332 lines
40 KiB
XML
332 lines
40 KiB
XML
<document id="461A869858F0A9A3B16F1224704BC288" ID-DOI="10.5281/zenodo.202649" ID-GBIF-Dataset="7d63c94c-6caa-47ba-aab1-7a201a52b616" ID-ISSN="1175-5326" ID-Zenodo-Dep="202649" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460253797476" checkinUser="plazi" docAuthor="Zaballos, Juan P. & Pérez-González, Sergio" docDate="2011" docId="786487BBFF85FFB8FF01F9E6FE00E57A" docLanguage="en" docName="zt02786p050.pdf" docOrigin="Zootaxa 2786" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Typhlocharis deferreri Zaballos & Perez Gonzalez, new species" docType="treatment" docVersion="10" lastPageNumber="47" masterDocId="845DFFC3FF84FFBDFF96FF98FFC9E74B" masterDocTitle="A new species of Typhlocharis Dieck, 1869 (Coleoptera: Carabidae: Anillini) from South Spain, with notes on the phylogenetic value of sexually related characters and the presence of stridulatory organ (pars stridens) in the genus" masterLastPageNumber="50" masterPageNumber="42" pageNumber="43" updateTime="1698243121341" updateUser="plazi">
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<mods:mods id="807FE075FE679763E51ABFC37E499BE8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BA96107971A588995878FEEC7775BE49">
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<mods:title id="B21CE52C2866B5D7E030026DDEBFB3CD">A new species of Typhlocharis Dieck, 1869 (Coleoptera: Carabidae: Anillini) from South Spain, with notes on the phylogenetic value of sexually related characters and the presence of stridulatory organ (pars stridens) in the genus</mods:title>
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</mods:titleInfo>
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<mods:name id="0D311CB072CB269B951F4CE300DDED46" type="personal">
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<mods:role id="53BEA5D11E9E5D6B4954DEFB090FB92D">
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<mods:roleTerm id="2CCAF65C8F4F01AD48FF632106B7B7E0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4B290820D69D55CEEE566B130439A3CC">Zaballos, Juan P.</mods:namePart>
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</mods:name>
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<mods:name id="C609A6262D0DBAA4B4AC300E17069AA9" type="personal">
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<mods:role id="D34993F124A0FB20042CEA8BBFC6F2E5">
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<mods:roleTerm id="FAACC0D6136D9DBFE4F9C2AC08FE0EDA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E24429C97F0D035525AA8EE091A88D8A">Pérez-González, Sergio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C5AB4423C4FDAD14465B126618A0AEDF">text</mods:typeOfResource>
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<mods:relatedItem id="82EACC3AD7669DF8A6CC31A02CD1BFAF" type="host">
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<mods:titleInfo id="E9279ADBA8F66DEB0B549E6B5D8BE816">
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<mods:title id="9E6FB4FFCDAA4F57898CD2EBA576C4CF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="833A213CE9BEF9F65300CD83D1F9CAA9">
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<mods:date id="F3822EDDC32B280CA156E4E75812913E">2011</mods:date>
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<mods:detail id="A68CC2992D5515703CA937715EFAC88B" type="volume">
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<mods:number id="37B0ED4E91DD39F3B20BAD43C810CB16">2786</mods:number>
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</mods:detail>
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<mods:extent id="93CB89478CBC4B23557740CEA2DC74A3" unit="page">
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<mods:start id="A023DAE7ED116B78979845BDCB3323D3">42</mods:start>
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<mods:end id="70EB39C56F4E4F4E0D34B27666C9930B">50</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="4EAC978AB0AC0A59C32304A396AF8389">journal article</mods:classification>
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<mods:identifier id="9B40522F2F102A6D1F1E356DF89C0B85" type="DOI">10.5281/zenodo.202649</mods:identifier>
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<mods:identifier id="602A1F56A3CDBEBC9ACA566682F2B6C4" type="GBIF-Dataset">7d63c94c-6caa-47ba-aab1-7a201a52b616</mods:identifier>
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<mods:identifier id="F16E06F1AAEDECEC1BD2DC4DDDE38F03" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="993F535F73E7BFB288F651229B269009" type="Zenodo-Dep">202649</mods:identifier>
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</mods:mods>
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<treatment id="786487BBFF85FFB8FF01F9E6FE00E57A" ID-DOI="http://doi.org/10.5281/zenodo.5667869" ID-GBIF-Taxon="119392135" ID-Zenodo-Dep="5667869" LSID="urn:lsid:plazi:treatment:786487BBFF85FFB8FF01F9E6FE00E57A" httpUri="http://treatment.plazi.org/id/786487BBFF85FFB8FF01F9E6FE00E57A" lastPageId="5" lastPageNumber="47" pageId="1" pageNumber="43">
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<subSubSection id="B8D76526FF85FFBCFF01F9E6FEDBE1F1" pageId="1" pageNumber="43" type="nomenclature">
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<paragraph id="F07236ADFF85FFBCFF01F9E6FC51E1D3" blockId="1.[151,920,1662,1722]" box="[151,920,1662,1688]" pageId="1" pageNumber="43">
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<heading id="AB3A81C1FF85FFBCFF01F9E6FC51E1D3" bold="true" box="[151,920,1662,1688]" fontSize="11" level="1" pageId="1" pageNumber="43" reason="1">
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<emphasis id="C2B9EABFFF85FFBCFF01F9E6FC51E1D3" bold="true" box="[151,920,1662,1688]" pageId="1" pageNumber="43">
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||
<taxonomicName id="37CD4D2EFF85FFBCFF01F9E6FCC9E1D3" ID-CoL="59VJL" authority="Zaballos & Perez Gonzalez" authorityName="Zaballos & Perez Gonzalez" box="[151,768,1662,1688]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="43" phylum="Arthropoda" rank="species" species="deferreri" status="sp. nov.">
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<emphasis id="C2B9EABFFF85FFBCFF01F9E6FE6FE1D3" bold="true" box="[151,422,1662,1688]" italics="true" pageId="1" pageNumber="43">Typhlocharis deferreri</emphasis>
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Zaballos & Pérez González
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</taxonomicName>
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<taxonomicNameLabel id="D98A57C4FF85FFBCFC9EF9E6FC51E1D3" box="[776,920,1662,1688]" pageId="1" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F07236ADFF85FFBCFF01F93AFEDBE1F1" blockId="1.[151,920,1662,1722]" box="[151,274,1698,1722]" pageId="1" pageNumber="43">
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(
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<figureCitation id="68F62A28FF85FFBCFF09F93AFEC0E1F1" box="[159,265,1698,1722]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,2000,2023]" captionStartId-1="4.[151,250,836,859]" captionStartId-2="4.[151,250,1805,1828]" captionTargetBox-1="[322,1263,199,802]" captionTargetBox-2="[203,1387,958,1777]" captionTargetId-1="figure@4.[316,1270,193,815]" captionTargetId-2="figure@4.[184,1403,939,1784]" captionTargetPageId-1="4" captionTargetPageId-2="4" captionText-0="FIGURE 1. Habitus 3 of Typhlocharis deferreri nov. sp." captionText-1="FIGURE 2. Abdomen (ventral view) of female and male, showing the median tubercle, the foveae, the chaetotaxy of the last segment and the “ belt ” of scaly microsculpture (shaded in grey)." captionText-2="FIGURE 3. Aedeagus in lateral and dorsal view, parameres in lateral view (a). Female genitalia in ventral view (b)." httpUri-1="https://zenodo.org/record/202650/files/figure.png" httpUri-2="https://zenodo.org/record/202651/files/figure.png" pageId="1" pageNumber="43">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B8D76526FF85FFBCFF01F973FEF5E0D9" pageId="1" pageNumber="43" type="materials_examined">
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<paragraph id="F07236ADFF85FFBCFF01F973FEF5E0D9" blockId="1.[151,1437,1770,2010]" pageId="1" pageNumber="43">
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<emphasis id="C2B9EABFFF85FFBCFF01F973FEEBE048" bold="true" box="[151,290,1770,1795]" pageId="1" pageNumber="43">
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<typeStatus id="2F76880FFF85FFBCFF01F973FF1BE048" box="[151,210,1771,1795]" pageId="1" pageNumber="43">Type</typeStatus>
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series.
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</emphasis>
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<typeStatus id="2F76880FFF85FFBCFEBEF972FE5CE049" box="[296,405,1770,1794]" pageId="1" pageNumber="43" type="holotype">Holotype</typeStatus>
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: 1 3 (nº 10),
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<date id="8473106DFF85FFBCFDBFF972FD64E049" box="[553,685,1770,1794]" pageId="1" pageNumber="43" value="1998-01-27">27-01-1998</date>
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, Cortijo La Mesa–La Nava. Chiclana. Cádiz,
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<quantity id="37359B48FF85FFBCFB3CF972FB2AE049" box="[1194,1251,1770,1794]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="43" unit="m" value="33.0">33 m</quantity>
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(
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<geoCoordinate id="95F9506AFF85FFBCFB67F972FA97E049" box="[1265,1374,1770,1794]" direction="north" orientation="latitude" pageId="1" pageNumber="43" precision="925" value="36.4">N 36º 24’</geoCoordinate>
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<geoCoordinate id="95F9506AFF85FFBCFAF2F973FF77E06D" direction="west" orientation="longitude" pageId="1" pageNumber="43" precision="925" value="-6.05">W 6º 03’</geoCoordinate>
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), J.P.Zaballos leg.
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<typeStatus id="2F76880FFF85FFBCFE19F897FDC9E06C" box="[399,512,1807,1831]" pageId="1" pageNumber="43" type="paratype">Paratypes</typeStatus>
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: 1 3,
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<date id="8473106DFF85FFBCFDD7F896FD71E06D" box="[577,696,1806,1830]" pageId="1" pageNumber="43" value="1997-02-09">9-02-1997</date>
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, same locality, J.P.Zaballos leg; 34 3, 34 ƤƤ,
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<date id="8473106DFF85FFBCFB44F896FA9EE06D" box="[1234,1367,1806,1830]" pageId="1" pageNumber="43" value="1998-01-27">27-01-1998</date>
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, same locality, J.P.Zaballos leg.
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<typeStatus id="2F76880FFF85FFBCFE2EF8AAFDE9E001" box="[440,544,1842,1866]" pageId="1" pageNumber="43" type="holotype">Holotype</typeStatus>
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in Coll. Zaballos
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<collectionCode id="96DCAE68FF85FFBCFD78F8AAFCF8E001" LSID="urn:lsid:biocol.org:col:34854" box="[750,817,1842,1866]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="1" pageNumber="43">UCM</collectionCode>
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,
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<typeStatus id="2F76880FFF85FFBCFCABF8ABFC63E000" box="[829,938,1843,1867]" pageId="1" pageNumber="43" type="paratype">Paratypes</typeStatus>
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in coll. Pérez-González (4 ex.), coll. Wrase (4 ex.), coll Ruíz-Tapiador (4 ex.) and
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<collectionCode id="96DCAE68FF85FFBCFDC0F8CEFD64E025" LSID="urn:lsid:biocol.org:col:35275" box="[598,685,1878,1902]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="1" pageNumber="43">MNCN</collectionCode>
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, Madrid (4 ex.) (ref. number 2107), rest of
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<typeStatus id="2F76880FFF85FFBCFB30F8CFFADAE024" box="[1190,1299,1879,1903]" pageId="1" pageNumber="43" type="paratype">Paratypes</typeStatus>
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in Zaballos
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<collectionCode id="96DCAE68FF85FFBCFF01F8E2FF1EE0D9" LSID="urn:lsid:biocol.org:col:34854" box="[151,215,1914,1938]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="1" pageNumber="43">UCM</collectionCode>
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(52 ex.),
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</paragraph>
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</subSubSection>
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<subSubSection id="B8D76526FF85FFBFFF53F806FE56E7BC" lastPageId="2" lastPageNumber="44" pageId="1" pageNumber="43" type="diagnosis">
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<paragraph id="F07236ADFF85FFBFFF53F806FE56E7BC" blockId="1.[151,1437,1770,2010]" lastBlockId="2.[151,1437,151,2035]" lastPageId="2" lastPageNumber="44" pageId="1" pageNumber="43">
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<emphasis id="C2B9EABFFF85FFBCFF53F806FEF6E0FC" bold="true" box="[197,319,1950,1975]" pageId="1" pageNumber="43">Diagnosis.</emphasis>
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Anophthalmous, small endogean beetle with yellow-brownish microreticulate integument with scattered pubescence. Body parallel and depressed. Elytra without teeth in the posterior margin, and 4+3 setae in the umbilicate series (fig. 1). Legs I and III with a marked internal angle in the femora. Sexually dimorphic. Abdominal sternum II of males with a median tubercle, absent in females which have deep foveae instead (fig. 2). Genitalia as in figure 3.
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</paragraph>
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</subSubSection>
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<subSubSection id="B8D76526FF86FFBFFF53FE9BFE99E1B3" pageId="2" pageNumber="44" type="description">
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<paragraph id="F07236ADFF86FFBFFF53FE9BFED6E572" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
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<emphasis id="C2B9EABFFF86FFBFFF53FE9BFE9FE657" bold="true" box="[197,342,259,284]" pageId="2" pageNumber="44">Description.</emphasis>
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Length of
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<typeStatus id="2F76880FFF86FFBFFE74FE9CFD86E657" box="[482,591,260,284]" pageId="2" pageNumber="44" type="holotype">Holotype</typeStatus>
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||
:
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<quantity id="37359B48FF86FFBFFDC8FE9CFD03E657" box="[606,714,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="44" unit="mm" value="1.35">1.35 mm</quantity>
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. Length of
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<typeStatus id="2F76880FFF86FFBFFCCFFE9DFC00E656" box="[857,969,261,285]" pageId="2" pageNumber="44" type="paratype">paratypes</typeStatus>
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:
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<quantity id="37359B48FF86FFBFFC4FFE9CFB48E657" box="[985,1153,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.41" metricValueMin="1.16" pageId="2" pageNumber="44" unit="mm" value="1.285" valueMax="1.41" valueMin="1.16">1.16–1.41 mm</quantity>
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(males),
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<quantity id="37359B48FF86FFBFFB62FE9CFA54E657" box="[1268,1437,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.48" metricValueMin="1.17" pageId="2" pageNumber="44" unit="mm" value="1.325" valueMax="1.48" valueMin="1.17">1.17–1.48 mm</quantity>
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(females). Head wider (
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<quantity id="37359B48FF86FFBFFE36FEBFFDFEE674" box="[416,567,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="44" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.3 mm</quantity>
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in males,
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<quantity id="37359B48FF86FFBFFD3DFEBFFC87E674" box="[683,846,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="2" pageNumber="44" unit="mm" value="0.28" valueMax="0.32" valueMin="0.24">0.24–0.32 mm</quantity>
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in females) than long (
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<quantity id="37359B48FF86FFBFFBC4FEBFFB20E674" box="[1106,1257,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="2" pageNumber="44" unit="mm" value="0.205" valueMax="0.21" valueMin="0.2">0.2–0.21 mm</quantity>
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in males,
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<quantity id="37359B48FF86FFBFFACBFEBFFF33E628" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.7999999999999998" pageId="2" pageNumber="44" unit="mm" value="0.195" valueMax="0.21" valueMin="0.18">0.18– 0.21 mm</quantity>
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in females), with irregular microsculpture, except in front, clypeus and labrum.
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<emphasis id="C2B9EABFFF86FFBFFBEFFED4FAC7E628" box="[1145,1294,331,355]" italics="true" pageId="2" pageNumber="44">Pars stridens</emphasis>
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present in the posterior region of the head (horizontal ridges near the vertex region). Labrum with a triangle-shaped area and a little bump of thicker cuticle. Mouth parts and antennae without apparent special features. Cephalic chaetotaxy (fig. 1): labrum with three pairs of setae and two minute pairs of setae associated; clypeus with two pairs of setae, those on the side much longer; frontal region with one pair close to the frontal sulcus, two pairs of supraocular setae (anterior and posterior), one supra-antennal pair and one genal pair; two pairs of temporal setae and sparse pubescence.
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</paragraph>
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<paragraph id="F07236ADFF86FFBFFF53FDDEFC7DE45B" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
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Pronotum slightly longer (
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<quantity id="37359B48FF86FFBFFE7CFDDDFD46E516" box="[490,655,581,605]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" pageId="2" pageNumber="44" unit="mm" value="0.36" valueMax="0.41" valueMin="0.31">0.31–0.41 mm</quantity>
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||
in males,
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||
<quantity id="37359B48FF86FFBFFC96FDDDFC5CE516" box="[768,917,581,605]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.2" metricValueMin="3.0" pageId="2" pageNumber="44" unit="mm" value="0.36" valueMax="0.42" valueMin="0.3">0.3–0.42 mm</quantity>
|
||
in females) than wide (
|
||
<quantity id="37359B48FF86FFBFFB0CFDDDFAF8E516" box="[1178,1329,581,605]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.5999999999999996" metricValueMin="3.0" pageId="2" pageNumber="44" unit="mm" value="0.32999999999999996" valueMax="0.36" valueMin="0.3">0.3–0.36 mm</quantity>
|
||
in males,
|
||
<quantity id="37359B48FF86FFBFFF01FDF1FEE8E5CA" box="[151,289,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="44" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
|
||
in females), subquadrate, posterior angles with 2–3 minor rounded denticles. Surface covered with subhexagonal to irregular microsculpture. There is one medial sulcus and one pair of lateral sulci. Chaetotaxy (fig. 1): one pair of setae in the anterior quarter of the pronotum, one pair in the posterior angles, five or six pairs in the anterior margin, and two pairs in the posterior margin. Pubescence in four or five rows on the disc. Margins with a row of pubescence, a little denser in the anterior and posterior margins.
|
||
</paragraph>
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<paragraph id="F07236ADFF86FFBFFF53FC84FAF9E31A" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
|
||
Elytra approximately twice as long (
|
||
<quantity id="37359B48FF86FFBFFDFEFC84FCC6E47F" box="[616,783,796,820]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="2" pageNumber="44" unit="mm" value="0.725" valueMax="0.8" valueMin="0.65">0.65–0.80 mm</quantity>
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||
in males,
|
||
<quantity id="37359B48FF86FFBFFC1FFC84FBE6E47F" box="[905,1071,796,820]" metricMagnitude="-4" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="2" pageNumber="44" unit="mm" value="0.765" valueMax="0.85" valueMin="0.68">0.68–0.85 mm</quantity>
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||
in females) as wide (
|
||
<quantity id="37359B48FF86FFBFFABCFC84FF0AE413" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.7" metricValueMin="3.1" pageId="2" pageNumber="44" unit="mm" value="0.33999999999999997" valueMax="0.37" valueMin="0.31">0.31–0.37 mm</quantity>
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||
in males,
|
||
<quantity id="37359B48FF86FFBFFEACFCD8FE17E413" box="[314,478,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="4.2" metricValueMin="3.1" pageId="2" pageNumber="44" unit="mm" value="0.365" valueMax="0.42" valueMin="0.31">0.31–0.42 mm</quantity>
|
||
in females), sides parallel, apex rounded. Lateral margins serrate, with 19–22 small teeth, decreasing in size posteriorly. Apical margin without teeth, but with the sutural angle slightly asymmetric. Surface covered with subhexagonal to irregular microsculpture. Scutellar organ present near the base of the suture. Disc flattened and with lateral carinae in the seventh stria. Chaetotaxy: umbilicate series with four setae in the anterior group and three in the posterior group (4+3). The distribution of the setae of the anterior group is characteristic: the third seta is separated by twice the distance between setae 1º and 2º and the 4º seta is separated twice the distance between 2º and 3º (fig. 1). Pubescence in five rows, lateral margins with one small seta in each denticle and apical margin with a row of small setae. There are two scutellary setae, but no discal or apical long setae.
|
||
</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53FBC6FC00E3D2" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">Legs similar in both sexes. Profemora and metafemora robust, the inner margin with a marked internal angle. The metatrochanters are rounded. There are no mesotibial hairs in males.</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53FB3DFEB3E207" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">Abdomen covered by irregular and large sized microsculpture, except the last segment which has a “belt” of scaly and serrate microsculpture (fig. 2). Females with a deep foveae in the sternum II and very slightly foveae in sterna III and IV. Males typically lack this foveae (if present, it is extremely superficial), but have a small, pointed median tubercle in the sternum II, with two small setae near the basis. Chaetotaxy of the last segment shows sexual dimorphism (fig. 2).</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53FAC0FE5EE2FC" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
|
||
Aedeagus (fig. 3a) with sickle-shaped median lobe, short and robust (
|
||
<quantity id="37359B48FF86FFBFFC41FAC0FB8BE23B" box="[983,1090,1368,1392]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="44" unit="mm" value="0.22">0.22 mm</quantity>
|
||
), arcuate, with a screw-shaped sclerite in the internal sac. In dorsal view the apex is bent to the left. Both parameres are subtriangular with two short setae in the apex.
|
||
</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53FA5BFBA2E10D" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">Female genitalia (fig. 3b) with long, tubular gonocoxites, with two fused apical setae in each one and scarce sensilae with some degree of individual variation. The gonoduct is short and shows the same thickness along its length, except at the connection with the spermatheca, which is funnel-shaped. The spermatheca is peanut-shaped in lateral view and subspherical in dorsal view. The spermathecal gland is cone-shaped.</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53F9C9FE99E1B3" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
|
||
<emphasis id="C2B9EABFFF86FFBFFF53F9C9FEE1E121" bold="true" box="[197,296,1617,1642]" pageId="2" pageNumber="44">Habitat.</emphasis>
|
||
The majority of the
|
||
<typeStatus id="2F76880FFF86FFBFFD98F9CAFDF7E121" box="[526,574,1618,1642]" pageId="2" pageNumber="44">type</typeStatus>
|
||
specimens were captured on a stormy day during
|
||
<date id="8473106DFF86FFBFFBF8F9CAFACDE122" box="[1134,1284,1617,1642]" pageId="2" pageNumber="44" value="1998-01">January 1998</date>
|
||
under deeply buried stones in open land with mainly clay soils (with little sand content), wet, but not saturated. The vegetation was mainly represented by fan palms (
|
||
<taxonomicName id="37CD4D2EFF86FFBFFDF4F901FC87E1FA" box="[610,846,1689,1713]" class="Liliopsida" family="Arecaceae" genus="Chamaerops" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="44" phylum="Tracheophyta" rank="species" species="humilis">
|
||
<emphasis id="C2B9EABFFF86FFBFFDF4F901FC87E1FA" box="[610,846,1689,1713]" italics="true" pageId="2" pageNumber="44">Chamaerops humilis</emphasis>
|
||
</taxonomicName>
|
||
L.), mastic trees (
|
||
<taxonomicName id="37CD4D2EFF86FFBFFBBEF902FB38E1FA" box="[1064,1265,1689,1713]" class="Magnoliopsida" family="Anacardiaceae" genus="Pistacia" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="44" phylum="Tracheophyta" rank="species" species="lentiscus">
|
||
<emphasis id="C2B9EABFFF86FFBFFBBEF902FB38E1FA" box="[1064,1265,1689,1713]" italics="true" pageId="2" pageNumber="44">Pistacia lentiscus</emphasis>
|
||
</taxonomicName>
|
||
L.) and white asphodel (
|
||
<taxonomicName id="37CD4D2EFF86FFBFFE9DF925FDEDE19E" authority="Miller" authorityName="Miller" box="[267,548,1725,1749]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Asphodelus" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="44" phylum="Tracheophyta" rank="species" species="albus">
|
||
<emphasis id="C2B9EABFFF86FFBFFE9DF925FE18E19E" box="[267,465,1725,1749]" italics="true" pageId="2" pageNumber="44">Asphodelus albus</emphasis>
|
||
Miller
|
||
</taxonomicName>
|
||
). In this habitat also lives another Anillini,
|
||
<taxonomicName id="37CD4D2EFF86FFBFFB84F925FE85E1B3" authority="Ganglbauer, 1900 (Zaballos 2005)" authorityName="Ganglbauer, 1900 (Zaballos" authorityYear="2005" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="korbi">
|
||
<emphasis id="C2B9EABFFF86FFBFFB84F925FB02E19E" box="[1042,1227,1725,1749]" italics="true" pageId="2" pageNumber="44">Geocharis korbi</emphasis>
|
||
Ganglbauer, 1900 (
|
||
<bibRefCitation id="945C4B5CFF86FFBFFF08F978FE8CE1B3" author="Zaballos" box="[158,325,1760,1784]" pageId="2" pageNumber="44" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">Zaballos 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8D76526FF86FFB8FF53F89DFE00E57A" lastPageId="5" lastPageNumber="47" pageId="2" pageNumber="44" type="etymology">
|
||
<paragraph id="F07236ADFF86FFBFFF53F89DFE67E00B" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
|
||
<emphasis id="C2B9EABFFF86FFBFFF53F89DFE82E056" bold="true" box="[197,331,1797,1821]" pageId="2" pageNumber="44">Etymology.</emphasis>
|
||
The new species is named
|
||
<emphasis id="C2B9EABFFF86FFBFFD0FF89EFCFBE056" box="[665,818,1798,1821]" italics="true" pageId="2" pageNumber="44">in memoriam</emphasis>
|
||
of Juan de Ferrer Andreu, great entomologist, great friend, a truly great man.
|
||
</paragraph>
|
||
<paragraph id="F07236ADFF86FFBFFF53F8D3FC1BE0B8" blockId="2.[151,1437,151,2035]" pageId="2" pageNumber="44">
|
||
<emphasis id="C2B9EABFFF86FFBFFF53F8D3FEF2E02F" bold="true" box="[197,315,1867,1892]" pageId="2" pageNumber="44">Affinities.</emphasis>
|
||
This new species can be assigned to the
|
||
<emphasis id="C2B9EABFFF86FFBFFCB7F8D4FCADE02F" box="[801,868,1868,1892]" italics="true" pageId="2" pageNumber="44">diecki</emphasis>
|
||
group (
|
||
<emphasis id="C2B9EABFFF86FFBFFC55F8D5FBC8E02F" box="[963,1025,1869,1892]" italics="true" pageId="2" pageNumber="44">sensu</emphasis>
|
||
<bibRefCitation id="945C4B5CFF86FFBFFB9DF8D4FA59E02F" author="Zaballos" box="[1035,1424,1868,1892]" pageId="2" pageNumber="44" refString="Zaballos, J. P. & Ruiz-Tapiador, I. (1997) Nuevos Typhlocharis Dieck (Coleoptera: Caraboidea: Trechidae) de Espana. Graellsia, 52, 95 - 106." type="journal article" year="1997">Zaballos and Ruíz-Tapiador 1997</bibRefCitation>
|
||
), because it shows the umbilicate series pattern (4+3); a group that currently includes a further 9 species (
|
||
<bibRefCitation id="945C4B5CFF86FFBFFAB5F8E8FF30E0E0" author="Andujar" pageId="2" pageNumber="44" refString="Andujar, C., Lencina, J. L. & Serrano, J. (2008) Typhlocharis Dieck, 1869 (Coleoptera: Carabidae: Anillina) a new species from the Iberian Peninsula, with notes about its relationships and the evolution of the diecki species group. Zootaxa, 1842, 35 - 44." type="journal article" year="2008">
|
||
Andújar
|
||
<emphasis id="C2B9EABFFF86FFBFFA1EF8E9FF64E0E0" italics="true" pageId="2" pageNumber="44">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="37CD4D2EFF86FFBFFE9AF80BFDC0E0E0" box="[268,521,1939,1963]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF86FFBFFE9AF80BFDC0E0E0" box="[268,521,1939,1963]" italics="true" pageId="2" pageNumber="44">Typhlocharis deferreri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C2B9EABFFF86FFBFFD84F80CFDA5E0E7" bold="true" box="[530,620,1940,1964]" pageId="2" pageNumber="44">nov. sp.</emphasis>
|
||
can be separated from all the other species of the group by having secondary sexual dimorphism on the ventral sternites: males have a median tubercle in the posterior margin of sternum II and females have quite deep posterolateral foveae in sternum II (fig. 2).
|
||
</paragraph>
|
||
<caption id="A4B26625FF87FFBEFF01F848FD14E0AD" box="[151,733,2000,2023]" pageId="3" pageNumber="45">
|
||
<paragraph id="F07236ADFF87FFBEFF01F848FD14E0AD" blockId="3.[151,733,2000,2023]" box="[151,733,2000,2023]" pageId="3" pageNumber="45">
|
||
<emphasis id="C2B9EABFFF87FFBEFF01F848FEDAE0AE" bold="true" box="[151,275,2000,2023]" pageId="3" pageNumber="45">FIGURE 1.</emphasis>
|
||
Habitus 3 of
|
||
<taxonomicName id="37CD4D2EFF87FFBEFE35F848FD4FE0AD" box="[419,646,2000,2022]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="45" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF87FFBEFE35F848FD4FE0AD" box="[419,646,2000,2022]" italics="true" pageId="3" pageNumber="45">Typhlocharis deferreri</emphasis>
|
||
</taxonomicName>
|
||
nov. sp.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A4B26625FF80FFB9FF01FCDCFCEDE432" httpUri="https://zenodo.org/record/202650/files/figure.png" pageId="4" pageNumber="46" targetBox="[322,1263,199,802]" targetPageId="4">
|
||
<paragraph id="F07236ADFF80FFB9FF01FCDCFCEDE432" blockId="4.[151,1436,836,889]" pageId="4" pageNumber="46">
|
||
<emphasis id="C2B9EABFFF80FFB9FF01FCDCFEDAE411" bold="true" box="[151,275,836,859]" pageId="4" pageNumber="46">FIGURE 2.</emphasis>
|
||
Abdomen (ventral view) of female and male, showing the median tubercle, the foveae, the chaetotaxy of the last segment and the “belt” of scaly microsculpture (shaded in grey).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A4B26625FF80FFB9FF01F895FAE4E068" box="[151,1325,1805,1828]" httpUri="https://zenodo.org/record/202651/files/figure.png" pageId="4" pageNumber="46" targetBox="[203,1387,958,1777]" targetPageId="4">
|
||
<paragraph id="F07236ADFF80FFB9FF01F895FAE4E068" blockId="4.[151,1325,1805,1828]" box="[151,1325,1805,1828]" pageId="4" pageNumber="46">
|
||
<emphasis id="C2B9EABFFF80FFB9FF01F895FEDBE069" bold="true" box="[151,274,1805,1828]" pageId="4" pageNumber="46">FIGURE 3.</emphasis>
|
||
Aedeagus in lateral and dorsal view, parameres in lateral view (a). Female genitalia in ventral view (b).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F07236ADFF80FFB8FF53F8D2FA42E799" blockId="4.[151,1436,1866,2031]" lastBlockId="5.[151,1436,151,561]" lastPageId="5" lastPageNumber="47" pageId="4" pageNumber="46">
|
||
This combination of traits (sexual dimorphism with presence of a median tubercle in males and 4+3 umbilicate series) is present also in other species of the genus that are included in different species groups:
|
||
<taxonomicName id="37CD4D2EFF80FFB9FB4BF8F7FE90E0E3" authority="Serrano and Aguiar, 2006" authorityName="Serrano and Aguiar" authorityYear="2006" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="bivari">
|
||
<emphasis id="C2B9EABFFF80FFB9FB4BF8F7FAF2E0CD" box="[1245,1339,1902,1926]" italics="true" pageId="4" pageNumber="46">T. bivari</emphasis>
|
||
Serrano and Aguiar, 2006
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37CD4D2EFF80FFB9FE06F80AFCEFE0E3" authority="Serrano and Aguiar, 2005" authorityName="Serrano and Aguiar" authorityYear="2005" box="[400,806,1936,1961]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="passosi">
|
||
<emphasis id="C2B9EABFFF80FFB9FE06F80AFE37E0E2" box="[400,510,1938,1961]" italics="true" pageId="4" pageNumber="46">T. passosi</emphasis>
|
||
<bibRefCitation id="945C4B5CFF80FFB9FD93F808FCEFE0E3" author="Serrano" box="[517,806,1936,1960]" pageId="4" pageNumber="46" refString="Serrano, A. R. M., Aguiar, C. A. & Proenca, S. J. (2005) Two new species of Typhlocharis Dieck of the gomezi species group from Portugal (Coleoptera: Carabidae). The Coleopterist Bulletin, 59, 239 - 249." type="journal article" year="2005">Serrano and Aguiar, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="C2B9EABFFF80FFB9FCA1F80AFC4FE0E2" box="[823,902,1938,1961]" italics="true" pageId="4" pageNumber="46">gomezi</emphasis>
|
||
group) differ by the presence of a median teeth on clypeus, two teeth in tibiae II and III of males and two teeth in the posterior margin of elytra (absent in the new species). On the other hand,
|
||
<taxonomicName id="37CD4D2EFF80FFB9FE77F840FC5CE0A5" authority="Zaballos and Wrase, 1998" authorityName="Zaballos and Wrase" authorityYear="1998" box="[481,917,2006,2031]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="navarica">
|
||
<emphasis id="C2B9EABFFF80FFB9FE77F840FDABE0A5" box="[481,610,2007,2031]" italics="true" pageId="4" pageNumber="46">T. navarica</emphasis>
|
||
<bibRefCitation id="945C4B5CFF80FFB9FDFDF84EFC5CE0A5" author="Zaballos" box="[619,917,2006,2030]" pageId="4" pageNumber="46" refString="Zaballos, J. P. & Wrase, D. W. (1998) Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia, 54, 43 - 52." type="journal article" year="1998">Zaballos and Wrase, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="37CD4D2EFF80FFB9FC3EF84EFBC5E0A5" box="[936,1036,2006,2030]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="outereloi">
|
||
<emphasis id="C2B9EABFFF80FFB9FC3EF84EFBC5E0A5" box="[936,1036,2006,2030]" italics="true" pageId="4" pageNumber="46">outereloi</emphasis>
|
||
</taxonomicName>
|
||
group),
|
||
<taxonomicName id="37CD4D2EFF80FFB8FBE6F840FEECE7E4" authority="Zaballos and Wrase, 1998" authorityName="Zaballos and Wrase" authorityYear="1998" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" lastPageId="5" lastPageNumber="47" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="monastica">
|
||
<emphasis id="C2B9EABFFF80FFB9FBE6F840FAC9E0A5" box="[1136,1280,2007,2031]" italics="true" pageId="4" pageNumber="46">T. monastica</emphasis>
|
||
<bibRefCitation id="945C4B5CFF80FFB8FA9CF84EFEECE7E4" author="Zaballos" lastPageId="5" lastPageNumber="47" pageId="4" pageNumber="46" refString="Zaballos, J. P. & Wrase, D. W. (1998) Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia, 54, 43 - 52." type="journal article" year="1998">Zaballos and Wrase, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37CD4D2EFF81FFB8FEC8FF01FCD0E7E4" authority="Zaballos and Wrase, 1998" authorityName="Zaballos and Wrase" authorityYear="1998" box="[350,793,151,176]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="peregrina">
|
||
<emphasis id="C2B9EABFFF81FFB8FEC8FF01FE21E7FB" box="[350,488,153,176]" italics="true" pageId="5" pageNumber="47">T. peregrina</emphasis>
|
||
<bibRefCitation id="945C4B5CFF81FFB8FE67FF0FFCD0E7E4" author="Zaballos" box="[497,793,151,175]" pageId="5" pageNumber="47" refString="Zaballos, J. P. & Wrase, D. W. (1998) Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia, 54, 43 - 52." type="journal article" year="1998">Zaballos and Wrase, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="37CD4D2EFF81FFB8FCBCFF01FC54E7FB" box="[810,925,153,176]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="monastica">
|
||
<emphasis id="C2B9EABFFF81FFB8FCBCFF01FC54E7FB" box="[810,925,153,176]" italics="true" pageId="5" pageNumber="47">monastica</emphasis>
|
||
</taxonomicName>
|
||
group) have a median tubercle in males, but they have 4+2 umbilicate series and teeth in the posterior margin of elytra (4+3 and unarmed in the new species).
|
||
</paragraph>
|
||
<paragraph id="F07236ADFF81FFB8FF53FF46FCF8E670" blockId="5.[151,1436,151,561]" pageId="5" pageNumber="47">
|
||
The angle of the inner margin of the profemora relates
|
||
<taxonomicName id="37CD4D2EFF81FFB8FCA6FF47FC65E7BD" box="[816,940,222,246]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF81FFB8FCA6FF47FC65E7BD" box="[816,940,222,246]" italics="true" pageId="5" pageNumber="47">T. deferreri</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="37CD4D2EFF81FFB8FC78FF47FB90E7BD" box="[1006,1113,223,246]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="armata">
|
||
<emphasis id="C2B9EABFFF81FFB8FC78FF47FB90E7BD" box="[1006,1113,223,246]" italics="true" pageId="5" pageNumber="47">T. armata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="37CD4D2EFF81FFB8FBFCFF47FACAE7BD" box="[1130,1283,222,246]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="silvanoides">
|
||
<emphasis id="C2B9EABFFF81FFB8FBFCFF47FACAE7BD" box="[1130,1283,222,246]" italics="true" pageId="5" pageNumber="47">T. silvanoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37CD4D2EFF81FFB8FAADFF47FF12E652" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="algarvensis">
|
||
<emphasis id="C2B9EABFFF81FFB8FAADFF47FF12E652" italics="true" pageId="5" pageNumber="47">T. algarvensis</emphasis>
|
||
</taxonomicName>
|
||
; which are geographically close. However,
|
||
<taxonomicName id="37CD4D2EFF81FFB8FD5AFE9AFC80E652" box="[716,841,257,281]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF81FFB8FD5AFE9AFC80E652" box="[716,841,257,281]" italics="true" pageId="5" pageNumber="47">T. deferreri</emphasis>
|
||
</taxonomicName>
|
||
is different from any of these by the combination of traits already commented, specially the sexual dimorphism.
|
||
</paragraph>
|
||
<paragraph id="F07236ADFF81FFB8FF53FEDEFE00E57A" blockId="5.[151,1436,151,561]" pageId="5" pageNumber="47">
|
||
The genitalia characters also support a close relationship of
|
||
<taxonomicName id="37CD4D2EFF81FFB8FCE3FED0FC3DE615" box="[885,1012,326,351]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF81FFB8FCE3FED0FC3DE615" box="[885,1012,326,351]" italics="true" pageId="5" pageNumber="47">T. deferreri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="37CD4D2EFF81FFB8FB93FED0FBBBE614" box="[1029,1138,328,351]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="armata">
|
||
<emphasis id="C2B9EABFFF81FFB8FB93FED0FBBBE614" box="[1029,1138,328,351]" italics="true" pageId="5" pageNumber="47">T. armata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37CD4D2EFF81FFB8FB38FED0FA80E615" box="[1198,1353,326,351]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="silvanoides">
|
||
<emphasis id="C2B9EABFFF81FFB8FB38FED0FA80E615" box="[1198,1353,326,351]" italics="true" pageId="5" pageNumber="47">T. silvanoides</emphasis>
|
||
</taxonomicName>
|
||
. These species share similar shape of gonocoxites, spermatheca and spermathecal gland, but in
|
||
<taxonomicName id="37CD4D2EFF81FFB8FB1DFEF2FAC3E6CA" box="[1163,1290,361,385]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="deferreri">
|
||
<emphasis id="C2B9EABFFF81FFB8FB1DFEF2FAC3E6CA" box="[1163,1290,361,385]" italics="true" pageId="5" pageNumber="47">T. deferreri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C2B9EABFFF81FFB8FA83FEF2FAB9E6C9" bold="true" box="[1301,1392,362,386]" pageId="5" pageNumber="47">nov. sp.</emphasis>
|
||
the gonoduct is a tube with regular thickness, while in the other two species the gonoduct has two well differentiated regions of different thickness. The aedeagus is very similar too, being shorter and more robust in the new species. Finally it is interesting to note that the distribution pattern of the anterior group of setae from the umbilicate series (fig. 1) is very similar to that of
|
||
<taxonomicName id="37CD4D2EFF81FFB8FD94FE6EFBC4E546" authority="Zaballos and Ruiz-Tapiador 1995" authorityName="Zaballos and Ruiz-Tapiador" authorityYear="1995" box="[514,1037,501,525]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="carmenae">
|
||
<emphasis id="C2B9EABFFF81FFB8FD94FE6EFD45E546" box="[514,652,502,525]" italics="true" pageId="5" pageNumber="47">T. carmenae</emphasis>
|
||
Zaballos and Ruíz-Tapiador 1995
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37CD4D2EFF81FFB8FBD2FE6EFA51E546" authority="Andujar et al. 2008" authorityName="Andujar et al." authorityYear="2008" box="[1092,1432,501,525]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="martini">
|
||
<emphasis id="C2B9EABFFF81FFB8FBD2FE6EFB7BE546" box="[1092,1202,502,525]" italics="true" pageId="5" pageNumber="47">T. martini</emphasis>
|
||
<bibRefCitation id="945C4B5CFF81FFB8FB2CFE6DFA51E546" author="Andujar" box="[1210,1432,501,525]" pageId="5" pageNumber="47" refString="Andujar, C., Lencina, J. L. & Serrano, J. (2008) Typhlocharis Dieck, 1869 (Coleoptera: Carabidae: Anillina) a new species from the Iberian Peninsula, with notes about its relationships and the evolution of the diecki species group. Zootaxa, 1842, 35 - 44." type="journal article" year="2008">
|
||
Andújar
|
||
<emphasis id="C2B9EABFFF81FFB8FA8BFE6EFA87E546" box="[1309,1358,501,525]" italics="true" pageId="5" pageNumber="47">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, both from the
|
||
<emphasis id="C2B9EABFFF81FFB8FEA0FD80FEB0E57B" box="[310,377,536,560]" italics="true" pageId="5" pageNumber="47">diecki</emphasis>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |