320 lines
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320 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.72474" ID-GBIF-Dataset="11e327fb-5ef8-4d85-bd77-e92a1dab8c3f" ID-GBIF-Taxon="190303381" ID-Pensoft-Pub="2535-0730-2-275" ID-Pensoft-UUID="671FAEEE86B25BE9A720A18193BC42B1" ID-ZooBank="9AC22770F3004265A21F841EA364FFD5" ModsDocID="2535-0730-5-2-275" checkinTime="1635891665621" checkinUser="pensoft" docAuthor="Framenau, Volker W., Baptista, Renner L. C., Oliveira, Francisca Samia M. & Castanheira, Pedro de S." docDate="2021" docId="B5D181AD1658587EB9157EC26439EF97" docLanguage="en" docName="EvolutSyst 5(2): 275-334" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-11-02" docSource="http://dx.doi.org/10.3897/evolsyst.5.72474" docTitle="Hortophora urbana Framenau & Baptista & Oliveira & Castanheira 2021, comb. nov." docType="treatment" docVersion="3" id="671FAEEE86B25BE9A720A18193BC42B1" lastPageNumber="275" masterDocId="671FAEEE86B25BE9A720A18193BC42B1" masterDocTitle="Taxonomic revision of the new spider genus Hortophora, the Australasian Garden Orb-weavers (Araneae, Araneidae)" masterLastPageNumber="334" masterPageNumber="275" pageId="0" pageNumber="275" updateTime="1637334715605" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the new spider genus Hortophora, the Australasian Garden Orb-weavers (Araneae, Araneidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Framenau, Volker W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
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<mods:affiliation>Harry Butler Institute, Murdoch University, Murdoch, Western Australia 6150, Australia & Department of Terrestrial Zoology, Western Australian Museum, Welshpool, Western Australia 6103, Australia & Zoological Museum Hamburg, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Centre for Taxonomy & Morphology, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">volker.framenau@murdoch.edu.au</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baptista, Renner L. C.</mods:namePart>
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<mods:affiliation>Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oliveira, Francisca Samia M.</mods:namePart>
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<mods:affiliation>Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Castanheira, Pedro de S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0623-1622</mods:nameIdentifier>
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<mods:affiliation>Harry Butler Institute, Murdoch University, Murdoch, Western Australia 6150, Australia & Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>275</mods:start>
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<mods:end>334</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.5.72474</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.5.72474</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-275</mods:identifier>
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<mods:identifier type="ZooBank">9AC22770F3004265A21F841EA364FFD5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">671FAEEE86B25BE9A720A18193BC42B1</mods:identifier>
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<treatment ID-GBIF-Taxon="190303381" LSID="urn:lsid:plazi:treatment:B5D181AD1658587EB9157EC26439EF97" httpUri="http://treatment.plazi.org/id/B5D181AD1658587EB9157EC26439EF97" lastPageNumber="275" pageId="0" pageNumber="275">
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<subSubSection pageId="0" pageNumber="275" type="nomenclature">
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName LSID="B5D181AD-1658-587E-B915-7EC26439EF97" authority="(Keyserling, 1887)" authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Keyserling" baseAuthorityYear="1887" family="Araneidae" genus="Hortophora" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hortophora urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana" status="comb. nov.">Hortophora urbana (Keyserling, 1887)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="275">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="description">
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<paragraph pageId="0" pageNumber="275">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hortophora gen. nov. species, second legs of male, prolateral view. A. H. biapicata comb. nov., right leg (HBI N 18501 - 1); B. H. cucullus sp. nov., right leg (SAM NN 19582); C. H. lodicula comb. nov., left leg (QM S 116469); D. H. megacantha sp. nov., left leg (QM S 116493); E. H. porongurup sp. nov., left leg (WAM T 75419); F. H. tatianeae sp. nov., left leg (MV K- 14612); G. H. transmarina comb. nov., right leg (QM S 20307); H. H. urbana comb. nov., left leg (QM S 111900); I. H. walesiana comb. nov., left leg (WAM T 75383); J. H. yesabah sp. nov., left leg (QM S 111896). Scale bars: 2 mm (A-C, E, F, H-J); 1 mm (D); 5 mm (G)." httpUri="https://binary.pensoft.net/fig/606182" pageId="0" pageNumber="275">Figs 2I</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">, 3D</figureCitation>
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">, 27</figureCitation>
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">, 28</figureCitation>
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution records of Hortophora urbana comb. nov. in Australia." httpUri="https://binary.pensoft.net/fig/606209" pageId="0" pageNumber="275">, 29</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="reference_group">
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName authority="Walckenaer, 1805" authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Framenau & Baptista & Oliveira & Castanheira" baseAuthorityYear="2021" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira urbana" order="Araneae" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="urbana" status="comb. nov.">Epeira urbana</taxonomicName>
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<bibRefCitation author="Keyserling, E" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="275" refId="B23" refString="Keyserling, E, 1887. Die Arachniden Australiens nach der Natur beschrieben und abgebildet. 2. Theil. 3. Lieferung. Bauer & Raspe, Nuernberg, pg. 153-232, pl. 12-20." title="Die Arachniden Australiens nach der Natur beschrieben und abgebildet. 2. Theil. 3. Lieferung. Bauer & Raspe, Nuernberg, pg. 153 - 232, pl. 12 - 20." year="1887">Keyserling 1887</bibRefCitation>
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: 160-161, plate 13, figs 5, 5a.
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</paragraph>
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus urbanus" order="Araneae" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="urbanus" status="comb. nov.">Araneus urbanus</taxonomicName>
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(Keyserling).-
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<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.9.1911.928" author="Rainbow, WJ" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="275" pagination="107 - 319" refId="B38" refString="Rainbow, WJ, 1911. A census of Australian Araneidae. Records of the Australian Museum 9: 107 - 319, DOI: https://doi.org/10.3853/j.0067-1975.9.1911.928" title="A census of Australian Araneidae." url="https://doi.org/10.3853/j.0067-1975.9.1911.928" volume="9" year="1911">Rainbow 1911</bibRefCitation>
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: 195.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="type material">
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<paragraph pageId="0" pageNumber="275">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="275">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="275">Holotype</emphasis>
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of
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<taxonomicName authority="Walckenaer, 1805" authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Framenau & Baptista & Oliveira & Castanheira" baseAuthorityYear="2021" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira urbana" order="Araneae" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="urbana">
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<emphasis italics="true" pageId="0" pageNumber="275">Epeira urbana</emphasis>
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</taxonomicName>
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Keyserling, 1887: Male, Sydney (
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<geoCoordinate degrees="33" direction="south" minutes="52" orientation="latitude" precision="925" value="-33.866665">33°52'S</geoCoordinate>
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,
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<geoCoordinate degrees="151" direction="east" minutes="12" orientation="longitude" precision="925" value="151.2">151°12'E</geoCoordinate>
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, New South Wales, Australia), Bradley collection (considered lost; see
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<bibRefCitation DOI="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="275" pagination="265 - 292" refId="B9" refString="Framenau, VW, 2005. The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae). Records of the Western Australian Museum 22: 265 - 292, DOI: https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" title="The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae)." url="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" volume="22" year="2005">Framenau (2005)</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="materials_examined">
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<paragraph pageId="0" pageNumber="275">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="275">See Appendix 1.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="diagnosis">
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<paragraph pageId="0" pageNumber="275">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="275">
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Males of
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<taxonomicName class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
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<emphasis italics="true" pageId="0" pageNumber="275">H. urbana</emphasis>
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</taxonomicName>
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comb. nov. can easily be distinguished from all other
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<taxonomicName authorityName="Framenau & Castanheira" authorityYear="2021" family="Araneidae" genus="Hortophora" kingdom="Animalia" lsidName="Hortophora" order="Araneae" pageId="0" pageNumber="275" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="275">Hortophora</emphasis>
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</taxonomicName>
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gen. nov. species by distinct shape of the median apophysis of male pedipalp. Whilst it is elongate transverse with a central protrusion as in many other species, both the central protrusion and the apical tip are blunt with only very small teeth apically (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C</figureCitation>
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), whereas these are generally pointed in other species of similar size, such as
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<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Framenau & Baptista & Oliveira & Castanheira" baseAuthorityYear="2021" class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. biapicata" order="Araneae" pageId="0" pageNumber="275" rank="species" species="biapicata">
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<emphasis italics="true" pageId="0" pageNumber="275">H. biapicata</emphasis>
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</taxonomicName>
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comb. nov. (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hortophora biapicata comb. nov., male and female genitalia. A. left male pedipalp, ventral view (HBI N 18501 - 1); B. left male pedipalp, ventral view (HBI N 18501 - 1); C. female epigyne, ventral view (HBI N 26183 - 1); D. female epigyne, lateral view (HBI 26183 - 1); E. female epigyne, posterior view (HBI N 26183 - 1); F. male left coxae of legs I and II showing coxal hooks, ventral view (HBI N 18501 - 1); G. female epigyne, ventral view (NHMD 12371); H. right male pedipalp, ventral view (NHMD 12380). Scale bars: 1 mm (A-E, G, H); 0.5 mm (F)." httpUri="https://binary.pensoft.net/fig/606187" pageId="0" pageNumber="275">7A, H</figureCitation>
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) and
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<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Framenau & Baptista & Oliveira & Castanheira" baseAuthorityYear="2021" class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. transmarina" order="Araneae" pageId="0" pageNumber="275" rank="species" species="transmarina">
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<emphasis italics="true" pageId="0" pageNumber="275">H. transmarina</emphasis>
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</taxonomicName>
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comb. nov. (Fig.
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Hortophora transmarina comb. nov., male (WAM T 67957). A. dorsal habitus; B. ventral habitus; C. left pedipalp, ventral view; D. left pedipalp, dorsal view. Scale bars: 5 mm (A, B); 1 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606204" pageId="0" pageNumber="275">24C</figureCitation>
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). The epigyne scape of
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<taxonomicName class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
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<emphasis italics="true" pageId="0" pageNumber="275">H. urbana</emphasis>
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</taxonomicName>
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comb. nov. is unlike that of any other species, as it is comparatively strong, centrally widened and has distinct transverse wrinkles (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28C-E</figureCitation>
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).
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275" start="Figure 27" startId="F27">
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<paragraph pageId="0" pageNumber="275">
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<emphasis bold="true" pageId="0" pageNumber="275">Figure 27.</emphasis>
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<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Keyserling" baseAuthorityYear="1887" family="Araneidae" genus="Hortophora" kingdom="Animalia" lsidName="Hortophora urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
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<emphasis italics="true" pageId="0" pageNumber="275">Hortophora urbana</emphasis>
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</taxonomicName>
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comb. nov., male (QM S111900).
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<emphasis bold="true" pageId="0" pageNumber="275">A.</emphasis>
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dorsal habitus;
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<emphasis bold="true" pageId="0" pageNumber="275">B.</emphasis>
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ventral habitus;
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<emphasis bold="true" pageId="0" pageNumber="275">C.</emphasis>
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right pedipalp, ventral view;
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<emphasis bold="true" pageId="0" pageNumber="275">D.</emphasis>
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right pedipalp, dorsal view. Scale bars: 2 mm (
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<emphasis bold="true" pageId="0" pageNumber="275">A, B</emphasis>
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); 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="275">C, D</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="description">
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<paragraph pageId="0" pageNumber="275">Description.</paragraph>
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<paragraph pageId="0" pageNumber="275">
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<emphasis bold="true" pageId="0" pageNumber="275">Male</emphasis>
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(QM S111900): Total length 10.0. Carapace 5.1 long, 4.7 wide, reddish-brown, anterior cephalic area somewhat lighter (Fig.
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27A</figureCitation>
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). Eye diameter AME 0.31, ALE 0.18, PME 0.20, PLE 0.18; row of eyes: AME 0.83, PME 0.59, PLE 2.48. Chelicerae brown; three promarginal teeth (median largest) and three retromarginal teeth (basal largest). Legs orange-brown (Fig.
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27A, B</figureCitation>
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). Tibiae of leg II with cluster of thick setae prolaterally, but no apico-ventral megaspur (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hortophora gen. nov. species, second legs of male, prolateral view. A. H. biapicata comb. nov., right leg (HBI N 18501 - 1); B. H. cucullus sp. nov., right leg (SAM NN 19582); C. H. lodicula comb. nov., left leg (QM S 116469); D. H. megacantha sp. nov., left leg (QM S 116493); E. H. porongurup sp. nov., left leg (WAM T 75419); F. H. tatianeae sp. nov., left leg (MV K- 14612); G. H. transmarina comb. nov., right leg (QM S 20307); H. H. urbana comb. nov., left leg (QM S 111900); I. H. walesiana comb. nov., left leg (WAM T 75383); J. H. yesabah sp. nov., left leg (QM S 111896). Scale bars: 2 mm (A-C, E, F, H-J); 1 mm (D); 5 mm (G)." httpUri="https://binary.pensoft.net/fig/606182" pageId="0" pageNumber="275">2H</figureCitation>
|
|
). Leg formula I> IV> II> III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I - 6.4 + 2.5 + 4.8 + 5.2 + 1.4 = 20.3, II - 5.8 + 2.2 + 4.5 + 0.9 + 1.4 = 14.8, III - 3.6 + 1.4 + 2.3 + 2.3 + 1.0 = 10.6, IV - 4.6 + 2.0 + 3.5 + 3.8 + 1.1 = 15.0. Labium 0.63 long, 0.85 wide, reddish-brown; endites dark brown (Fig.
|
|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27B</figureCitation>
|
|
). Sternum 2.3 long, 1.9 wide, irregular light brown-brown, covered with white setae (Fig.
|
|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27B</figureCitation>
|
|
). Abdomen 5.0 long, 4.5 wide, dorsum with indistinct humeral humps, beige with darker folium pattern and central brown line (Fig.
|
|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27A</figureCitation>
|
|
); venter olive-grey with indistinct lateral white lines (Fig.
|
|
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Hortophora walesiana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsalview. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606210" pageId="0" pageNumber="275">30B</figureCitation>
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|
). Pedipalp length of segments (femur + patella + tibia + cymbium = total length): 1.2 + 0.5 + 0.4 + 2.1 = 4.2; paracymbium elongated and curved, ending on a hook-like rounded tip (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C, D</figureCitation>
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|
); median apophysis elongated transverse with blunt rounded central protrusion and indistinct tips (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C</figureCitation>
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); conductor lobe of standard shape and size (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C</figureCitation>
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); terminal apophysis deflated, ending in an elongated, sclerotised tip (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C</figureCitation>
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|
); conductor heavily sclerotized ending in two rounded projections (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
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,
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|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C, D</figureCitation>
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); embolus elongated and sinuous (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hortophora gen. nov., left male pedipalps, ventral view A. H. biapicata comb. nov. expanded (WAM T 77043); B. H. biapicata comb. nov. (WAM T 68036); C. H. tatianeae sp. nov. (WAM T 68012); D. H. urbana comb. nov. (AM KS 66276). Scale bars: 1 mm (A), 2 mm (B-D)." httpUri="https://binary.pensoft.net/fig/606183" pageId="0" pageNumber="275">3D</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Hortophora urbana comb. nov., male (QM S 111900). A. dorsal habitus; B. ventral habitus; C. right pedipalp, ventral view; D. right pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.5 mm (C, D)." httpUri="https://binary.pensoft.net/fig/606207" pageId="0" pageNumber="275">27C</figureCitation>
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).
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|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" pageId="0" pageNumber="275">Female</emphasis>
|
|
(QM S111901): Total length 17.00. Carapace 4.5 long, 6.1 wide; very dark brown, cephalic area lighter and covered with white setae (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28A</figureCitation>
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|
). Eye diameter AME 0.90, PME 0.67, PLE 3.52. Chelicerae reddish-brown; four promarginal teeth (apical and third largest) and three retrolateral (similar size). Leg colouration as in male (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28A, B</figureCitation>
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|
). Pedipalp length of segments (femur + patella + tibia + tarsus = total length): 2.2 + 1.0 + 1.3 + 2.7 = 7.2. Leg formula I> IV> II> III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I - 7.5 + 3.8 + 6.3 + 6.6 + 2.0 = 26.2, II - 6.5 + 3.5 + 5.8 + 0.9 + 1.9 = 1.6, III - 5.0 + 1.7 + 3.0 + 3.0 + 1.3 = 14.00, IV - 6.7 + 3.1 + 4.5 + 5.0 + 1.6 = 20.9. Labium 0.90 long, 1.53 wide, as in male; endites as in male (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28B</figureCitation>
|
|
). Sternum 3.2 long, 2.7 wide, reddish-brown, centrally somewhat darker (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28B</figureCitation>
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). Abdomen 13.0 long, 11.0 wide, similar as male but colouration poorly preserved (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28A, B</figureCitation>
|
|
). Epigyne (QM S111902) (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28C-E</figureCitation>
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) wider than long; scape thick and wrinkled, constricted anteriorly with very sclerotized and black lateral edges.
|
|
</paragraph>
|
|
<caption httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275" start="Figure 28" startId="F28">
|
|
<paragraph pageId="0" pageNumber="275">
|
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<emphasis bold="true" pageId="0" pageNumber="275">Figure 28.</emphasis>
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<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Keyserling" baseAuthorityYear="1887" family="Araneidae" genus="Hortophora" kingdom="Animalia" lsidName="Hortophora urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
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<emphasis italics="true" pageId="0" pageNumber="275">Hortophora urbana</emphasis>
|
|
</taxonomicName>
|
|
comb. nov., female.
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A.</emphasis>
|
|
dorsal habitus (QM S111901);
|
|
<emphasis bold="true" pageId="0" pageNumber="275">B.</emphasis>
|
|
ventral habitus (QM S111901);
|
|
<emphasis bold="true" pageId="0" pageNumber="275">C.</emphasis>
|
|
epigyne, ventral view (QM S111901);
|
|
<emphasis bold="true" pageId="0" pageNumber="275">D.</emphasis>
|
|
epigyne, lateral view (QM S111902);
|
|
<emphasis bold="true" pageId="0" pageNumber="275">E.</emphasis>
|
|
epigyne, posterior view (QM S111902). Scale bars: 5 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A, B</emphasis>
|
|
); 0.5mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">C-E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="variation">
|
|
<paragraph pageId="0" pageNumber="275">Variation.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Size variation: total length males 10.0-13.1 (n=4), females 13.1-20.0 (n=8). The epigyne scape was broken off in half the females measured here. Abdominal colour patterns are fairly variable in
|
|
<taxonomicName class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">H. urbana</emphasis>
|
|
</taxonomicName>
|
|
comb. nov. male and females, from dark specimens to very light ones as illustrated here. The folium pattern is generally not very distinct, but light guanine spot and lines are frequent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="remarks">
|
|
<paragraph pageId="0" pageNumber="275">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
The holotype of
|
|
<taxonomicName authority="Walckenaer, 1805" authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Framenau & Baptista & Oliveira & Castanheira" baseAuthorityYear="2021" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira urbana" order="Araneae" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="urbana">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Epeira urbana</emphasis>
|
|
</taxonomicName>
|
|
Keyserling, 1887 was part of the Bradley collection and is considered lost (
|
|
<bibRefCitation DOI="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="275" pagination="265 - 292" refId="B9" refString="Framenau, VW, 2005. The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae). Records of the Western Australian Museum 22: 265 - 292, DOI: https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" title="The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae)." url="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" volume="22" year="2005">Framenau. 2005</bibRefCitation>
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|
,
|
|
<bibRefCitation DOI="https://doi.org/10.3897/evolsyst.3.33454" author="Framenau, VW" journalOrPublisher="Evolutionary Systematics" pageId="0" pageNumber="275" pagination="1 - 27" refId="B11" refString="Framenau, VW, 2019. Generic and family transfers, and nomina dubia for orb-weaving spiders (Araneae, Araneidae) in the Australasian, Oriental and Pacific regions. Evolutionary Systematics 3: 1 - 27, DOI: https://doi.org/10.3897/evolsyst.3.33454" title="Generic and family transfers, and nomina dubia for orb-weaving spiders (Araneae, Araneidae) in the Australasian, Oriental and Pacific regions." url="https://doi.org/10.3897/evolsyst.3.33454" volume="3" year="2019">2019</bibRefCitation>
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|
), but we do not consider it necessary to designate a neotype, as the original illustrations of the male pedipalp in combination with the distribution of the species allow an accurate identification.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
We described the best preserved female available to us, but as the scape of this specimens was broken off (Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hortophora urbana comb. nov., female. A. dorsal habitus (QM S 111901); B. ventral habitus (QM S 111901); C. epigyne, ventral view (QM S 111901); D. epigyne, lateral view (QM S 111902); E. epigyne, posterior view (QM S 111902). Scale bars: 5 mm (A, B); 0.5 mm (C-E)." httpUri="https://binary.pensoft.net/fig/606208" pageId="0" pageNumber="275">28B</figureCitation>
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), we described an intact scape of a different female.
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|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="life history and habitat preferences">
|
|
<paragraph pageId="0" pageNumber="275">Life history and habitat preferences.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Mature males of
|
|
<taxonomicName class="Arachnida" family="Araneidae" genus="H." kingdom="Animalia" lsidName="H. urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">H. urbana</emphasis>
|
|
</taxonomicName>
|
|
comb. nov. were collected in November and December and mature females from December to April indicating summer-maturity (equivalent to the wet season in northern latitudes). Habitat descriptions with museum specimens include rainforest and softwood scrub, but a single record is from a suburban clothes line.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="distribution">
|
|
<paragraph pageId="0" pageNumber="275">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Keyserling" baseAuthorityYear="1887" family="Araneidae" genus="Hortophora" kingdom="Animalia" lsidName="Hortophora urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Hortophora urbana</emphasis>
|
|
</taxonomicName>
|
|
comb. nov. has mainly been found along the east coast of Australia from southern New South Wales to northern Queensland, but occurs into northern Western Australia (Fig.
|
|
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution records of Hortophora urbana comb. nov. in Australia." httpUri="https://binary.pensoft.net/fig/606209" pageId="0" pageNumber="275">29</figureCitation>
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|
).
|
|
</paragraph>
|
|
<caption httpUri="https://binary.pensoft.net/fig/606209" pageId="0" pageNumber="275" start="Figure 29" startId="F29">
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" pageId="0" pageNumber="275">Figure 29.</emphasis>
|
|
Distribution records of
|
|
<taxonomicName authorityName="Framenau & Baptista & Oliveira & Castanheira" authorityYear="2021" baseAuthorityName="Keyserling" baseAuthorityYear="1887" family="Araneidae" genus="Hortophora" kingdom="Animalia" lsidName="Hortophora urbana" order="Araneae" pageId="0" pageNumber="275" rank="species" species="urbana">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Hortophora urbana</emphasis>
|
|
</taxonomicName>
|
|
comb. nov. in Australia.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
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|
</document> |