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<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce &amp; Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFAFFFEBFF6D6D18CCA3FE88" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Allenbycimbex Archibald &amp; Rasnitsyn 2023, n. gen." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, Symphyta) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="5" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Cimbicidae (Hymenoptera, Symphyta) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 &amp; Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America &amp; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia &amp; Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-05-04</mods:number>
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<mods:detail type="volume">
<mods:number>5278</mods:number>
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<treatment LSID="urn:lsid:plazi:treatment:A148D509FFAFFFEBFF6D6D18CCA3FE88" httpUri="http://treatment.plazi.org/id/A148D509FFAFFFEBFF6D6D18CCA3FE88" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
<paragraph blockId="4.[151,1189,1613,1712]" box="[151,420,1613,1640]" pageId="4" pageNumber="5">
<heading bold="true" box="[151,420,1613,1640]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis bold="true" box="[151,420,1613,1640]" pageId="4" pageNumber="5">
<taxonomicName authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[151,330,1613,1640]" class="Insecta" family="Cimbicidae" genus="Allenbycimbex" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="n. gen.">
<emphasis bold="true" box="[151,330,1613,1640]" italics="true" pageId="4" pageNumber="5">Allenbycimbex</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[338,420,1614,1640]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[151,1189,1613,1712]" box="[151,1189,1650,1676]" pageId="4" pageNumber="5">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection box="[151,223,1686,1712]" pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[151,1189,1613,1712]" box="[151,223,1686,1712]" pageId="4" pageNumber="5">
<figureCitation box="[151,220,1686,1712]" captionStart="FIGURE 4" captionStartId="6.[151,250,1327,1351]" captionTargetBox="[155,1430,525,1300]" captionTargetId="figure-85@6.[151,1436,525,1304]" captionTargetPageId="6" captionText="FIGURE 4. Allenbycimbex morrisae, holotype RBCM.EH2007.002.0001: A, photograph; B, drawing; C, drawing of right wing; D, drawing of left wing. Scale bar for A and B is 5 mm (as the fossil has been stretched by geologic shear forces, C and D are altered to the presumed unstretched configuration)." figureDoi="http://doi.org/10.5281/zenodo.7894842" httpUri="https://zenodo.org/record/7894842/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,606,1754,1778]" pageId="4" pageNumber="5" type="type_taxon">
<paragraph blockId="4.[151,606,1754,1778]" box="[151,606,1754,1778]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[151,291,1754,1778]" pageId="4" pageNumber="5">
<typeStatus box="[151,204,1754,1778]" pageId="4" pageNumber="5">Type</typeStatus>
species.
</emphasis>
<taxonomicName authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[298,544,1754,1778]" class="Insecta" family="Cimbicidae" genus="Allenbycimbex" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="morrisae" status="n. sp.">
<emphasis box="[298,544,1754,1778]" italics="true" pageId="4" pageNumber="5">Allenbycimbex morrisae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[551,606,1755,1778]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="diagnosis">
<paragraph blockId="4.[151,1437,1824,1995]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[151,274,1824,1850]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
Distinguished from
<taxonomicName authorityName="Nel" authorityYear="2004" box="[499,637,1824,1850]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[499,637,1824,1850]" italics="true" pageId="4" pageNumber="5">Cenocimbex</emphasis>
</taxonomicName>
by R lacking dark triangular sclerite before base of pterostigma reaching 1-Rs+M near 1m-cu; wing narrower (length/width
<emphasis box="[728,754,1861,1886]" italics="true" pageId="4" pageNumber="5">ca</emphasis>
. 3.0 as reconstructed) [
<taxonomicName authorityName="Nel" authorityYear="2004" box="[1021,1159,1860,1886]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[1021,1159,1860,1886]" italics="true" pageId="4" pageNumber="5">Cenocimbex</emphasis>
</taxonomicName>
: 2.5]; cell 3rm long (2rm to 3r-m distinctly longer than length of 3r-m); hind wing with m-cu placed much basally than in
<taxonomicName authorityName="Nel" authorityYear="2004" box="[1290,1428,1896,1922]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[1290,1428,1896,1922]" italics="true" pageId="4" pageNumber="5">Cenocimbex</emphasis>
</taxonomicName>
; distinguished from
<taxonomicName authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[368,506,1933,1958]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis box="[368,506,1933,1958]" italics="true" pageId="4" pageNumber="5">Leptostigma</emphasis>
</taxonomicName>
by pterostigma broad, maximum height distinctly exceeding height of C to R basal to pterostigma.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7894836" ID-Zenodo-Dep="7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1833,1857]" targetBox="[192,1395,191,1809]" targetPageId="5" targetType="figure">
<paragraph blockId="5.[151,1437,1833,2037]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[151,278,1833,1857]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
Heads of
<taxonomicName authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[393,562,1833,1857]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Cenocimbicinae">Cenocimbicinae</taxonomicName>
: A,
<taxonomicName authority="GSC" authorityName="GSC" box="[607,910,1833,1857]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="brevilatum">
<emphasis box="[607,852,1833,1857]" italics="true" pageId="5" pageNumber="6">Leptostigma brevilatum</emphasis>
GSC
</taxonomicName>
142839, drawing; B,
<taxonomicName authority="GSC" authorityName="GSC" box="[1146,1344,1833,1857]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="brevilatum">
<emphasis box="[1146,1285,1833,1857]" italics="true" pageId="5" pageNumber="6">L. brevilatum</emphasis>
GSC
</taxonomicName>
142839, photograph; C,
<taxonomicName authority="GSC" authorityName="GSC" box="[326,671,1869,1893]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longipallidum">
<emphasis box="[326,607,1869,1893]" italics="true" pageId="5" pageNumber="6">Leptostigma longipallidum</emphasis>
GSC
</taxonomicName>
142841, drawing; D,
<taxonomicName authority="GSC" authorityName="GSC" box="[926,1166,1869,1893]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longipallidum">
<emphasis box="[926,1103,1869,1893]" italics="true" pageId="5" pageNumber="6">L. longipallidum</emphasis>
GSC
</taxonomicName>
142841, photograph; E,
<taxonomicName authorityName="Nel, 2004 Nel" authorityYear="2004" box="[151,399,1905,1929]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="menatensis">
<emphasis box="[151,399,1905,1929]" italics="true" pageId="5" pageNumber="6">Cenocimbex menatensis</emphasis>
</taxonomicName>
MNHN-LP-R.11195, drawing; F,
<taxonomicName authorityName="Nel, 2004 Nel" authorityYear="2004" box="[763,909,1905,1929]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="menatensis">
<emphasis box="[763,909,1905,1929]" italics="true" pageId="5" pageNumber="6">C. menatensis</emphasis>
</taxonomicName>
MNHN-LP-R.11195, photograph; G,
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longipallidum">
<emphasis italics="true" pageId="5" pageNumber="6">Leptostigma longipallidum</emphasis>
</taxonomicName>
F-771, drawing; H,
<taxonomicName box="[503,672,1941,1965]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longipallidum">
<emphasis box="[503,672,1941,1965]" italics="true" pageId="5" pageNumber="6">L. longipallidum</emphasis>
</taxonomicName>
F-771, photograph; I,
<taxonomicName authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[904,1202,1941,1965]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longitenebricum">
<emphasis box="[904,1202,1941,1965]" italics="true" pageId="5" pageNumber="6">Leptostigma longitenebricum</emphasis>
</taxonomicName>
F-1565, drawing; J,
<taxonomicName authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longitenebricum">
<emphasis italics="true" pageId="5" pageNumber="6">L. longitenebricum</emphasis>
</taxonomicName>
F-1565, photograph; K,
<taxonomicName authority="RBCM. EH" authorityName="RBCM. EH" box="[581,943,1977,2001]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longiclava">
<emphasis box="[581,822,1977,2001]" italics="true" pageId="5" pageNumber="6">Leptostigma longiclava</emphasis>
RBCM.EH
</taxonomicName>
2017.050.4085.0001, drawing; L,
<taxonomicName authority="RBCM. EH" authorityName="RBCM. EH" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="longiclava">
<emphasis box="[1300,1436,1977,2001]" italics="true" pageId="5" pageNumber="6">L. longiclava</emphasis>
RBCM.EH
</taxonomicName>
2017.050.4085.0001, photograph. Scale bars 2 mm.
</paragraph>
</caption>
<paragraph blockId="6.[151,1436,150,321]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[199,402,150,177]" pageId="6" pageNumber="7">Locality and age.</emphasis>
<materialsCitation country="Canada" county="Species" location="Vermilion Bluffs Member" municipality="Princeton" pageId="6" pageNumber="7" specimenCount="1" stateProvince="British Columbia">
<location LSID="urn:lsid:plazi:treatment:A148D509FFAFFFEBFF6D6D18CCA3FE88:2C3E32C4FFADFFEBFE626BC2CAC1FF78" box="[408,697,151,177]" country="Canada" county="Species" municipality="Princeton" name="Vermilion Bluffs Member" pageId="6" pageNumber="7" stateProvince="British Columbia">Vermilion Bluffs Member</location>
of the
<location LSID="urn:lsid:plazi:treatment:A148D509FFAFFFEBFF6D6D18CCA3FE88:2C3E32C4FFADFFEBFCF96BC2CBA1FF78" box="[771,985,151,177]" country="Canada" county="Species" municipality="Princeton" name="Allenby Formation" pageId="6" pageNumber="7" stateProvince="British Columbia">Allenby Formation</location>
south of the village of
<collectingMunicipality box="[1239,1349,151,177]" pageId="6" pageNumber="7">Princeton</collectingMunicipality>
,
<collectingRegion country="Canada" name="British Columbia" pageId="6" pageNumber="7">British Columbia</collectingRegion>
,
<collectingCountry box="[276,364,187,213]" name="Canada" pageId="6" pageNumber="7">Canada</collectingCountry>
; mid-Ypresian.
<emphasis bold="true" box="[199,403,222,248]" pageId="6" pageNumber="7">
<collectingCounty box="[199,286,222,248]" pageId="6" pageNumber="7">Species</collectingCounty>
included.
</emphasis>
<typeStatus box="[409,465,223,249]" pageId="6" pageNumber="7">Type</typeStatus>
only.
<emphasis bold="true" box="[199,334,259,285]" pageId="6" pageNumber="7">Etymology.</emphasis>
Named for the Allenby Formation, where its only specimen was found. The name is to be treated with masculine grammatical gender since it is derived from
<taxonomicName authorityName=": Cockerell" authorityYear="1922" box="[824,910,295,321]" class="Insecta" family="Cimbicidae" genus="Cimbex" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis box="[824,910,295,321]" italics="true" pageId="6" pageNumber="7">Cimbex</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[968,1230,295,321]" pageId="6" pageNumber="7" pagination="1 - 1064" refId="ref19654" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
Taeger
<emphasis box="[1051,1107,295,321]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2010: 188
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>