166 lines
18 KiB
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166 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.60525" ID-GBIF-Dataset="217bea89-5bd1-4ae7-bfce-fb945a933fc4" ID-GBIF-Taxon="176088088" ID-Pensoft-Pub="2535-0730-2-119" ID-Pensoft-UUID="6301B37BBB7D51A185FB132ED24E4E81" ID-ZooBank="8BD62DBF438C46508C4037C837FD573F" LSID="urn:lsid:zoobank.org:act:AA3F0556-CAB5-477C-A42B-0AFFC9E84561" ModsDocID="2535-0730-2-119" checkinTime="1608157809646" checkinUser="pensoft" docAuthor="Ingrisch, Sigfrid" docDate="2020" docId="C56666737F0258B4B4E862C87B96C192" docLanguage="en" docName="EvolutSyst 4(2): 119-132" docOrigin="Evolutionary Systematics 4 (2)" docPubDate="2020-12-15" docSource="http://dx.doi.org/10.3897/evolsyst.4.60525" docTitle="Liara (Acanthocoryphus) durata Ingrisch 2020, sp. nov." docType="treatment" docUuid="AA3F0556-CAB5-477C-A42B-0AFFC9E84561" docUuidSource="ZooBank" docVersion="3" id="6301B37BBB7D51A185FB132ED24E4E81" lastPageNumber="119" masterDocId="6301B37BBB7D51A185FB132ED24E4E81" masterDocTitle="New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections" masterLastPageNumber="132" masterPageNumber="119" pageId="0" pageNumber="119" updateTime="1643451831480" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingrisch, Sigfrid</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8624-0472</mods:nameIdentifier>
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<mods:affiliation>Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier type="email">s.ingrisch@leibniz-zfmk.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>119</mods:start>
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<mods:end>132</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.4.60525</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.4.60525</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-119</mods:identifier>
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<mods:identifier type="ZooBank">8BD62DBF438C46508C4037C837FD573F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6301B37BBB7D51A185FB132ED24E4E81</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="176088088" LSID="urn:lsid:zoobank.org:act:AA3F0556-CAB5-477C-A42B-0AFFC9E84561" httpUri="http://treatment.plazi.org/id/C56666737F0258B4B4E862C87B96C192" lastPageNumber="119" pageId="0" pageNumber="119">
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<subSubSection pageId="0" pageNumber="119" type="nomenclature">
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<paragraph pageId="0" pageNumber="119">
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<taxonomicName LSID="http://zoobank.org/AA3F0556-CAB5-477C-A42B-0AFFC9E84561" authority="Ingrisch, 2020" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara (Acanthocoryphus) durata" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="durata" status="sp. nov." subGenus="Acanthocoryphus">Liara (Acanthocoryphus) durata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="119">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">Fig. 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="holotype">
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<paragraph pageId="0" pageNumber="119">Holotype</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">(male).</emphasis>
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Hongkong [without precision], in or before 1911, coll. F.W. Terry - depository: BMNH (Lo031S001; B.M.1911-359). The type is unique.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="etymology">
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<paragraph pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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The new species is named for the sclerotised central convexity of the male titillators, modified from Latin
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<emphasis italics="true" pageId="0" pageNumber="119">durare</emphasis>
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to harden, hardened.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="diagnosis">
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<paragraph pageId="0" pageNumber="119">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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The titillators of the new species are intermediate between those of the subgenera
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<taxonomicName authorityName="Redtenbacher" authorityYear="1891" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Liara</emphasis>
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</taxonomicName>
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s.str. and
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<taxonomicName authorityName="Kocak & Kemal" authorityYear="2009" class="Insecta" family="Tettigoniidae" genus="Unalianus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Unalianus" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Unalianus</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Koçak">Kocak</normalizedToken>
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& Kemal, 2009 in that they are fused as in
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<taxonomicName authorityName="Kocak & Kemal" authorityYear="2009" class="Insecta" family="Tettigoniidae" genus="Unalianus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Unalianus" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Unalianus</emphasis>
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</taxonomicName>
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but of general shape as in
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<taxonomicName authorityName="Redtenbacher" authorityYear="1891" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Liara</emphasis>
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</taxonomicName>
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; the membranous lateral expansions are sclerotised but the apical areas not as strongly prolonged as in
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<taxonomicName authorityName="Kocak & Kemal" authorityYear="2009" class="Insecta" family="Tettigoniidae" genus="Unalianus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Unalianus" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Unalianus</emphasis>
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</taxonomicName>
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. Judging from the shapes of the cerci and titillators, the new species comes close to
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<taxonomicName authorityName="Ingrisch" authorityYear="1998" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara (Acanthocoryphus) brevis" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="brevis" subGenus="Acanthocoryphus">Liara (Acanthocoryphus) brevis</taxonomicName>
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Ingrisch, 1998. It differs by the long and upcurved fastigium verticis which is almost twice as long as the scapus instead of only little longer and has a strong tubercle at base. Further differences are the fused titillators, the very small styli, and shorter wings. The cerci are similar in both species but in the new species less wide at base and the apical branches more curved.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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A short and stout, strongly brachypterous species (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4A</figureCitation>
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). Fastigium verticis much longer than scapus, strongly upcurved; with a large tubercle at top near base, immediately followed by a smaller tubercle anteriorly; apex acute; separated from fastigium frontis by a narrow seam (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4C, D</figureCitation>
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). Frons rugose (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4B</figureCitation>
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). Pronotum rugose, short funnel-shaped, only in apical area little shouldered; anterior margin in middle truncate; posterior margin truncate; transverse sulci only indicated; paranota with anterior angle rounded, posterior angle broadly rounded; auditory swelling very weak; humeral sinus weak. Tegmen surpassing apex of third abdominal tergite, hind wings of almost same length (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4A</figureCitation>
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). Prosternum mute; mesosternal lobes long and stout; metasternal lobes obtuse; intermedial plate with small spines at posterior angles (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4F</figureCitation>
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). Femora with the following number of spines on ventral margins: profemur no external, 2-3 internal; mesofemur 4 external, no internal; postfemur 6-7 external, no internal. Knee lobes of profemur obtuse on external, short spinose on internal side; of mesofemur acute on external, spinose on internal side; of postfemur spinose on both sides.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 4.</emphasis>
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<taxonomicName authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara (Acanthocoryphus) durata" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="durata" subGenus="Acanthocoryphus">Liara (Acanthocoryphus) durata</taxonomicName>
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sp. nov. male holotype:
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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habitus lateral view;
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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face,
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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head dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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fastigium verticis lateral view;
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<emphasis bold="true" pageId="0" pageNumber="119">E</emphasis>
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stridulatory file on underside of left tegmen;
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<emphasis bold="true" pageId="0" pageNumber="119">F</emphasis>
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thoracic sternites, the arrow points at one of the prosternal spines;
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<emphasis bold="true" pageId="0" pageNumber="119">G-I</emphasis>
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titillators of male phallus in ventro-apical view (
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<emphasis bold="true" pageId="0" pageNumber="119">G</emphasis>
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), do. as seen from base (
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<emphasis bold="true" pageId="0" pageNumber="119">H</emphasis>
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), do. in oblique lateral view (
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<emphasis bold="true" pageId="0" pageNumber="119">I</emphasis>
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);
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<emphasis bold="true" pageId="0" pageNumber="119">J, K</emphasis>
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male cercus in external (
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<emphasis bold="true" pageId="0" pageNumber="119">J</emphasis>
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) and internal (
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<emphasis bold="true" pageId="0" pageNumber="119">K</emphasis>
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) views.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Male.</emphasis>
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Tenth abdominal tergite little prolonged; apical margin converging on both sides, broadly truncate in middle. Epiproct triangularly rounded with shallow medial furrow. Paraprocts with short obtuse compressed projections. Cerci short, at apex divided into a broad falcate dorsal and a narrow falcate ventral projection, both curved up; on internal side with a weak furrow from dividing point of apical projections to base (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4J, K</figureCitation>
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). Subgenital plate with straight converging lateral margins; apex broad, deep triangularly excised; styli very small. Titillators fused in middle; basal areas furrowed, pointing laterad; fused area at apex faintly bilobate; lateral expansions of central parts ("lateral sacculi" in
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<bibRefCitation author="Ingrisch, S" journalOrPublisher="Courier Forschungsinstitut Senckenberg" pageId="0" pageNumber="119" pagination="1 - 387" refId="B8" refString="Ingrisch, S, 1998. Monograph of the Oriental Agraeciini (Insecta, Ensifera, Tettigoniidae): Taxonomic revision, phylogeny, stridulation, and development. Courier Forschungsinstitut Senckenberg 206: 1 - 387" title="Monograph of the Oriental Agraeciini (Insecta, Ensifera, Tettigoniidae): Taxonomic revision, phylogeny, stridulation, and development." volume="206" year="1998">Ingrisch 1998</bibRefCitation>
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) sclerotised, bulging, then curved and diminishing, largely surpassing tip of fused parts, terminating in an acute tip (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Liara (Acanthocoryphus) durata sp. nov. male holotype: A habitus lateral view; B face, C head dorsal view; D fastigium verticis lateral view; E stridulatory file on underside of left tegmen; F thoracic sternites, the arrow points at one of the prosternal spines; G-I titillators of male phallus in ventro-apical view (G), do. as seen from base (H), do. in oblique lateral view (I); J, K male cercus in external (J) and internal (K) views." httpUri="https://binary.pensoft.net/fig/487061" pageId="0" pageNumber="119">4G-I</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="coloration">
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<paragraph pageId="0" pageNumber="119">Coloration.</paragraph>
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<paragraph pageId="0" pageNumber="119">Uniformely yellowish brown. Frons reddish brown; mandibles black; tip of fastigium verticis especially on ventral side black. Pronotum with indication of brown lateral bands near hind margin only. Tegmen with dark spots. Meso- and metasternal lobes dark brown with tips of lobes yellowish brown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Measurements of male.</paragraph>
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<paragraph pageId="0" pageNumber="119">Body w/o wings: 19; pronotum: 7.6; tegmen: 8.2, hind femur: 14 mm.</paragraph>
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</subSubSection>
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</treatment>
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</document> |