332 lines
44 KiB
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332 lines
44 KiB
XML
<document id="16D415D5DF301D86EF50117585A7196A" ID-DOI="10.5281/zenodo.4650641" ID-ISSN="1638-9395" ID-Zenodo-Dep="4650641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617194300556" checkinUser="felipe" docAuthor="Badiola, Ainara, Astibia, Humberto, Suberbiola, Xabier Pereda & Murelaga, Xabier" docDate="2002" docId="C503CB7EFF8EFFD5FE87FA44FED8F2C0" docLanguage="en" docName="Geodiversitas.24.4.841-848.pdf" docOrigin="Geodiversitas 24 (4)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Leptolophus Remy 1965" docType="treatment" docVersion="3" lastPageNumber="846" masterDocId="393AB306FF8FFFD0FFD6FFEFFF9DF208" masterDocTitle="First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe" masterLastPageNumber="848" masterPageNumber="841" pageNumber="842" updateTime="1698897231207" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="6AC1852C6A2D6720E25B8CBCA3E66DD6">First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe</mods:title>
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<mods:namePart id="460AF16093DD6D1B9F8F27B808607D81">Badiola, Ainara</mods:namePart>
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<mods:affiliation id="4C803BBADE09FB0728C139240FCF0305">Universidad del País Vasco / Euskal Facultad de Ciencias, Departamento de Apartado 644,</mods:affiliation>
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<mods:roleTerm id="50A90890269F04E90E70D1D41C6E5DED">Author</mods:roleTerm>
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<mods:namePart id="934BD46BD2D18BBBEC1A17AC34FAA05D">Astibia, Humberto</mods:namePart>
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<mods:affiliation id="CAB10130D8C7F14C48C80CE8D56FE0C8">Herriko Unibertsitatea, Estratigrafía y Paleontología ,, E- 48080 Bilbao (Spain) gpbbakoa @ lg. ehu. es gppasayh @ lg. ehu. es</mods:affiliation>
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<mods:nameIdentifier id="AF7319F81BF7E90A4419D870A4BC7D3F" type="email">gpbbakoa@lg.ehu.es</mods:nameIdentifier>
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<mods:roleTerm id="C1C2ED645F62BDD269C35D449DC3A788">Author</mods:roleTerm>
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<mods:namePart id="115ED5E404CD92CBA73408E393711EFC">Suberbiola, Xabier Pereda</mods:namePart>
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<mods:affiliation id="05349B92AF1C006C5AC501BF5B167732">Universidad del País Vasco / Euskal Herriko Unibertsitatea, Facultad de Ciencias, Departamento de Estratigrafía y Paleontología, Apartado 644, E- 48080 Bilbao (Spain) Muséum national d’Histoire naturelle, Département Histoire de la Terre, 8 rue Buffon, F- 75231 Paris cedex 05 (France) gpppesuf @ lg. ehu. es</mods:affiliation>
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<mods:roleTerm id="105FAEEB30D639E0051F861EA9CB796B">Author</mods:roleTerm>
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<mods:namePart id="44E4AB565CD1C7C983FB8AD76C78845C">Murelaga, Xabier</mods:namePart>
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<mods:affiliation id="33135C6EDAEC2216600A005350B50636">Universidad del País Vasco / Euskal Herriko Unibertsitatea, Facultad de Ciencias, Departamento de Estratigrafía y Paleontología, Apartado 644, E- 48080 Bilbao (Spain) gpbmubej @ lg. ehu. es ..</mods:affiliation>
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<mods:typeOfResource id="B7EF3553F2539B4B3AC5BC00663D5007">text</mods:typeOfResource>
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<mods:title id="9C824BD313AF01F9EE245EDBD2346777">Geodiversitas</mods:title>
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<mods:part id="93A6C044AB8497C54CA581A9E4D0FDBC">
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<mods:date id="C55E68EA3A4D1053F924EE921D8C201F">2002</mods:date>
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<mods:number id="BD87528146F24002976FCA48ECB6857F">24</mods:number>
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<mods:start id="C63FF401BE3270B3597A2AD6E4F27A3F">841</mods:start>
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<mods:identifier id="CD1D6D1E705D4C6FD5A55898E7EAF757" type="DOI">10.5281/zenodo.4650641</mods:identifier>
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<mods:identifier id="8A9A23AE47A50AD7004BF7E416E54347" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="4F3418B8D77951FB118D975E35F92208" type="Zenodo-Dep">4650641</mods:identifier>
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<treatment id="C503CB7EFF8EFFD5FE87FA44FED8F2C0" LSID="urn:lsid:plazi:treatment:C503CB7EFF8EFFD5FE87FA44FED8F2C0" httpUri="http://treatment.plazi.org/id/C503CB7EFF8EFFD5FE87FA44FED8F2C0" lastPageId="5" lastPageNumber="846" pageId="1" pageNumber="842">
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<subSubSection id="05B029E3FF8EFFD1FE87FA44FE4BF7ED" pageId="1" pageNumber="842" type="nomenclature">
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<paragraph id="4D157A68FF8EFFD1FE87FA44FE4BF7ED" blockId="1.[337,497,1450,1509]" pageId="1" pageNumber="842">
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<heading id="165DCD04FF8EFFD1FE87FA44FE4BF7ED" centered="true" fontSize="11" level="2" pageId="1" pageNumber="842" reason="2">
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<taxonomicName id="8AAA01EBFF8EFFD1FE87FA44FE56F7CD" authorityName="Remy" authorityYear="1965" box="[337,459,1451,1477]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="842" phylum="Chordata" rank="genus">
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<emphasis id="7FDEA67AFF8EFFD1FE87FA44FE56F7CD" box="[337,459,1451,1477]" italics="true" pageId="1" pageNumber="842">Leptolophus</emphasis>
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</taxonomicName>
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sp. (
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<figureCitation id="D59166EDFF8EFFD1FEA2FA24FE51F7ED" box="[372,460,1483,1509]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="2.[151,162,1264,1281]" captionStartId-1="3.[151,162,883,900]" captionStartId-2="4.[151,162,1318,1335]" captionTargetBox-0="[221,1173,169,1228]" captionTargetBox-1="[275,1117,175,847]" captionTargetBox-2="[311,1090,172,1276]" captionTargetId-0="figure-182@2.[212,1185,156,1252]" captionTargetId-1="figure-361@3.[265,1132,173,835]" captionTargetId-2="figure-141@4.[295,1102,176,1291]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIG. 1. — Upper series M1 and M2 of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right M1 (MCNA 9947); A, labial view; B, occlusal view; C, lingual view; D-F, right M2 (MCNA 9948); D, labial view; E, occlusal view; F, lingual view. Scale bar: 5 cm." captionText-1="FIG. 2. — Upper teeth of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right P4? (MCNA 9963); A, labial view; B, occlusal view; C, lingual view; D, E, left fragment of upper tooth (MCNA 9964); D, labial view; E, occlusal view. Scale bar: 5 cm." captionText-2="FIG. 3. — Left mandible (MCNA 9002) showing the m1-2 series of Leptolophus sp. from Zambrana (Alava, Basque Country); A, occlusal view; B, labial view (modified from Astibia et al. 2000). Scale bar: 5 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4650643" figureDoi-1="http://doi.org/10.5281/zenodo.4650645" figureDoi-2="http://doi.org/10.5281/zenodo.4650647" httpUri-0="https://zenodo.org/record/4650643/files/figure.png" httpUri-1="https://zenodo.org/record/4650645/files/figure.png" httpUri-2="https://zenodo.org/record/4650647/files/figure.png" pageId="1" pageNumber="842">Figs 1-3</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="05B029E3FF8EFFD1FF41F9E9FB15F2D3" pageId="1" pageNumber="842" type="materials_examined">
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<paragraph id="4D157A68FF8EFFD1FF41F9E9FEAEF48C" blockId="1.[151,684,1542,1668]" pageId="1" pageNumber="842">
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MATERIAL EXAMINED. — Right M1 (
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<collectionCode id="2BBBE2ADFF8EFFD1FDC7F9E8FDFDF417" box="[529,608,1543,1567]" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="1" pageNumber="842">MCNA</collectionCode>
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9947); right M2 (
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<collectionCode id="2BBBE2ADFF8EFFD1FEDEF9CFFEC4F430" box="[264,345,1568,1592]" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="1" pageNumber="842">MCNA</collectionCode>
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9948); right upper (P4?) tooth (
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<collectionCode id="2BBBE2ADFF8EFFD1FF48F9D5FF72F45A" box="[158,239,1594,1618]" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="1" pageNumber="842">MCNA</collectionCode>
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9963); fragment of upper tooth (
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<collectionCode id="2BBBE2ADFF8EFFD1FD8DF9D5FD31F45A" box="[603,684,1594,1618]" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="1" pageNumber="842">MCNA</collectionCode>
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9964); left mandible with the m1-2 molar series (
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<collectionCode id="2BBBE2ADFF8EFFD1FF4BF983FF76F48C" box="[157,235,1644,1668]" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="1" pageNumber="842">MCNA</collectionCode>
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9002).
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</paragraph>
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<paragraph id="4D157A68FF8EFFD1FD1CFF46FB15F2D3" blockId="1.[714,1247,169,219]" pageId="1" pageNumber="842">
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LOCALITY AND AGE. — Zambrana, Alava; late Eocene (Priabonian,
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<collectionCode id="2BBBE2ADFF8EFFD1FC9CFF2CFCF2F2D2" box="[842,879,195,218]" country="0" name="Mohonk Preserve, Inc." pageId="1" pageNumber="842">MP</collectionCode>
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18) (see
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<bibRefCitation id="293B0799FF8EFFD1FC1EFF2CFBE0F2D3" author="ASTIBIA H. & ARANBURU A. & SUBERBIOLA X. & MURELAGA X. & SESE C. & CUESTA M. A. & SOLA S. & BACETA J. I. & BADIOLA A. & KOHLER M." box="[968,1149,195,219]" pageId="1" pageNumber="842" pagination="233 - 248" refId="ref2799" refString="ASTIBIA H., ARANBURU A., PEREDA- SUBERBIOLA X., MURELAGA X., SESE C., CUESTA M. A., MOYA- SOLA S., BACETA J. I., BADIOLA A. & KOHLER M. 2000. - Un nouveau gisement a vertebres continentaux de l'Eocene superieur de Zambrana (Bassin de Miranda-Trevino, Alava, Pays Basque). Geobios 32: 233 - 248." type="journal article" year="2000">
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Astibia
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<emphasis id="7FDEA67AFF8EFFD1FBC4FF2CFBA4F2D3" box="[1042,1081,195,219]" italics="true" pageId="1" pageNumber="842">et al</emphasis>
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. 2000
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</bibRefCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="05B029E3FF8EFFD1FD1CFEE5FC11F02D" pageId="1" pageNumber="842" type="discussion">
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<paragraph id="4D157A68FF8EFFD1FD1CFEE5FCB3F32A" blockId="1.[714,1247,266,549]" box="[714,814,266,292]" pageId="1" pageNumber="842">REMARKS</paragraph>
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<paragraph id="4D157A68FF8EFFD1FD1CFEC4FC11F02D" blockId="1.[714,1247,266,549]" pageId="1" pageNumber="842">
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The classical systematic nomenclature considers
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<taxonomicName id="8AAA01EBFF8EFFD1FD1CFEA4FC66F36D" authority="Pomel, 1847" authorityName="Pomel" authorityYear="1847" box="[714,1019,331,357]" class="Magnoliopsida" family="Asteraceae" genus="Plagiolophus" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="842" phylum="Tracheophyta" rank="genus">
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<emphasis id="7FDEA67AFF8EFFD1FD1CFEA4FCC4F36D" box="[714,857,331,357]" italics="true" pageId="1" pageNumber="842">Plagiolophus</emphasis>
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Pomel, 1847
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</taxonomicName>
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and
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<taxonomicName id="8AAA01EBFF8EFFD1FBECFEA4FCC3F38D" authority="Cuvier, 1804" authorityName="Cuvier" authorityYear="1804" class="Mammalia" family="Palaeotheriidae" genus="Palaeotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="842" phylum="Chordata" rank="genus">
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<emphasis id="7FDEA67AFF8EFFD1FBECFEA4FB42F36D" box="[1082,1247,331,357]" italics="true" pageId="1" pageNumber="842">Palaeotherium</emphasis>
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Cuvier, 1804
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</taxonomicName>
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as members of either the
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<taxonomicName id="8AAA01EBFF8EFFD1FB53FE84FB42F38D" authorityName="Gray" authorityYear="1821" box="[1157,1247,363,389]" class="Mammalia" family="Equidae" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="842" phylum="Chordata" rank="family">Equidae</taxonomicName>
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or the Palaeotheridae (
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<bibRefCitation id="293B0799FF8EFFD1FC6CFE64FBE3F3AD" author="SAVAGE D. E. & RUSSELL D. E. & LOUIS P." box="[954,1150,394,421]" pageId="1" pageNumber="842" pagination="1 - 94" refId="ref3401" refString="SAVAGE D. E., RUSSELL D. E. & LOUIS P. 1965. - European Eocene Equidae (Perissodactyla). University of California Publications, Geological Sciences 56: 1 - 94." type="journal article" year="1965">
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Savage
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<emphasis id="7FDEA67AFF8EFFD1FBDCFE65FBA8F3AD" box="[1034,1077,394,421]" italics="true" pageId="1" pageNumber="842">et al</emphasis>
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. 1965
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</bibRefCitation>
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;
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<bibRefCitation id="293B0799FF8EFFD1FB5EFE65FC99F3CD" author="FRANZEN J. L." pageId="1" pageNumber="842" refId="ref3249" refString="FRANZEN J. L. 1968. - Revision der Gattung Palaeotherium Cuvier, 1804 (Palaeotheriidae, Perissodactyla, Mammalia). Dissertation Albert- Ludwigs-Universitat zu Freiburg, Freiburg, Germany, 2 vols, 181 p." type="book" year="1968">Franzen 1968</bibRefCitation>
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;
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<bibRefCitation id="293B0799FF8EFFD1FCD8FE45FC12F3CD" author="REMY J. A." box="[782,911,426,453]" pageId="1" pageNumber="842" refId="ref3327" refString="REMY J. A. 1976. - Etude comparative des structures dentaires chez les Paleotheriidae et divers autres perissodactyles fossiles. These 3 e cycle, Universite Louis Pasteur, Strasbourg I, France, 207 p." type="book" year="1976">Remy 1976</bibRefCitation>
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). Here we follow the systematic proposition of
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<bibRefCitation id="293B0799FF8EFFD1FCA4FE24FB91F3ED" author="CUESTA M. A." box="[882,1036,459,485]" pageId="1" pageNumber="842" pagination="87 - 109" refId="ref3166" refString="CUESTA M. A. 1993. - Los Palaeotheriidae (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, Espana). Estudios Geologicos 49: 87 - 109." type="journal article" year="1993">Cuesta (1993</bibRefCitation>
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,
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<bibRefCitation id="293B0799FF8EFFD1FBCCFE24FBC9F3ED" author="CUESTA M. A." box="[1050,1108,459,485]" pageId="1" pageNumber="842" pagination="253 - 279" refId="ref3204" refString="CUESTA M. A. 1994. - Los Plagiolophinae (Remy, 1976) nuevo rango (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, Espana). Estudios Geologicos 50: 253 - 279." type="journal article" year="1994">1994</bibRefCitation>
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||
) who classified
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<taxonomicName id="8AAA01EBFF8EFFD1FCDDFE04FC39F00D" authorityName="Pomel" authorityYear="1847" box="[779,932,491,517]" class="Magnoliopsida" family="Asteraceae" genus="Plagiolophus" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="842" phylum="Tracheophyta" rank="genus">
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<emphasis id="7FDEA67AFF8EFFD1FCDDFE04FC39F00D" box="[779,932,491,517]" italics="true" pageId="1" pageNumber="842">Plagiolophus</emphasis>
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</taxonomicName>
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and closer relatives as
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<taxonomicName id="8AAA01EBFF8EFFD1FD1CFDE4FC15F02D" authorityName="Pavlow" authorityYear="1888" box="[714,904,523,549]" class="Mammalia" family="Pachynolophidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="842" phylum="Chordata" rank="family">Pachynolophidae</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="05B029E3FF8EFFD5FD1CFDA5FED8F2C0" lastPageId="5" lastPageNumber="846" pageId="1" pageNumber="842" type="description">
|
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<paragraph id="4D157A68FF8EFFD1FD1CFDA5FCC6F06A" blockId="1.[714,1248,586,1669]" box="[714,859,586,612]" pageId="1" pageNumber="842">DESCRIPTION</paragraph>
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<paragraph id="4D157A68FF8EFFD1FD1CFD84FB28F0ED" blockId="1.[714,1248,586,1669]" pageId="1" pageNumber="842">The upper tooth remains which we described were found at the same level and in close proximity to each other. They probably belong to the same species and even to the same individual.</paragraph>
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<paragraph id="4D157A68FF8EFFD1FD1CFD04FD64F6ED" blockId="1.[714,1248,586,1669]" pageId="1" pageNumber="842">
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The molar series exhibits a strong height gradient (
|
||
<figureCitation id="D59166EDFF8EFFD1FD04FCE4FC8CF12D" box="[722,785,779,805]" captionStart="FIG" captionStartId="2.[151,162,1264,1281]" captionTargetBox="[221,1173,169,1228]" captionTargetId="figure-182@2.[212,1185,156,1252]" captionTargetPageId="2" captionText="FIG. 1. — Upper series M1 and M2 of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right M1 (MCNA 9947); A, labial view; B, occlusal view; C, lingual view; D-F, right M2 (MCNA 9948); D, labial view; E, occlusal view; F, lingual view. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650643" httpUri="https://zenodo.org/record/4650643/files/figure.png" pageId="1" pageNumber="842">Fig. 1</figureCitation>
|
||
). The size of the M1 is much smaller than that of the M2. The crown of M1 is very worn and less high (
|
||
<quantity id="8A52D78DFF8EFFD1FCA2FCA4FC7FF16C" box="[884,994,842,869]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="842" unit="mm" value="17.5">17.5 mm</quantity>
|
||
) than that of the M2 (
|
||
<quantity id="8A52D78DFF8EFFD1FD04FC84FCBBF18C" box="[722,806,874,901]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="842" unit="mm" value="31.0">31 mm</quantity>
|
||
). The M1 is very molariform and it has a trapezoidal outline, with the lingual wall being shorter than the labial one (
|
||
<figureCitation id="D59166EDFF8EFFD1FB91FC44FB0FF1CD" box="[1095,1170,939,965]" captionStart="FIG" captionStartId="2.[151,162,1264,1281]" captionTargetBox="[221,1173,169,1228]" captionTargetId="figure-182@2.[212,1185,156,1252]" captionTargetPageId="2" captionText="FIG. 1. — Upper series M1 and M2 of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right M1 (MCNA 9947); A, labial view; B, occlusal view; C, lingual view; D-F, right M2 (MCNA 9948); D, labial view; E, occlusal view; F, lingual view. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650643" httpUri="https://zenodo.org/record/4650643/files/figure.png" pageId="1" pageNumber="842">Fig. 1</figureCitation>
|
||
A-C). Morphologically, the M1 is similar to the M2, but its cingula are more developed. The labial cingulum is W-shaped, quite thick and prominent. The anterolingual cingulum is also well developed and it ascends up to the occlusal surface. Moreover, the development of the parastyle and metastyle is decreasing in the molar series. These styles are less developed in M1 than in the M2.
|
||
</paragraph>
|
||
<paragraph id="4D157A68FF8EFFD2FD1CFB04FB1AF7CC" blockId="1.[714,1248,586,1669]" lastBlockId="2.[714,1247,1386,1669]" lastPageId="2" lastPageNumber="843" pageId="1" pageNumber="842">
|
||
The M2 (
|
||
<figureCitation id="D59166EDFF8EFFD1FCEDFB04FCE1F70D" box="[827,892,1259,1285]" captionStart="FIG" captionStartId="2.[151,162,1264,1281]" captionTargetBox="[221,1173,169,1228]" captionTargetId="figure-182@2.[212,1185,156,1252]" captionTargetPageId="2" captionText="FIG. 1. — Upper series M1 and M2 of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right M1 (MCNA 9947); A, labial view; B, occlusal view; C, lingual view; D-F, right M2 (MCNA 9948); D, labial view; E, occlusal view; F, lingual view. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650643" httpUri="https://zenodo.org/record/4650643/files/figure.png" pageId="1" pageNumber="842">Fig. 1</figureCitation>
|
||
D-F) is moderately worn. It is very hypsodont, with its internal wall being as high as the external one. The crown is trapezoidal, with the basal part being narrower than the occlusal surface. The ectoloph is anteroposteriorly flat, showing an external wall that is lingually inclined and slightly concave on both sides of the mesostyle. The latter is very sharp and prominent. The ribs of the paracone and metacone are absent. In contrast, the parastyle and metastyle are well developed. The parastyle protrudes notably on the labial side, while the metastyle extends more posteriorly. Protoloph and metaloph are quite slender and oblique relative to the ectoloph. The latter is incomplete but it seems that the metaloph is connected to it, so that the two transversal valleys are separate. Owing to the oblique metaloph, a short and slend- er crista (hypoloph?) is present between the ectoloph and metaloph; it is sligtly convex and closes the posterior fossa. The labial cingulum is quite incomplete, but it continues and ascends up to the occlusal surface. The basal part of the internal surface is also incomplete, but it seems that the internal cingulum is not present.
|
||
</paragraph>
|
||
<caption id="19D52AE0FF8DFFD2FF41FB1FFF5AF724" ID-DOI="http://doi.org/10.5281/zenodo.4650643" ID-Zenodo-Dep="4650643" httpUri="https://zenodo.org/record/4650643/files/figure.png" pageId="2" pageNumber="843" startId="2.[151,162,1264,1281]" targetBox="[221,1173,169,1228]" targetPageId="2">
|
||
<paragraph id="4D157A68FF8DFFD2FF41FB1FFF5AF724" blockId="2.[151,1249,1264,1324]" pageId="2" pageNumber="843">
|
||
FIG. 1. — Upper series M1 and M2 of
|
||
<taxonomicName id="8AAA01EBFF8DFFD2FE18FB1FFDA8F709" authorityName="Remy" authorityYear="1965" box="[462,565,1264,1281]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="2" pageNumber="843" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8DFFD2FE18FB1FFDA8F709" box="[462,565,1264,1281]" italics="true" pageId="2" pageNumber="843">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
sp. from Zambrana (Alava, Basque Country);
|
||
<emphasis id="7FDEA67AFF8DFFD2FC78FB1FFC4DF709" bold="true" box="[942,976,1264,1281]" pageId="2" pageNumber="843">A -C</emphasis>
|
||
, right M1 (MCNA 9947);
|
||
<emphasis id="7FDEA67AFF8DFFD2FB4BFB1FFB37F709" bold="true" box="[1181,1194,1264,1281]" pageId="2" pageNumber="843">A</emphasis>
|
||
, labial view;
|
||
<emphasis id="7FDEA67AFF8DFFD2FF1FFAE9FF4BF71F" bold="true" box="[201,214,1286,1303]" pageId="2" pageNumber="843">B</emphasis>
|
||
, occlusal view;
|
||
<emphasis id="7FDEA67AFF8DFFD2FEB4FAE9FEEDF71F" bold="true" box="[354,368,1286,1303]" pageId="2" pageNumber="843">C</emphasis>
|
||
, lingual view;
|
||
<emphasis id="7FDEA67AFF8DFFD2FE3BFAE9FD93F71F" bold="true" box="[493,526,1286,1303]" pageId="2" pageNumber="843">D -F</emphasis>
|
||
, right M2 (MCNA 9948);
|
||
<emphasis id="7FDEA67AFF8DFFD2FD3EFAE9FD6BF71F" bold="true" box="[744,758,1286,1303]" pageId="2" pageNumber="843">D</emphasis>
|
||
, labial view;
|
||
<emphasis id="7FDEA67AFF8DFFD2FCBEFAE9FCE9F71F" bold="true" box="[872,884,1286,1303]" pageId="2" pageNumber="843">E</emphasis>
|
||
, occlusal view;
|
||
<emphasis id="7FDEA67AFF8DFFD2FBD6FAE9FB96F71F" bold="true" box="[1024,1035,1286,1303]" pageId="2" pageNumber="843">F</emphasis>
|
||
, lingual view. Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D157A68FF8DFFD3FD1CFA24FD17F66D" blockId="2.[714,1247,1386,1669]" lastBlockId="3.[151,684,970,1509]" lastPageId="3" lastPageNumber="844" pageId="2" pageNumber="843">
|
||
MCNA 9963 and 9964 look like the M1 and M2 described above, but they are much smaller and less hypsodont than these teeth (
|
||
<figureCitation id="D59166EDFF8DFFD2FBFCF9E4FBF0F42D" box="[1066,1133,1547,1573]" captionStart="FIG" captionStartId="3.[151,162,883,900]" captionTargetBox="[275,1117,175,847]" captionTargetId="figure-361@3.[265,1132,173,835]" captionTargetPageId="3" captionText="FIG. 2. — Upper teeth of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right P4? (MCNA 9963); A, labial view; B, occlusal view; C, lingual view; D, E, left fragment of upper tooth (MCNA 9964); D, labial view; E, occlusal view. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650645" httpUri="https://zenodo.org/record/4650645/files/figure.png" pageId="2" pageNumber="843">Fig. 2</figureCitation>
|
||
). The best preserved tooth (MCNA 9963) is quite molariform (
|
||
<figureCitation id="D59166EDFF8DFFD2FCDAF9A4FCD7F46D" box="[780,842,1611,1637]" captionStart="FIG" captionStartId="3.[151,162,883,900]" captionTargetBox="[275,1117,175,847]" captionTargetId="figure-361@3.[265,1132,173,835]" captionTargetPageId="3" captionText="FIG. 2. — Upper teeth of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right P4? (MCNA 9963); A, labial view; B, occlusal view; C, lingual view; D, E, left fragment of upper tooth (MCNA 9964); D, labial view; E, occlusal view. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650645" httpUri="https://zenodo.org/record/4650645/files/figure.png" pageId="2" pageNumber="843">Fig. 2</figureCitation>
|
||
A-C). The protoloph is slender, arches anteriorly and posteriorly; it is connected to the ectoloph and exhibits a U shape. The development of the styles is very similar to that of the M1. The ribs of the paracone and metacone are absent. These teeth are reminiscent of the premolars (probably the P4) of
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FEA2FBA4FD18F66D" authority="(Remy 1998)" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[372,645,1099,1125]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="844" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FEA2FBA4FE73F66D" box="[372,494,1099,1125]" italics="true" pageId="3" pageNumber="844">Leptolophus</emphasis>
|
||
(
|
||
<bibRefCitation id="293B0799FF8CFFD3FE2DFBA4FDE3F66D" author="REMY J. A." box="[507,638,1099,1125]" pageId="3" pageNumber="844" pagination="45 - 108" refId="ref3366" refString="REMY J. A. 1998. - Le genre Leptolophus (Perissodactyla, Mammalia): Morphologie et histologie dentaires, anatomie cranienne, implications fonctionnelles. Palaeovertebrata 27: 45 - 108." type="journal article" year="1998">Remy 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="19D52AE0FF8CFFD3FF41FC9CFB4FF192" ID-DOI="http://doi.org/10.5281/zenodo.4650645" ID-Zenodo-Dep="4650645" httpUri="https://zenodo.org/record/4650645/files/figure.png" pageId="3" pageNumber="844" startId="3.[151,162,883,900]" targetBox="[275,1117,175,847]" targetPageId="3">
|
||
<paragraph id="4D157A68FF8CFFD3FF41FC9CFB4FF192" blockId="3.[151,1248,883,922]" pageId="3" pageNumber="844">
|
||
FIG. 2. — Upper teeth of
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FEA6FC9CFE44F18C" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[368,473,883,900]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="844" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FEA6FC9CFE44F18C" box="[368,473,883,900]" italics="true" pageId="3" pageNumber="844">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
sp. from Zambrana (Alava, Basque Country);
|
||
<emphasis id="7FDEA67AFF8CFFD3FCB0FC9CFC15F18C" bold="true" box="[870,904,883,900]" pageId="3" pageNumber="844">A -C</emphasis>
|
||
, right P4? (MCNA 9963);
|
||
<emphasis id="7FDEA67AFF8CFFD3FBB1FC9CFBE9F18C" bold="true" box="[1127,1140,883,900]" pageId="3" pageNumber="844">A</emphasis>
|
||
, labial view;
|
||
<emphasis id="7FDEA67AFF8CFFD3FF41FC66FF39F192" bold="true" box="[151,164,905,922]" pageId="3" pageNumber="844">B</emphasis>
|
||
, occlusal view;
|
||
<emphasis id="7FDEA67AFF8CFFD3FEFEFC66FEABF192" bold="true" box="[296,310,905,922]" pageId="3" pageNumber="844">C</emphasis>
|
||
, lingual view;
|
||
<emphasis id="7FDEA67AFF8CFFD3FE7AFC66FE27F192" bold="true" box="[428,442,905,922]" pageId="3" pageNumber="844">D</emphasis>
|
||
,
|
||
<emphasis id="7FDEA67AFF8CFFD3FE12FC66FE4DF192" bold="true" box="[452,464,905,922]" pageId="3" pageNumber="844">E</emphasis>
|
||
, left fragment of upper tooth (MCNA 9964);
|
||
<emphasis id="7FDEA67AFF8CFFD3FCE9FC66FCD0F192" bold="true" box="[831,845,905,922]" pageId="3" pageNumber="844">D</emphasis>
|
||
, labial view;
|
||
<emphasis id="7FDEA67AFF8CFFD3FC6EFC66FC59F192" bold="true" box="[952,964,905,922]" pageId="3" pageNumber="844">E</emphasis>
|
||
, occlusal view. Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D157A68FF8CFFD3FF41FB84FD90F7ED" blockId="3.[151,684,970,1509]" pageId="3" pageNumber="844">
|
||
The mandible MCNA 9002 preserves only the m1 and m2 (
|
||
<figureCitation id="D59166EDFF8CFFD3FEFFFB64FEF6F6AD" box="[297,363,1163,1189]" captionStart="FIG" captionStartId="4.[151,162,1318,1335]" captionTargetBox="[311,1090,172,1276]" captionTargetId="figure-141@4.[295,1102,176,1291]" captionTargetPageId="4" captionText="FIG. 3. — Left mandible (MCNA 9002) showing the m1-2 series of Leptolophus sp. from Zambrana (Alava, Basque Country); A, occlusal view; B, labial view (modified from Astibia et al. 2000). Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650647" httpUri="https://zenodo.org/record/4650647/files/figure.png" pageId="3" pageNumber="844">Fig. 3</figureCitation>
|
||
). These teeth are hypsodont, narrow and exhibit a strong height gradient: the size of the m2 is much larger and longer than the m1. The crowns are trapezoidal and the internal wall is as high as the external one. In lateral view, the lophids are rounded and have a U shape, with a concave aspect in the internal wall. The paralophid is well developed and it protrudes notably on the lingual side. The m2 has a small hypoconulid, but does not ascend up to the occlusal surface; in m1 it is absent.
|
||
</paragraph>
|
||
<paragraph id="4D157A68FF8CFFD3FF41F9E4FEACF42A" blockId="3.[151,684,1547,1669]" box="[151,305,1547,1573]" pageId="3" pageNumber="844">COMPARISONS</paragraph>
|
||
<paragraph id="4D157A68FF8CFFD3FF41F9C4FD31F48D" blockId="3.[151,684,1547,1669]" pageId="3" pageNumber="844">
|
||
The M1 and M2 of Zambrana are typical of the genus
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FF31F9A4FEE9F46D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[231,372,1611,1637]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="844" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FF31F9A4FEE9F46D" box="[231,372,1611,1637]" italics="true" pageId="3" pageNumber="844">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
and differ from those of
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FF41F984FEBBF48D" authorityName="Pomel" authorityYear="1847" box="[151,294,1643,1669]" class="Magnoliopsida" family="Asteraceae" genus="Plagiolophus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="844" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FF41F984FEBBF48D" box="[151,294,1643,1669]" italics="true" pageId="3" pageNumber="844">Plagiolophus</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics:
|
||
</paragraph>
|
||
<paragraph id="4D157A68FF8CFFD3FD1CFC24FCA4F70C" blockId="3.[714,1247,970,1669]" pageId="3" pageNumber="844">1) trapezoidal outline, with the basal part being more narrower than the occlusal surface; 2) proto- and metaloph much longer and more slender, very lophodont and oriented more obliquely to the ectoloph; 3) less developed intermediate conules; 4) much higher internal cusp, with the hypsodonty of the labial and lingual surfaces being almost equal; 5) ectoloph anteroposteriorly flat; and 6) absence of ribs of the paracone and metacone.</paragraph>
|
||
<paragraph id="4D157A68FF8CFFD3FD1CFAE4FB57F42D" blockId="3.[714,1247,970,1669]" pageId="3" pageNumber="844">
|
||
The mandible MCNA 9002, previously referred to as
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FD32FAC4FB22F74D" authority="Cuesta, 1994" authorityName="Cuesta" authorityYear="1994" box="[740,1215,1322,1349]" class="Magnoliopsida" family="Asteraceae" genus="Plagiolophus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="844" phylum="Tracheophyta" rank="species" species="mazateronensis">
|
||
<emphasis id="7FDEA67AFF8CFFD3FD32FAC4FCF9F74D" box="[740,868,1323,1349]" italics="true" pageId="3" pageNumber="844">Plagiolophus</emphasis>
|
||
aff.
|
||
<emphasis id="7FDEA67AFF8CFFD3FC44FAC5FBB1F74B" box="[914,1068,1322,1347]" italics="true" pageId="3" pageNumber="844">mazateronensis</emphasis>
|
||
<bibRefCitation id="293B0799FF8CFFD3FBE7FAC4FB22F74D" author="CUESTA M. A." box="[1073,1215,1323,1349]" pageId="3" pageNumber="844" pagination="253 - 279" refId="ref3204" refString="CUESTA M. A. 1994. - Los Plagiolophinae (Remy, 1976) nuevo rango (Perissodactyla, Mammalia) del Eoceno de la Cuenca del Duero (Castilla y Leon, Espana). Estudios Geologicos 50: 253 - 279." type="journal article" year="1994">Cuesta, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="293B0799FF8CFFD3FD1CFAA4FC4FF76D" author="ASTIBIA H. & ARANBURU A. & SUBERBIOLA X. & MURELAGA X. & SESE C. & CUESTA M. A. & SOLA S. & BACETA J. I. & BADIOLA A. & KOHLER M." box="[714,978,1354,1381]" pageId="3" pageNumber="844" pagination="233 - 248" refId="ref2799" refString="ASTIBIA H., ARANBURU A., PEREDA- SUBERBIOLA X., MURELAGA X., SESE C., CUESTA M. A., MOYA- SOLA S., BACETA J. I., BADIOLA A. & KOHLER M. 2000. - Un nouveau gisement a vertebres continentaux de l'Eocene superieur de Zambrana (Bassin de Miranda-Trevino, Alava, Pays Basque). Geobios 32: 233 - 248." type="journal article" year="2000">
|
||
Astibia
|
||
<emphasis id="7FDEA67AFF8CFFD3FCF8FAA5FCF9F76D" box="[814,868,1354,1381]" italics="true" pageId="3" pageNumber="844">et al</emphasis>
|
||
. (2000)
|
||
</bibRefCitation>
|
||
, differs from that of
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FD1CFA84FCC8F78D" authorityName="Pomel" authorityYear="1847" box="[714,853,1387,1413]" class="Magnoliopsida" family="Asteraceae" genus="Plagiolophus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="844" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FD1CFA84FCC8F78D" box="[714,853,1387,1413]" italics="true" pageId="3" pageNumber="844">Plagiolophus</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: 1) the height gradient of the m1-2 series is well developed; 2) the teeth have a trapezoidal outline; 3) the internal wall is as high as the external one; and 4) m1 without hypoconulid, m2 with a small one which does not ascend up to the occlusal surface.
|
||
</paragraph>
|
||
<paragraph id="4D157A68FF8CFFD4FD1CF9C4FC7CF48C" blockId="3.[714,1247,970,1669]" lastBlockId="4.[714,1247,1418,1669]" lastPageId="4" lastPageNumber="845" pageId="3" pageNumber="844">
|
||
The Zambrana upper teeth are larger and more hypsodont than those of the
|
||
<taxonomicName id="8AAA01EBFF8CFFD3FBC6F9A4FB11F46D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[1040,1164,1611,1637]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="844" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8CFFD3FBC6F9A4FB11F46D" box="[1040,1164,1611,1637]" italics="true" pageId="3" pageNumber="844">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
species from the late middle Eocene of southern
|
||
<collectingCountry id="35BD3AF8FF8CFFD3FB40F985FB42F48C" box="[1174,1247,1642,1668]" name="France" pageId="3" pageNumber="844">France</collectingCountry>
|
||
and
|
||
<collectingCountry id="35BD3AF8FF8BFFD4FF18FA64FEFDF7AD" box="[206,352,1419,1445]" name="Switzerland" pageId="4" pageNumber="845">Switzerland</collectingCountry>
|
||
(
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FEA0FA65FE6EF7AD" authorityName="Remy" authorityYear="1965" box="[374,499,1418,1445]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="species" species="stehlini">
|
||
<emphasis id="7FDEA67AFF8BFFD4FEA0FA65FE6EF7AD" box="[374,499,1418,1445]" italics="true" pageId="4" pageNumber="845">L. stehlini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FDE3FA65FEABF7CD" authority="Stehlin, 1904" authorityName="Stehlin" authorityYear="1904" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="species" species="nouleti">
|
||
<emphasis id="7FDEA67AFF8BFFD4FDE3FA65FD31F7AD" box="[565,684,1418,1445]" italics="true" pageId="4" pageNumber="845">L. nouleti</emphasis>
|
||
Stehlin, 1904
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="293B0799FF8BFFD4FEB9FA45FE68F7CD" author="REMY J. A." box="[367,501,1450,1477]" pageId="4" pageNumber="845" pagination="45 - 108" refId="ref3366" refString="REMY J. A. 1998. - Le genre Leptolophus (Perissodactyla, Mammalia): Morphologie et histologie dentaires, anatomie cranienne, implications fonctionnelles. Palaeovertebrata 27: 45 - 108." type="journal article" year="1998">Remy 1998</bibRefCitation>
|
||
) (
|
||
<figureCitation id="D59166EDFF8BFFD4FDC6FA44FDC5F7CD" box="[528,600,1451,1477]" captionStart="FIG" captionStartId="5.[151,162,1148,1165]" captionTargetBox="[305,1091,229,1113]" captionTargetId="graphics-148@5.[364,816,247,1075]" captionTargetPageId="5" captionText="FIG. 4. — A, diagram d/D (in mm) of the M1 of several Pachynolophidae from the Eocene of Europe, D, largest diagonal (from the parastyle to the hypocone) of the upper teeth, d, diagonal perpendicular to D; B, diagram l/L (in mm) of the M2 of several Pachynolophidae from the Eocene of Europe, l, maximum width, L, average length of the external wall of the ectoloph. Measurements from Casanovas 1975 and Casanovas et al. 1998 for Plagiolophus annectens (Owen, 1848); Cuesta 1994 for Plagiolophus mazateronensis Cuesta, 1994 and Leptolophus sp.; Remy 1998 for Leptolophus stehlini Remy, 1965 and Leptolophus nouleti Stehlin, 1904." figureDoi="http://doi.org/10.5281/zenodo.4650649" httpUri="https://zenodo.org/record/4650649/files/figure.png" pageId="4" pageNumber="845">Fig. 4</figureCitation>
|
||
). Morphologically, the teeth are comparable to those of
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FF41FA04FE8BF40D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[151,278,1515,1541]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8BFFD4FF41FA04FE8BF40D" box="[151,278,1515,1541]" italics="true" pageId="4" pageNumber="845">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
sp. from Mazaterón, but the Zambrana teeth are clearly larger and more hypsodont. The large size and the more accentuated hypsodonty are considered as derived characters compared to
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FEFBF984FE37F48D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[301,426,1643,1669]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8BFFD4FEFBF984FE37F48D" box="[301,426,1643,1669]" italics="true" pageId="4" pageNumber="845">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
from Mazaterón. This interpretation is coherent with the younger age of the Zambrana locality (MP 18) relatively to Mazaterón (MP 15-16) (see
|
||
<bibRefCitation id="293B0799FF8BFFD4FC2BFA24FB4EF7ED" author="ANTUNES M. & CASANOVAS M. L. & CUESTA M. A. & CHECA L. & SANTAFE J. V. & AGUSTI J." box="[1021,1235,1482,1509]" pageId="4" pageNumber="845" pagination="97" refId="ref2721" refString="ANTUNES M., CASANOVAS M. L., CUESTA M. A., CHECA L., SANTAFE J. V. & AGUSTI J. 1997. - Eocene mammals from Iberian Peninsula, in AGUILAR J. P., LEGENDRE S. & MICHAUX J. (eds), Actes du Congres BiochroM' 97. Memoires et" type="book chapter" year="1997">
|
||
Antunes
|
||
<emphasis id="7FDEA67AFF8BFFD4FB88FA25FB0CF7ED" box="[1118,1169,1482,1509]" italics="true" pageId="4" pageNumber="845">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
). On the basis of these differences, the Zambrana specimens may belong to a new species of
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FD1CF9C4FCCCF44D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[714,849,1579,1605]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8BFFD4FD1CF9C4FCCCF44D" box="[714,849,1579,1605]" italics="true" pageId="4" pageNumber="845">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
. By pending a revision of all the material of
|
||
<emphasis id="7FDEA67AFF8BFFD4FC9AF9A4FC52F46D" box="[844,975,1611,1637]" italics="true" pageId="4" pageNumber="845">
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FC9AF9A4FC51F46D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[844,972,1611,1637]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">Leptolophus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the Zambrana teeth are referred to
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FC94F984FC21F48D" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[834,956,1643,1669]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8BFFD4FC94F984FC21F48D" box="[834,956,1643,1669]" italics="true" pageId="4" pageNumber="845">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
<caption id="19D52AE0FF8BFFD4FF41FAC9FCDBF745" ID-DOI="http://doi.org/10.5281/zenodo.4650647" ID-Zenodo-Dep="4650647" httpUri="https://zenodo.org/record/4650647/files/figure.png" pageId="4" pageNumber="845" startId="4.[151,162,1318,1335]" targetBox="[311,1090,172,1276]" targetPageId="4">
|
||
<paragraph id="4D157A68FF8BFFD4FF41FAC9FCDBF745" blockId="4.[151,1249,1318,1357]" pageId="4" pageNumber="845">
|
||
FIG. 3. — Left mandible (MCNA 9002) showing the m1-2 series of
|
||
<taxonomicName id="8AAA01EBFF8BFFD4FD34FAC9FCD3F73F" baseAuthorityName="Remy" baseAuthorityYear="1998" box="[738,846,1318,1335]" class="Mammalia" family="Pachynolophidae" genus="Leptolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="845" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FDEA67AFF8BFFD4FD34FAC9FCD3F73F" box="[738,846,1318,1335]" italics="true" pageId="4" pageNumber="845">Leptolophus</emphasis>
|
||
</taxonomicName>
|
||
sp. from Zambrana (Alava, Basque Country);
|
||
<emphasis id="7FDEA67AFF8BFFD4FF41FAD3FF39F745" bold="true" box="[151,164,1340,1357]" pageId="4" pageNumber="845">A</emphasis>
|
||
, occlusal view;
|
||
<emphasis id="7FDEA67AFF8BFFD4FEFEFAD3FEA8F745" bold="true" box="[296,309,1340,1357]" pageId="4" pageNumber="845">B</emphasis>
|
||
, labial view (modified from
|
||
<bibRefCitation id="293B0799FF8BFFD4FDCFFAD3FD2CF745" author="ASTIBIA H. & ARANBURU A. & SUBERBIOLA X. & MURELAGA X. & SESE C. & CUESTA M. A. & SOLA S. & BACETA J. I. & BADIOLA A. & KOHLER M." box="[537,689,1340,1357]" pageId="4" pageNumber="845" pagination="233 - 248" refId="ref2799" refString="ASTIBIA H., ARANBURU A., PEREDA- SUBERBIOLA X., MURELAGA X., SESE C., CUESTA M. A., MOYA- SOLA S., BACETA J. I., BADIOLA A. & KOHLER M. 2000. - Un nouveau gisement a vertebres continentaux de l'Eocene superieur de Zambrana (Bassin de Miranda-Trevino, Alava, Pays Basque). Geobios 32: 233 - 248." type="journal article" year="2000">
|
||
Astibia
|
||
<emphasis id="7FDEA67AFF8BFFD4FD81FAD3FDE7F745" box="[599,634,1340,1357]" italics="true" pageId="4" pageNumber="845">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
). Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D157A68FF8AFFD5FEE5FF40FED8F2C0" blockId="5.[307,325,175,200]" box="[307,325,175,200]" pageId="5" pageNumber="846">A</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |