140 lines
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140 lines
14 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFA8FFE9FF6D6C04CA1FFDD0" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cenocimbex Nel 2004" docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="4" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
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<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894838" ID-Zenodo-Dep="7894838" LSID="urn:lsid:plazi:treatment:A148D509FFA8FFE9FF6D6C04CA1FFDD0" httpUri="http://treatment.plazi.org/id/A148D509FFA8FFE9FF6D6C04CA1FFDD0" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection box="[151,512,1873,1900]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph blockId="3.[151,512,1873,1936]" box="[151,512,1873,1900]" pageId="3" pageNumber="4">
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<heading bold="true" box="[151,512,1873,1900]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis bold="true" box="[151,512,1873,1900]" pageId="3" pageNumber="4">
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Genus
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<taxonomicName authority="Nel, 2004" authorityName="Nel" authorityYear="2004" box="[238,512,1873,1900]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[238,388,1873,1900]" italics="true" pageId="3" pageNumber="4">Cenocimbex</emphasis>
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<bibRefCitation author="Nel, A." box="[396,512,1874,1900]" pageId="3" pageNumber="4" pagination="253 - 269" refId="ref18757" refString="Nel, A. (2004) New and poorly known Cenozoic sawflies of France (Hymenoptera, Tenthredinoidea, Pamphilioidea). Deutsche Entomologische Zeitschrift, 51, 253 - 269. https: // doi. org / 10.1002 / mmnd. 20040510208" type="journal article" year="2004">Nel, 2004</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,307,1910,1936]" pageId="3" pageNumber="4" type="description">
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<paragraph blockId="3.[151,512,1873,1936]" box="[151,307,1910,1936]" pageId="3" pageNumber="4">
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<figureCitation box="[151,229,1910,1936]" captionStart="FIGURE 2" captionStartId="4.[151,250,1147,1171]" captionTargetBox="[151,1391,564,1120]" captionTargetId="figure-185@4.[151,1436,561,1124]" captionTargetPageId="4" captionText="FIGURE 2. Cenocimbex menatensis Nel, holotype MNHN-LP-R.11195: A, extract from a reflectance transformation imaging (RTI) file (Cultural Heritage Imaging 2012); B, drawing of left forewing; C, drawing of right forewing preserved portion; D, left hind wing. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894834" httpUri="https://zenodo.org/record/7894834/files/figure.png" pageId="3" pageNumber="4">
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Figs.
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<geoCoordinate box="[213,229,1910,1936]" degrees="2, 3" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5555" value="2.3">2</geoCoordinate>
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</figureCitation>
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,
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<figureCitation box="[241,303,1910,1936]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="3" pageNumber="4">3E–F</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="type_taxon">
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<paragraph blockId="3.[151,547,1978,2002]" box="[151,547,1978,2002]" pageId="3" pageNumber="4">
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<emphasis bold="true" box="[151,286,1978,2002]" pageId="3" pageNumber="4">
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<typeStatus box="[151,204,1978,2002]" pageId="3" pageNumber="4">Type</typeStatus>
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species
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</emphasis>
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<taxonomicName authority="Nel, 2004" authorityName="Nel, 2004 Nel" authorityYear="2004" box="[291,543,1978,2002]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="menatensis">
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<emphasis box="[291,434,1979,2002]" italics="true" pageId="3" pageNumber="4">C. menatensis</emphasis>
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<bibRefCitation author="Nel, A." box="[440,543,1978,2002]" pageId="3" pageNumber="4" pagination="253 - 269" refId="ref18757" refString="Nel, A. (2004) New and poorly known Cenozoic sawflies of France (Hymenoptera, Tenthredinoidea, Pamphilioidea). Deutsche Entomologische Zeitschrift, 51, 253 - 269. https: // doi. org / 10.1002 / mmnd. 20040510208" type="journal article" year="2004">Nel, 2004</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="4.[151,1436,150,537]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[151,388,150,177]" pageId="4" pageNumber="5">Emended diagnosis.</emphasis>
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Separated from all other
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<taxonomicName authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[669,851,151,177]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Cenocimbicinae">Cenocimbicinae</taxonomicName>
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by R before base of pterostigma with dark triangular sclerite reaching 1-Rs+M near 1m-cu [vs. sclerotization not apparent, either absent or pale, small]; by wing broad, estimated length/width 2.5 [all others
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<emphasis box="[575,601,223,248]" italics="true" pageId="4" pageNumber="5">ca</emphasis>
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. 2.8 or more]; by cell 3rm short (2r-m to 3r-m length ≤ 3r-m [vs. distinctly longer]); from
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[318,456,259,284]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus" status="n. gen.">
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<emphasis box="[318,456,259,284]" italics="true" pageId="4" pageNumber="5">Leptostigma</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[463,542,259,285]" pageId="4" pageNumber="5">
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<taxonomicNameLabel box="[463,542,259,285]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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also by pterostigma broad, maximum height distinctly more than height of C to R basal to pterostigma [
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<taxonomicName authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[419,557,295,320]" class="Magnoliopsida" family="Rubiaceae" genus="Leptostigma" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus" status="n. gen.">
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<emphasis box="[419,557,295,320]" italics="true" pageId="4" pageNumber="5">Leptostigma</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[564,643,295,321]" pageId="4" pageNumber="5">
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<taxonomicNameLabel box="[564,643,295,321]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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: similar in height to C to R basal to pterostigma]. Hind wing with r-m, m-cu and cu-a only moderately distant from each other (for about length of m-cu, so cu-a at about midlength cell mcu) [in other genera usually well before mid-length].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="discussion">
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<paragraph blockId="4.[151,1436,150,537]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[199,315,403,429]" pageId="4" pageNumber="5">Remarks.</emphasis>
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Sclerotisation on R near the pterostigma, even though of much lesser size, is clearly seen in an extant cimbicine, see
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<bibRefCitation author="Vilhelmsen, L. & Smith, D. R. & Malagon-Aldana, L. A." box="[318,575,439,465]" pageId="4" pageNumber="5" pagination="1 - 36" refId="ref19867" refString="Vilhelmsen, L., Smith, D. R. & Malagon-Aldana, L. A. (2018) A review of the South American genera of Cimbicidae (Insecta, Hymenoptera). European Journal of Taxonomy, 482, 1 - 36. https: // doi. org / 10.5852 / ejt. 2018.482" type="journal article" year="2018">
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Vilhelmsen
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<emphasis box="[453,509,439,465]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
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2018
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</bibRefCitation>
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,
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<figureCitation box="[586,673,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1147,1171]" captionTargetBox="[151,1391,564,1120]" captionTargetId="figure-185@4.[151,1436,561,1124]" captionTargetPageId="4" captionText="FIGURE 2. Cenocimbex menatensis Nel, holotype MNHN-LP-R.11195: A, extract from a reflectance transformation imaging (RTI) file (Cultural Heritage Imaging 2012); B, drawing of left forewing; C, drawing of right forewing preserved portion; D, left hind wing. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894834" httpUri="https://zenodo.org/record/7894834/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[151,1436,150,537]" box="[199,1001,474,501]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[199,404,474,501]" pageId="4" pageNumber="5">Locality and age.</emphasis>
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Menat Formation; Puy-de-Dôme,
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<collectingCountry box="[792,872,475,501]" name="France" pageId="4" pageNumber="5">France</collectingCountry>
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; Selandian.
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</paragraph>
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<paragraph blockId="4.[151,1436,150,537]" box="[199,615,510,537]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[199,396,510,536]" pageId="4" pageNumber="5">Species included</emphasis>
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.
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<typeStatus box="[409,465,511,537]" pageId="4" pageNumber="5">Type</typeStatus>
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species only.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |