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<document ID-DOI="http://dx.doi.org/10.3897/jhr.87.72842" ID-GBIF-Dataset="3e3ca27b-44ad-4ea2-af6b-516245293ccf" ID-Pensoft-Pub="1314-2607-87-323" ID-Pensoft-UUID="D03A96052A8550F9918BB08ACA344FB9" ID-Zenodo-Dep="5811493" ID-ZooBank="E343379ED04447ABA1ED47B3F01F3E59" ModsDocID="1314-2607-87-323" checkinTime="1640348264987" checkinUser="pensoft" docAuthor="Talamas, Elijah J., Bremer, Jonathan S., Moore, Matthew R., Bon, Marie-Claude, Lahey, Zachary, Roberts, Cheryl G., Combee, Lynn A., McGathey, Natalie, van Noort, Simon, Timokhov, Alexander V., Hougardy, Evelyne &amp; Hogg, Brian" docDate="2021" docId="8F262AD752325B1B80F7C78C4024F0A3" docLanguage="en" docName="JourHymenoptRes 87: 323-480" docOrigin="Journal of Hymenoptera Research 87" docPubDate="2021-12-23" docSource="http://dx.doi.org/10.3897/jhr.87.72842" docTitle="Hadronotus Foerster 1856" docType="treatment" docVersion="4" id="D03A96052A8550F9918BB08ACA344FB9" lastPageNumber="323" masterDocId="D03A96052A8550F9918BB08ACA344FB9" masterDocTitle="A maximalist approach to the systematics of a biological control agent: Gryon aetherium Talamas, sp. nov. (Hymenoptera, Scelionidae)" masterLastPageNumber="480" masterPageNumber="323" pageNumber="323" updateTime="1678757109313" updateUser="pensoft">
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<mods:title>A maximalist approach to the systematics of a biological control agent: Gryon aetherium Talamas, sp. nov. (Hymenoptera, Scelionidae)</mods:title>
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<mods:namePart>Talamas, Elijah J.</mods:namePart>
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<mods:affiliation>Florida Department of Agriculture and Consumer Services, Gainesville, FL, USA</mods:affiliation>
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<mods:namePart>Moore, Matthew R.</mods:namePart>
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<mods:namePart>Bon, Marie-Claude</mods:namePart>
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<mods:namePart>Lahey, Zachary</mods:namePart>
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<mods:affiliation>Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, Columbus, OH, USA</mods:affiliation>
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<mods:namePart>Roberts, Cheryl G.</mods:namePart>
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<mods:namePart>McGathey, Natalie</mods:namePart>
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<mods:affiliation>Iziko South African Museum, Cape Town, South Africa</mods:affiliation>
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<mods:namePart>Timokhov, Alexander V.</mods:namePart>
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<treatment ID-GBIF-Taxon="191737334" LSID="urn:lsid:plazi:treatment:8F262AD752325B1B80F7C78C4024F0A3" httpUri="http://treatment.plazi.org/id/8F262AD752325B1B80F7C78C4024F0A3" lastPageNumber="323" pageId="0" pageNumber="323">
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<taxonomicName LSID="8F262AD7-5232-5B1B-80F7-C78C4024F0A3" authority="Foerster" authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">
Hadronotus
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="323" type="reference_group">
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Hadronotus</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1856
<emphasis bold="true" pageId="0" pageNumber="323">stat. rev.</emphasis>
: 101, 105 (original description. Type:
<taxonomicName authorityName="Foerster" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus exsculptus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="exsculptus">Hadronotus exsculptus</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, first included species, keyed. Synonymized by
<bibRefCitation DOI="https://doi.org/10.1080/03745481.1936.10801393" author="Nixon, GEJ" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="323" pagination="114 - 191" refId="B201" refString="Nixon, GEJ, 1936. The African species of Teleasinae (Hym., Proctotrupoidea, Fam. Scelionidae). Annals and Magazine of Natural History 17: 114 - 191, DOI: https://doi.org/10.1080/03745481.1936.10801393" title="The African species of Teleasinae (Hym., Proctotrupoidea, Fam. Scelionidae)." url="https://doi.org/10.1080/03745481.1936.10801393" volume="17" year="1936">Nixon (1936)</bibRefCitation>
,
<bibRefCitation author="Masner, L" journalOrPublisher="Casopis Ceskoslovenske Spolecnosti Entomologicke" pageId="0" pageNumber="323" pagination="157 - 168" refId="B149" refString="Masner, L, 1961. The genera Gryon Hal., Idris Foerst. and Hemisius Westw. (Hym., Scelionidae). Casopis Ceskoslovenske Spolecnosti Entomologicke 58: 157 - 168" title="The genera Gryon Hal., Idris Foerst. and Hemisius Westw. (Hym., Scelionidae)." volume="58" year="1961">Masner (1961)</bibRefCitation>
); Walker, 1874: 10 (keyed); Howard, 1886: 172 (keyed); Cresson, 1887: 248, 314 (catalog of species of U.S. and Canada); Cresson, 1887: 84 (keyed); Ashmead, 1893: 210, 211, 229 (description, keyed, key to species of U.S. and Canada); Ashmead, 1894: 217, 229 (key to species of St. Vincent, keyed); Ashmead, 1896: 265 (keyed); Dalla Torre, 1898: 498 (catalog of species); Ashmead, 1900: 328 (list of species of West Indies); Ashmead, 1903: 92, 94 (keyed); Kieffer, 1908: 119 (keyed); Brues, 1908: 27, 28, 37, 51 (diagnosis, list of species, keyed); Kieffer, 1910: 65, 81 (description, list of species, keyed); Brues, 1910: 47 (key to species of North America); Kieffer, 1912: 56 (key to species of Seychelles); Dodd, 1913a: 131 (keyed); Kieffer, 1913: 230, 235 (description, key to species of Europe and Algeria); Dodd, 1915: 18 (key to species of Australia, Java, and Fiji); Brues, 1916: 543, 544 (keyed); Kieffer, 1926: 271, 453 (description, keyed, key to species); Nixon, 1934: 1 (description, key to new species described); Nixon, 1934: 290 (description, key to species of Africa); Jansson, 1939: 172 (keyed); Maneval, 1940: 112, 113 (keyed); Mani, 1941: 20, 26 (catalog of species of India, keyed); Risbec, 1950: 585, 591 (key to species of Ethiopian region, keyed); Muesebeck &amp; Walkley, 1951: 704 (catalog of species of U.S. and Canada); Muesebeck &amp; Walkley, 1956: 357 (citation of type species); Masner, 1958: 42 (status of subgenera, delimitation of species groups); Masner, 1961: 158 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday);
<normalizedToken originalValue="Szabó">Szabo</normalizedToken>
, 1966: 421, 429 (description, key to Palearctic species known to the author, keyed); Baltazar, 1966: 182 (cataloged, catalog of species of the Philippines);
<normalizedToken originalValue="Hellén">Hellen</normalizedToken>
, 1971: 5, 22 (description, keyed); Carpenter, 1992: 471 (fossil references).
<emphasis bold="true" pageId="0" pageNumber="323">Comments.</emphasis>
The holotype specimen of
<taxonomicName authorityName="Foerster" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus exsculptus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="exsculptus">Hadronotus exsculptus</taxonomicName>
is missing its head (Figures
<figureCitation captionStart="Figures 9294" captionStartId="F26" captionText="Figures 92 - 94. Hadronotus exsculptus, holotype female (NHMW-HYM # 0002996), mesosoma and metasoma 92 posterodorsal view 93 dorsal view 94 lateral view." figureDoi="10.3897/jhr.87.72842.figures92-94" httpUri="https://binary.pensoft.net/fig/627193" pageId="0" pageNumber="323">92-94</figureCitation>
), but the morphology of the mesosoma and metasoma clearly match the generic concept that we associate with Clade B: T1 without lateral pit (Figure
<figureCitation captionStart="Figures 9294" captionStartId="F26" captionText="Figures 92 - 94. Hadronotus exsculptus, holotype female (NHMW-HYM # 0002996), mesosoma and metasoma 92 posterodorsal view 93 dorsal view 94 lateral view." figureDoi="10.3897/jhr.87.72842.figures92-94" httpUri="https://binary.pensoft.net/fig/627193" pageId="0" pageNumber="323">93</figureCitation>
), hind tibia without subgenual spines (Figure
<figureCitation captionStart="Figures 9294" captionStartId="F26" captionText="Figures 92 - 94. Hadronotus exsculptus, holotype female (NHMW-HYM # 0002996), mesosoma and metasoma 92 posterodorsal view 93 dorsal view 94 lateral view." figureDoi="10.3897/jhr.87.72842.figures92-94" httpUri="https://binary.pensoft.net/fig/627193" pageId="0" pageNumber="323">94</figureCitation>
), metapleuron setose (Figure
<figureCitation captionStart="Figures 9294" captionStartId="F26" captionText="Figures 92 - 94. Hadronotus exsculptus, holotype female (NHMW-HYM # 0002996), mesosoma and metasoma 92 posterodorsal view 93 dorsal view 94 lateral view." figureDoi="10.3897/jhr.87.72842.figures92-94" httpUri="https://binary.pensoft.net/fig/627193" pageId="0" pageNumber="323">94</figureCitation>
). The nearly parallel arrangement of the acetabular carina and mesopleural carina, and transverse shape of foveae in the prespecular sulcus are characters known to us from other species of
<taxonomicName authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Hadronotus</taxonomicName>
and will be useful for treating
<taxonomicName family="Scelionidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="H. exsculptus" order="Hymenoptera" pageId="0" pageNumber="323" rank="species" species="exsculptus">H. exsculptus</taxonomicName>
at the species level in future studies.
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName class="Insecta" family="Scelionidae" genus="Muscidea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Muscidea" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Muscidea</taxonomicName>
Motschoulsky, 1863
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 70 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Muscidea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Muscidea pubescens" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="pubescens">Muscidea pubescens</taxonomicName>
Motschoulsky, by monotypy. Synonymized by
<bibRefCitation DOI="https://doi.org/10.4039/entm10897fv" author="Masner, L" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="323" pagination="1 - 87" refId="B152" refString="Masner, L, 1976. Revisionary notes and keys to world genera of Scelionidae (Hymenoptera: Proctotrupoidea). Memoirs of the Entomological Society of Canada 97: 1 - 87, DOI: https://doi.org/10.4039/entm10897fv" title="Revisionary notes and keys to world genera of Scelionidae (Hymenoptera: Proctotrupoidea)." url="https://doi.org/10.4039/entm10897fv" volume="97" year="1976">Masner (1976)</bibRefCitation>
); Ashmead, 1904a: 326 (keyed); Masner, 1976: 57 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday).
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Dodd" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Hadronotoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hadronotoides" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Hadronotoides</taxonomicName>
Dodd, 1913b
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 171 (original description. Type:
<taxonomicName authorityName="Dodd" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus pentatomus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="pentatomus">Hadronotus pentatomus</taxonomicName>
Dodd, by monotypy and original designation. Treated as junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
by Caleca (1990)); Kieffer, 1926: 266, 474 (description, keyed, key to species); Brues, 1940: 81 (description); Mani, 1941: 19, 27 (catalog of species of India, keyed); Muesebeck &amp; Walkley, 1956: 357 (citation of type species); Masner, 1976: 7, 59 (description, keyed); Mani &amp; Sharma, 1982: 151 (keyed); Mineo &amp; Villa, 1982b: 175 (taxonomic value of pleural structures, clypeus, and antennal sensilla); Mineo &amp; Villa, 1982a: 139 (taxonomic value of structures on the posterior surface of the head); Galloway &amp; Austin, 1984: 6, 81 (diagnosis, list of species described from Australia, keyed); Johnson, 1992: 398 (cataloged, catalog of world species); Carpenter, 1992: 471 (fossil references).
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Dodd" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Platyteleia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Platyteleia" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Platyteleia</taxonomicName>
Dodd, 1913a
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 131, 153 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Platyteleia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Platyteleia latipennis" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="latipennis">Platyteleia latipennis</taxonomicName>
Dodd, by monotypy and original designation); Dodd, 1914b: 79 (description); Kieffer, 1926: 269, 408 (description, keyed, key to species); Muesebeck &amp; Walkley, 1956: 386 (citation of type species); Masner, 1958: 42 (status of subgenera, delimitation of species groups); Masner, 1961: 158 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday);
<normalizedToken originalValue="Szabó">Szabo</normalizedToken>
, 1966: 421, 429 (description, key to Palearctic species known to the author, keyed); Baltazar, 1966: 182 (cataloged, catalog of species of the Philippines);
<normalizedToken originalValue="Hellén">Hellen</normalizedToken>
, 1971: 5, 22 (description, keyed); Galloway &amp; Austin, 1984: 78 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday); Carpenter, 1992: 471 (fossil references).
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Dodd" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Telenomoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telenomoides" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Telenomoides</taxonomicName>
Dodd, 1913a
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 158, 168 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Telenomoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telenomoides flavipes" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="flavipes">Telenomoides flavipes</taxonomicName>
Dodd, by original designation. Key to species of Australia, keyed); Muesebeck &amp; Walkley, 1956: 402 (citation of type species).
<emphasis bold="true" pageId="0" pageNumber="323">Comments.</emphasis>
<bibRefCitation author="Mineo, G" journalOrPublisher="Frustula Entomologica" pageId="0" pageNumber="323" pagination="171 - 188" refId="B178" refString="Mineo, G, 1990a. Studies on the Scelionidae (Hym. Proctotrupoidea) XXV. Material for a revision of Gryon Haliday with description of six new species: Gryon crassifemoratum, G. gryonis, G. minimum, G. pecki, G. scorsonis and G. sulawense. Frustula Entomologica 11: 171 - 188" title="Studies on the Scelionidae (Hym. Proctotrupoidea) XXV. Material for a revision of Gryon Haliday with description of six new species: Gryon crassifemoratum, G. gryonis, G. minimum, G. pecki, G. scorsonis and G. sulawense." volume="11" year="1990 a">Mineo (1990a)</bibRefCitation>
treated
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Telenomoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telenomoides flavipes" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="flavipes">Telenomoides flavipes</taxonomicName>
as a junior synonym of
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon orestes" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="orestes">Gryon orestes</taxonomicName>
(Dodd), implicitly making
<taxonomicName authorityName="Dodd" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Telenomoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telenomoides" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Telenomoides</taxonomicName>
a junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
. Examination of the holotype specimen leads us to treat
<taxonomicName authorityName="Dodd" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Telenomoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telenomoides" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Telenomoides</taxonomicName>
as a junior synonym of
<taxonomicName authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Hadronotus</taxonomicName>
based on the presence of five clavomeres, the shape of the clypeus, and the form of foveae along anterior T1.
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Brethes" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Notilena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Notilena" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Notilena</taxonomicName>
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, 1913
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 84 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Notilena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Notilena Gallardoi" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="Gallardoi">Notilena Gallardoi</taxonomicName>
<normalizedToken originalValue="Brèthes">Brethes</normalizedToken>
, by monotypy and original designation); Muesebeck &amp; Walkley, 1956: 375 (citation of type species); De Santis &amp; Esquivel, 1966: 96 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday).
<emphasis bold="true" pageId="0" pageNumber="323">Comments.</emphasis>
We remove
<taxonomicName authorityName="Brethes" authorityYear="1913" class="Insecta" family="Scelionidae" genus="Notilena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Notilena" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Notilena</taxonomicName>
from
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
and treat it as a synonym of
<taxonomicName authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Hadronotus</taxonomicName>
based on characters in the original description, &quot;Capite punctato-umbilicato, facie longitrorsum impressa, utrinque transverse striata et in medio antennas versus longitrorsum cristata,&quot; which we interpret to indicate that the sculpture of the head is punctate-umbilicate and that the antennal scrobe has transverse striation.
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Dodd" authorityYear="1914" class="Insecta" family="Scelionidae" genus="Austroscelio" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Austroscelio" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Austroscelio</taxonomicName>
Dodd, 1914c
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 93 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Sparasion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sparasion nigricoxa" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="nigricoxa">Sparasion nigricoxa</taxonomicName>
Dodd, by original designation. Synonymized by Galloway, in Galloway &amp; Austin (1984)); Kieffer, 1926: 266, 473 (description, keyed, key to species); Muesebeck &amp; Walkley, 1956: 334 (citation of type species); Galloway &amp; Austin, 1984: 78 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday).
</paragraph>
<paragraph pageId="0" pageNumber="323">
<taxonomicName authorityName="Kieffer" authorityYear="1926" family="Scelionidae" genus="Hadrophanurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hadrophanurus" order="Hymenoptera" pageId="0" pageNumber="323" rank="genus">Hadrophanurus</taxonomicName>
Kieffer, 1926
<emphasis bold="true" pageId="0" pageNumber="323">syn. n.</emphasis>
: 15, 130 (original description. Type:
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Telenomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telenomus pennsylvanicus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="pennsylvanicus">Telenomus pennsylvanicus</taxonomicName>
Ashmead, by monotypy, keyed. Synonymized by
<bibRefCitation author="Masner, L" journalOrPublisher="Casopis Ceskoslovenske Spolecnosti Entomologicke" pageId="0" pageNumber="323" pagination="157 - 168" refId="B149" refString="Masner, L, 1961. The genera Gryon Hal., Idris Foerst. and Hemisius Westw. (Hym., Scelionidae). Casopis Ceskoslovenske Spolecnosti Entomologicke 58: 157 - 168" title="The genera Gryon Hal., Idris Foerst. and Hemisius Westw. (Hym., Scelionidae)." volume="58" year="1961">Masner (1961)</bibRefCitation>
); Muesebeck &amp; Walkley, 1951: 694 (catalog of species of U.S. and Canada); Muesebeck &amp; Walkley, 1956: 357 (citation of type species); Masner, 1961: 158 (junior synonym of
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">Gryon</taxonomicName>
Haliday); Subba Rao &amp; Chacko, 1962: 479 (key to species).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="323" type="diagnosis">
<paragraph pageId="0" pageNumber="323">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="323">Sculpture of head and mesosoma highly variable, ranging from coriaceous microsculpture to coarsely areolate or rugose; mandibular dentition variable, teeth of unequal size; clypeus not projecting; ventral frons without facial striae; antennal scrobe with macrosculpture ranging from transversely striate to areolate rugose; antennal scrobe often delimited by carinae; female antenna with 10 flagellomeres, four to seven clavomeres; sculpture of mesoscutum and mesoscutellum variable, ranging from coriaceous microsculpture to coarsely areolate, striate or rugose; epomial carina variable, sometimes extending dorsally to pronotal shoulder; netrion absent; mesoscutal humeral sulcus and mesoscutal suprahumeral sulcus variable: absent or indicated by a furrow or line of foveae; mesoscutum with or without humeral pit; sculpture of axillula variable, sometimes with parallel carina between coarse foveae, but not distinctly striate; metapleuron divided dorsoventrally by a change in sculpture or setation; hind tibia without subgenual spines; foveae along anterior T1 decreasing in size laterally, not bordered laterally by a carina or pit.</paragraph>
<paragraph pageId="0" pageNumber="323">
<emphasis bold="true" pageId="0" pageNumber="323">Comments.</emphasis>
<taxonomicName authorityName="Foerster" authorityYear="1856" class="Insecta" family="Scelionidae" genus="Hadronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadronotus" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="323">Hadronotus</emphasis>
</taxonomicName>
is morphologically variable and to our knowledge is not united by any single character.
</paragraph>
</subSubSection>
</treatment>
</document>