219 lines
18 KiB
XML
219 lines
18 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFBEFFFBFF6D6DEACA93FD4D" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma alaemacula Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="23" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="22" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894880" ID-Zenodo-Dep="7894880" LSID="urn:lsid:plazi:treatment:A148D509FFBEFFFBFF6D6DEACA93FD4D" httpUri="http://treatment.plazi.org/id/A148D509FFBEFFFBFF6D6DEACA93FD4D" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection pageId="21" pageNumber="22" type="nomenclature">
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<paragraph blockId="21.[151,1200,1726,1825]" box="[151,521,1726,1753]" pageId="21" pageNumber="22">
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<heading bold="true" box="[151,521,1726,1753]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis bold="true" box="[151,521,1726,1753]" pageId="21" pageNumber="22">
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,447,1726,1753]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="alaemacula" status="n. sp.">
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<emphasis bold="true" box="[151,447,1726,1753]" italics="true" pageId="21" pageNumber="22">Leptostigma alaemacula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[454,521,1727,1753]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="21.[151,1200,1726,1825]" box="[151,1200,1763,1790]" pageId="21" pageNumber="22">
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http://zoobank.org/
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<uri box="[370,1200,1763,1790]" pageId="21" pageNumber="22">
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urn:lsid:zoobank.org:act:
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<uuid box="[640,1200,1763,1790]" pageId="21" pageNumber="22">CB061CC9-BB17-4FED-B536-2A35796FCBBA</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,238,1799,1825]" pageId="21" pageNumber="22" type="description">
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<paragraph blockId="21.[151,1200,1726,1825]" box="[151,238,1799,1825]" pageId="21" pageNumber="22">
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<figureCitation box="[151,234,1799,1825]" captionStart="FIGURE 21" captionStartId="22.[151,250,1502,1526]" captionTargetBox="[151,1436,668,1465]" captionTargetId="figure-232@22.[151,1436,668,1478]" captionTargetPageId="22" captionText="FIGURE 21. Leptostigma alaemaculum holotype SR 92-18-08, forewing: A, photograph; B, drawing. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894914" httpUri="https://zenodo.org/record/7894914/files/figure.png" pageId="21" pageNumber="22">Fig. 21</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="materials_examined">
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<paragraph blockId="21.[151,1437,1871,2041]" pageId="21" pageNumber="22">
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<materialsCitation collectingDate="1992-09-25" collectionCode="A, SR" collectorName="Carma Henry & Republic" pageId="21" pageNumber="22" specimenCode="SR 92-18, 08" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[151,260,1871,1897]" pageId="21" pageNumber="22">
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<typeStatus box="[151,260,1871,1897]" pageId="21" pageNumber="22">Holotype</typeStatus>
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</emphasis>
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.
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<specimenCode box="[273,379,1871,1897]" collectionCode="SR" country="China" lsid="urn:lsid:biocol.org:col:14050" name="Sichuan Institute of Natural Resources" pageId="21" pageNumber="22" type="Herbarium">SR 92-18</specimenCode>
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-
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<specimenCode box="[387,417,1871,1897]" collectionCode="SR" country="China" lsid="urn:lsid:biocol.org:col:14050" name="Sichuan Institute of Natural Resources" pageId="21" pageNumber="22" type="Herbarium">08</specimenCode>
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(
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<figureCitation box="[431,514,1871,1897]" captionStart="FIGURE 21" captionStartId="22.[151,250,1502,1526]" captionTargetBox="[151,1436,668,1465]" captionTargetId="figure-232@22.[151,1436,668,1478]" captionTargetPageId="22" captionText="FIGURE 21. Leptostigma alaemaculum holotype SR 92-18-08, forewing: A, photograph; B, drawing. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894914" httpUri="https://zenodo.org/record/7894914/files/figure.png" pageId="21" pageNumber="22">Fig. 21</figureCitation>
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).
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<collectionCode box="[532,551,1871,1897]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="21" pageNumber="22" type="Herbarium">A</collectionCode>
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forewing, collected by
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<collectorName box="[812,963,1871,1897]" pageId="21" pageNumber="22">Carma Henry</collectorName>
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at
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<collectorName box="[996,1097,1871,1897]" pageId="21" pageNumber="22">Republic</collectorName>
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on
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<date box="[1138,1364,1871,1898]" pageId="21" pageNumber="22" value="1992-09-25">
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<collectingDate box="[1138,1364,1871,1898]" pageId="21" pageNumber="22" value="1992-09-25">September 25, 1992</collectingDate>
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</date>
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, in the collections of the
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<collectionCode box="[351,389,1907,1933]" country="China" lsid="urn:lsid:biocol.org:col:14050" name="Sichuan Institute of Natural Resources" pageId="21" pageNumber="22" type="Herbarium">SR</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph blockId="21.[151,1437,1871,2041]" pageId="21" pageNumber="22">
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<emphasis bold="true" box="[199,322,1943,1969]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
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Distinguished from other species of
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<taxonomicName authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[732,870,1944,1969]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="genus">
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<emphasis box="[732,870,1944,1969]" italics="true" pageId="21" pageNumber="22">Leptostigma</emphasis>
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</taxonomicName>
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by a combination of forewing dark in cell 1r+2rm. Tentatively diagnostic: R+M shorter than 1-M; 2r-rs joins cell 3rm distinctly near its middle; 2r-m joins M scarcely basal to 2m-cu; anal cells widely separated by fusion of 1A and 2A+3A, without crossvein.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="description">
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<paragraph blockId="22.[151,1437,150,645]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[199,345,150,176]" pageId="22" pageNumber="23">Description.</emphasis>
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<typeStatus box="[350,458,150,176]" pageId="22" pageNumber="23">Holotype</typeStatus>
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. Forewing length
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<quantity box="[653,753,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="22" pageNumber="23" unit="mm" value="14.2">14.2 mm</quantity>
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, maximum width
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<quantity box="[950,1036,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="22" pageNumber="23" unit="mm" value="4.1">4.1 mm</quantity>
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, length/width 3.5; membrane lightly infuscate throughout except darker posterior to 1r-rs; pterostigma dark; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r; 2r-rs shorter than Rs between 2r-m, 2r-rs, joining cell 3rm near its middle; 2r-m slightly curved, joins Rs notably basal to 2m-cu joining M, aligned with 2mcu; 1-M almost straight, sub-parallel to 1m-cu; 3r-m distinctly bowed toward base; cell 2mcu with M longer than 1m-cu; 2m-cu curved distinctly toward base; cu-a postfurcal; anal cells separated for distance greater than width of cell 2a, without crossvein between them.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="discussion">
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<paragraph blockId="22.[151,1437,150,645]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[199,315,366,392]" pageId="22" pageNumber="23">Remarks.</emphasis>
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Differs from
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[467,616,366,392]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="brevilatum" status="n. sp.">
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<emphasis box="[467,616,366,392]" italics="true" pageId="22" pageNumber="23">L. brevilatum</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[623,685,366,392]" pageId="22" pageNumber="23">
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<taxonomicNameLabel box="[623,685,366,392]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[698,845,366,392]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="longiclava" status="n. sp.">
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<emphasis box="[698,845,366,392]" italics="true" pageId="22" pageNumber="23">L. longiclava</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[851,913,366,392]" pageId="22" pageNumber="23">
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<taxonomicNameLabel box="[851,913,366,392]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[927,1113,366,392]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="longipallidum" status="n. sp.">
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<emphasis box="[927,1113,366,392]" italics="true" pageId="22" pageNumber="23">L. longipallidum</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1119,1181,366,392]" pageId="22" pageNumber="23">
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<taxonomicNameLabel box="[1119,1181,366,392]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1195,1408,366,392]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="longitenebricum" status="n. sp.">
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<emphasis box="[1195,1408,366,392]" italics="true" pageId="22" pageNumber="23">L. longitenebricum</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="22" pageNumber="23">
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<taxonomicNameLabel pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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by its forewing dark posterior to 1r-rs and 2r-m joins M close to 2m-cu. Differs from
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1148,1289,403,428]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="proxivena" status="n. sp.">
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<emphasis box="[1148,1289,403,428]" italics="true" pageId="22" pageNumber="23">L. proxivena</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1296,1359,402,428]" pageId="22" pageNumber="23">
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<taxonomicNameLabel box="[1296,1359,402,428]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="fasciatum" status="n. sp.">
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<emphasis italics="true" pageId="22" pageNumber="23">L. fasciatum</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[267,330,438,464]" pageId="22" pageNumber="23">
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<taxonomicNameLabel box="[267,330,438,464]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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in which the anal cells are not separated by a fusion of 1A with 2A+3A.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,429,474,501]" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph blockId="22.[151,1437,150,645]" box="[199,429,474,501]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[199,303,474,500]" pageId="22" pageNumber="23">Material</emphasis>
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.
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<typeStatus box="[317,425,474,500]" pageId="22" pageNumber="23">Holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="etymology">
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<paragraph blockId="22.[151,1437,150,645]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[199,334,510,536]" pageId="22" pageNumber="23">Etymology.</emphasis>
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From the Latin
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<emphasis box="[515,551,510,536]" italics="true" pageId="22" pageNumber="23">ala</emphasis>
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, meaning wing, and
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<emphasis box="[785,867,510,536]" italics="true" pageId="22" pageNumber="23">macula</emphasis>
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, spot, referring to the forewing spot. The epithet is a noun in apposition.
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</paragraph>
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<paragraph blockId="22.[151,1437,150,645]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[199,402,582,608]" pageId="22" pageNumber="23">Locality and age.</emphasis>
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<materialsCitation collectionCode="A" country="United States of America" county="Mountain Formation" location="Republic" municipality="Klondike Mountain" pageId="22" pageNumber="23" specimenCode="A0307" specimenCount="1" stateProvince="Washington">
|
||
<location LSID="urn:lsid:plazi:treatment:A148D509FFBEFFFBFF6D6DEACA93FD4D:2C3E32C4FFBDFFFBFE636913CA79FDA9" box="[409,513,582,608]" country="United States of America" county="Mountain Formation" municipality="Klondike Mountain" name="Republic" pageId="22" pageNumber="23" stateProvince="Washington">Republic</location>
|
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,
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<collectingRegion box="[523,659,582,608]" country="United States of America" name="Washington" pageId="22" pageNumber="23">Washington</collectingRegion>
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,
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<collectingCountry box="[669,952,582,608]" name="United States of America" pageId="22" pageNumber="23">United States of America</collectingCountry>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:A148D509FFBEFFFBFF6D6DEACA93FD4D:2C3E32C4FFBDFFFBFC386912CC90FDA9" box="[962,1256,582,609]" country="United States of America" county="Mountain Formation" municipality="Klondike Mountain" name="Tom Thumb Tuff Member" pageId="22" pageNumber="23" stateProvince="Washington">Tom Thumb Tuff Member</location>
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of the
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<location LSID="urn:lsid:plazi:treatment:A148D509FFBEFFFBFF6D6DEACA93FD4D:2C3E32C4FFBDFFFBFACE6913C9F8FD4D" country="United States of America" county="Mountain Formation" municipality="Klondike Mountain" name="Klondike Mountain Formation" pageId="22" pageNumber="23" stateProvince="Washington">
|
||
<collectingMunicipality pageId="22" pageNumber="23">
|
||
Klondike
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<collectingCounty box="[151,260,618,644]" pageId="22" pageNumber="23">Mountain</collectingCounty>
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</collectingMunicipality>
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Formation
|
||
</location>
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||
exposure
|
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<specimenCode box="[497,577,618,644]" collectionCode="A" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="22" pageNumber="23" type="Herbarium">A0307</specimenCode>
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; mid-Ypresian
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||
</materialsCitation>
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||
.
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |