journal article</mods:classification>
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<mods:identifier id="7D43B17FDFBD0C8ED8E849E1D4CDCBF7" type="DOI">http://dx.doi.org/10.3897/zookeys.689.12021</mods:identifier>
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<treatment id="DA39A95003381D8C9A8563E27AE9D513" ID-GBIF-Taxon="132648718" LSID="urn:lsid:plazi:treatment:DA39A95003381D8C9A8563E27AE9D513" httpUri="http://treatment.plazi.org/id/DA39A95003381D8C9A8563E27AE9D513" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection id="CFDD29E3C6A9E6E65263312CE0F604F6" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="E417A523704AF27FAFE159C6269BDE91" pageId="39" pageNumber="40">
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<taxonomicName id="9D3A3F9BAF1172DFFD7D9EE8F4A199D5" ID-CoL="5J342" authority="Schmidt, 1890" authorityName="Schmidt" authorityYear="1890" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Baeckmanniolus) varians" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="varians" subGenus="Baeckmanniolus" subSpecies="varians">Hypocaccus (Baeckmanniolus) varians varians (Schmidt, 1890)</taxonomicName>
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Figs 183, 184-192, 193-201, 756
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</paragraph>
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</subSubSection>
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<subSubSection id="E0FB7CB89A0F07011E40A56C6D52EBFA" pageId="39" pageNumber="40" type="reference_group">
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<paragraph id="2C9F1EF996752D38884F68B6B188E450" pageId="39" pageNumber="40">
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<taxonomicName id="F8953C48048D7B1FC618000914428883" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus varians" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="varians">Saprinus varians</taxonomicName>
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Schmidt, 1890: 55.
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</paragraph>
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</subSubSection>
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<subSubSection id="28C350A35731833233FA5BC5932CBA93" pageId="39" pageNumber="40" type="type locality">
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<paragraph id="66B91581C5659B4C4B890292CB8DCD8A" pageId="39" pageNumber="40">Type locality.</paragraph>
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<paragraph id="37BF62560B27A876DDBEF133C7F35C3F" pageId="39" pageNumber="40">Japan.</paragraph>
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</subSubSection>
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<subSubSection id="6C29FE80F90327EFEEF6DAAA7D6CF8F6" pageId="39" pageNumber="40" type="type material">
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<paragraph id="A166C67DEFCEFA67E26EF8D13E4E8A48" pageId="39" pageNumber="40">Type material examined.</paragraph>
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<paragraph id="870203FE9AA8C37B5857E0D04C6B4B1F" pageId="39" pageNumber="40">
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<taxonomicName id="A60C28F66DA2586FF6CC970AA063FE76" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus varians" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="varians">Saprinus varians</taxonomicName>
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Schmidt, 1890: Lectotype, present designation: ♂, glued on the mounting card, with the following labels:
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<normalizedToken id="B5E70DEF057421CF04AFD4B5BCBDC6AB" originalValue="“Japan”">"Japan"</normalizedToken>
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(written); followed by:
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<normalizedToken id="F652E034E97E78F792523B6F1948DC0A" originalValue="“Type”">"Type"</normalizedToken>
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(brick-red label, printed); followed by: "coll. J. Schmidt" (printed); followed by: "
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<taxonomicName id="F73785462FB8A0F62C7FD9647A398A3F" lsidName="varians" pageId="39" pageNumber="40" rank="species" species="varians">varians</taxonomicName>
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/ m." (black-yellow margined label, written); followed by: "
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<taxonomicName id="393CAFDD451CF032E5DFC8173681B9C2" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="9ABEFCF0057CB39407A9AE466064103D" lsidName="varians" pageId="39" pageNumber="40" rank="species" species="varians">varians</taxonomicName>
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Schmidt / Coll. Schmidt-Bickhardt" (printed); followed by: "
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<taxonomicName id="BB0EAEE9D7FC97EE3F97350CB80A5906" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus varians" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="varians">Saprinus varians</taxonomicName>
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/ Schmidt, 1890 / LECTOTYPE 2014 / des. Lackner & Leschen" (red label, written) (ZMHUB). Paralectotypes, present designation: 4 specs., probably ♀♀, carrying the same hand-written labels
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<normalizedToken id="5A2CF57FC7A5C0FB3526F2787535E410" originalValue="“Japan”">"Japan"</normalizedToken>
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and
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<normalizedToken id="37A793C977FE2F47589D563856FB7132" originalValue="“Type”">"Type"</normalizedToken>
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and the printed label "
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<taxonomicName id="CF6F3ED31D8234D5BFFEDA204F99C063" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="0E76E482633739F9B95E837E9802B4BF" lsidName="varians" pageId="39" pageNumber="40" rank="species" species="varians">varians</taxonomicName>
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Schmidt / Coll. Schmidt-Bickhardt", followed by the paralectotype labels identical to that of the lectotype (ZMHUB). This species has been described from unknown number of specimens and the lectotype designation fixes the taxonomic identity of the species.
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</paragraph>
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<caption id="7B8C9076D26615EE90073A241FDF3B71" pageId="39" pageNumber="40">
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<paragraph id="33A39DD579F0836CF96E5617C237F1DC" pageId="39" pageNumber="40">
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Figures 184-192. 184
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<taxonomicName id="C25EFD3A23C45947B39CFB8354DB06C8" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Baeckmanniolus) varians" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="varians" subGenus="Baeckmanniolus" subSpecies="varians">Hypocaccus (Baeckmanniolus) varians varians</taxonomicName>
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(Schmidt, 1890) head, dorsal view 185 antennal club, ventral view 186 mentum, ventral view 187 propygidium + pygidium 188 prosternum 189 mesoventrite 190 lateral disc of metaventrite + metepisternum 191 protibia, dorsal view 192 ditto, ventral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="214A14AFB502162115C9D6A7D922D56D" pageId="40" pageNumber="41" type="materials_examined">
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<paragraph id="9B540981DCF2C6EEE039ACC323C745F6" pageId="40" pageNumber="41">
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<pageBreakToken id="C8CC3008487640B6CA27A51B87282FF9" pageId="40" pageNumber="41" start="start">Additional</pageBreakToken>
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material examined.
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</paragraph>
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<paragraph id="6C45A7317385818782F21A6E5EA85983" pageId="40" pageNumber="41">AUSTRALIA. Western Australia: 1 spec., Monte Bello Island (ANIC). Northern Territory: 1 ♂ & 1 ♀, Groote Eylandt, N.B. Tindale (SAMA). Queensland: 1 ♀, Cairns (SAMA).</paragraph>
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</subSubSection>
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<subSubSection id="F7245B25FEB30761CC9A308A9FEFCB31" pageId="40" pageNumber="41" type="biology_ecology">
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<paragraph id="E9F8DACA65C990660BA6E0AFE82F135F" pageId="40" pageNumber="41">Biology.</paragraph>
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<paragraph id="3430D552C7FC7F46DEF22A7DAB93F21C" pageId="40" pageNumber="41">See above.</paragraph>
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<caption id="343C48BF72CC7CF8805A925A57CE26DF" pageId="40" pageNumber="41">
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<paragraph id="3909AA79208730AFEF53566E7ACF5903" pageId="40" pageNumber="41">
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Figures 193-201. 193
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<taxonomicName id="A5E992DC1B6F2718265B030D3E985405" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Baeckmanniolus) varians" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="varians" subGenus="Baeckmanniolus" subSpecies="varians">Hypocaccus (Baeckmanniolus) varians varians</taxonomicName>
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(Schmidt, 1890) male terminalia: 8th sternite + 8th tergite, ventral view 194 ditto, dorsal view 195 ditto, lateral view 196 male terminalia: 9th + 10th tergites, dorsal view 197 ditto, lateral view 198 male terminalia: spiculum gastrale, ventral view 199 ditto, lateral view 200 male terminalia: aedeagus, dorsal view 201 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="31EDAAB4D0566A41A10199FBDEF34AD5" pageId="40" pageNumber="41" type="distribution">
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<paragraph id="BE2E8928FB13228156AF6DB0EFCB8D58" pageId="40" pageNumber="41">Distribution.</paragraph>
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<paragraph id="2FF4EF78BE6CD330EB669C9D821B7621" pageId="40" pageNumber="41">
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Described from Japan, the nominotypical subspecies is found also on Sakhalin Island, Taiwan, Philippines, Vietnam, Sri Lanka, and Australia. The two other subspecies are
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<taxonomicName id="42CBB2552C781B309BA8A50F5DC72865" lsidName="H. (B.) varians" pageId="40" pageNumber="41" rank="subSpecies" species="varians" subGenus="B." subSpecies="hatsune">H. (B.) varians hatsune</taxonomicName>
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Nakane, 1977, described from the Ogasawara Islands (southern Japan) and
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<taxonomicName id="02CF90585DA1F65881A326A7C4EF4149" lsidName="H. (B.) varians" pageId="40" pageNumber="41" rank="subSpecies" species="varians" subGenus="B." subSpecies="continentalis">H. (B.) varians continentalis</taxonomicName>
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Reichardt, 1941 described from Russian Far East (Primorsky Kray) and found also in eastern China (Shandong) (
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<bibRefCitation id="8BAD6E8CC8E1FFFA5FAEB49B21E2F9B7" pageId="40" pageNumber="41">Mazur 2011</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="23FD7D3C2E71BAB199F23F0FA60AF425" pageId="40" pageNumber="41" type="remarks">
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<paragraph id="A83FE31261B5AD297B5007BBA390E1D4" pageId="40" pageNumber="41">Remarks.</paragraph>
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<paragraph id="378257784FCA0C24AE0C134B741FD066" pageId="40" pageNumber="41">
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The species
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<taxonomicName id="FFBF966C591BAF449B05842CBE980FB3" lsidName="H. (B.) varians" pageId="40" pageNumber="41" rank="species" species="varians" subGenus="B.">H. (B.) varians</taxonomicName>
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has a faint pronotal punctation indicating that it should be transferred into the nominotypical subgenus
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<taxonomicName id="E7A95DACAE0699CECC3B02EAD07B4FB4" lsidName="(Hypocaccus)" pageId="40" pageNumber="41" rank="subGenus" subGenus="Hypocaccus">Hypocaccus</taxonomicName>
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. According to
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<bibRefCitation id="05888E7BF5564D546BED8410FCA9169D" author="Bousquet, Y" journalOrPublisher="NRC Research Press, Ottawa, Ontario, Canada" pageId="213" pageNumber="214" title="The Insects and Arachnids of Canada. Part 24. ColeopteraHisteridae." year="2006">Bousquet and Laplante (2006</bibRefCitation>
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; see above) the outer margin of metatibia of the subgenus
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<taxonomicName id="08D730120D249B445C1195F6720156A0" lsidName="(Baeckmanniolus)" pageId="40" pageNumber="41" rank="subGenus" subGenus="Baeckmanniolus">Baeckmanniolus</taxonomicName>
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has three rows of denticles instead of two rows present in
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<taxonomicName id="689B539A342AD15CC3B7DBE994F6EDAA" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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. The metatibia of
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<taxonomicName id="5072750FF0B597D8E257ED2FBF7698DA" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (B.) varians" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="varians" subGenus="B.">Hypocaccus (B.) varians</taxonomicName>
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possesses two rows on the metatibia supplemented by another 2 (occasionally 3) denticles between the two rows. The presence of vague and weakened pronotal punctation and by possessing denticles between the two rows on the outer margin of metatibia, this species represents a transitional form between the two subgenera and its exact taxonomic position should be determined by a thorough morphological study of all known species of
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<taxonomicName id="DFC370EB8A66E25A522E9FFBBAF5DDED" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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s. l.
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<taxonomicName id="1FC95B27AB546FB7AC03B4573A6FFB7F" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (B.) varians" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="varians" subGenus="B." subSpecies="varians">Hypocaccus (B.) varians varians</taxonomicName>
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is diagnosed above, as well as provided with a differential diagnosis that separates it from other Australopacific taxa. Most important differences between the members of the genus
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<taxonomicName id="D32CAB7D42DBAB6D788DDA2D7C6F33DF" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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are outlined in the key to the subgenera above. We chose not to fully re-describe
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<taxonomicName id="E5304203CB1A540473E94D5F4A84BFFD" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (B.) varians" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="varians" subGenus="B." subSpecies="varians">Hypocaccus (B.) varians varians</taxonomicName>
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here, leaving its re-description to the revision of the genus
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<taxonomicName id="E569485C11BD107E53351C2DA848013C" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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. For the sake of the better species recognition, however, we decided to depict it here, including its male terminalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |