274 lines
28 KiB
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274 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="41536018B75F2CD16AF90D3E23C37A17" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole manantenensis Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="2A00A2C0-FA00-48C1-AEFB-847A94ABE490" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777007" LSID="urn:lsid:zoobank.org:act:2A00A2C0-FA00-48C1-AEFB-847A94ABE490" httpUri="http://treatment.plazi.org/id/41536018B75F2CD16AF90D3E23C37A17" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/2A00A2C0-FA00-48C1-AEFB-847A94ABE490" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole manantenensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manantenensis" status="sp. nov.">Pheidole manantenensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">Figs 37A-F</figureCitation>
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85F</figureCitation>
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<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87J</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 12 Dec 2005; Fisher et al. leg.; BLF13478, CASENT0068456 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •1w., 1q.; same data as for holotype; CASENT0068457, CASENT0923218 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •2w.; Makirovana Forest;
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<geoCoordinate degrees="14.104" direction="south" orientation="latitude" precision="55" value="-14.104">-14.104</geoCoordinate>
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,
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<geoCoordinate degrees="50.03574" direction="east" orientation="longitude" precision="1" value="50.03574">50.03574</geoCoordinate>
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; alt. 225 m; 4 May 2011; B.L. Fisher et al. leg.; CASENT0230806, CASENT0230809 (CASC). •1w.; Makirovana Forest;
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<geoCoordinate degrees="14.16044" direction="south" orientation="latitude" precision="1" value="-14.16044">-14.16044</geoCoordinate>
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,
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<geoCoordinate degrees="49.95216" direction="east" orientation="longitude" precision="1" value="49.95216">49.95216</geoCoordinate>
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; alt. 550 m; 1 May 2011; B.L. Fisher et al. leg.; CASENT0231430 (CASC). •4w.; Makirovana Forest; -14.16666, 49.95; alt. 715 m; 1 May 2011; B.L. Fisher et al. leg.; CASENT0243103, CASENT0243118, CASENT0243120, CASENT0243129 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1w., 1s.; Montagne
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<normalizedToken originalValue="d’Akirindro">d'Akirindro</normalizedToken>
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7.6 km 341°NNW Ambinanitelo;
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<geoCoordinate degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
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,
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<geoCoordinate degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
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; alt. 600 m; 17 Mar 2003; B.L. Fisher et al. leg.; CASENT0038921, CASENT0039038 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Head in full-face view cordate, widened posteriorly; sides of the head with moderately dense, short, suberect to erect pilosity; antennal scrobes present, weakly impressed, and indistinctly delimited ventrally and posteriorly by carinulae; scrobe surface foveolate, with indistinct, thick, moderately sparse, irregular rugae; propodeal spines long (PSL: 0.2-0.26 (0.24)); gaster shagreened; inner hypostomal teeth distinct, high, narrow, and triangular, closely spaced, with rounded apex; outer hypostomal teeth approximately as high as inner hypostomal teeth, but thinner and with wider base, lobe-like.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head and mesosoma foveolate; propodeal spines long (PSL: 0.14-0.17 (0.15)); promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 0.96-1.13 (1.07); HW: 0.95-1.15 (1.07); SL: 0.52-0.58 (0.55); EL: 0.1-0.12 (0.12); WL: 0.79-0.94 (0.88); PSL: 0.2-0.26 (0.24); MTL: 0.49-0.57 (0.52); PNW: 0.45-0.54 (0.51); PTW: 0.12-0.15 (0.14); PPW: 0.33-0.44 (0.4); CI: 98.2-102.5 (100.5); SI: 48.3-54.7 (51.1); PSLI: 20.0-24.5 (22.1); PPI: 31.9-38.2 (34.8); PNI: 45.6-48.9 (47.1); MTI: 45.6-51.3 (48.2).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view cordate, slightly longer than wide, anterior of eyes slightly convex, posterior of eyes convex, occipital margins of lobes convex (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with moderately dense, short, suberect to erect pilosity; whole head with moderately dense, long, suberect to erect pilosity. Antennal scrobes impressed and delimited ventrally and posteriorly by carinulae; scrobe surface foveolate, with indistinct, thick, moderately sparse, irregular rugae. Occipital lobes shiny, foveolate, with additional thick, sparse, irregular rugae, rugae weakening posteriorly; frons with moderately sparse, thick, longitudinal rugae, on the posterior part rugae longitudinal to irregular, interspaces superficially foveolate; genae foveolate, with additional, indistinct, and irregular rugulae; malar area with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose. Centre of clypeus shiny and with indistinct, short rugulae, lateral sides with longitudinal rugae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37B, D</figureCitation>
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). Inner hypostomal teeth distinct, high, narrow, and triangular, closely spaced, with rounded apex; outer hypostomal teeth approximately as high as inner hypostomal teeth, but thinner and with wider base, lobe-like (Fig.
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum low and arched, dorsal mesonotum slightly concave, posterior mesonotum relatively steep, with small tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines long, triangular, thin, with sharp apex and relatively narrow base; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D</figureCitation>
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). Surface foveolate, with additional sparse and indistinct rugoreticulation; dorsoventral propodeum with fading sculpture. Pilosity sparse, long, erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle moderately long, foveolate, with distinct, short horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny and foveolate; in dorsal view sides with moderately long, wide, acute, and triangular projections; pilosity moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny and shagreened; pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D, F</figureCitation>
|
||
).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Brown to dark brown; legs yellow; malar area brighter that the rest of the head (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole manantenensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manantenensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole manantenensis</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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||
) of paratype minor worker (CASENT0923218) and full-face view (
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||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0068456).
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</paragraph>
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</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.47-0.51 (0.49); HW: 0.41-0.46 (0.44); SL: 0.45-0.54 (0.49); EL: 0.08-0.11 (0.09); WL: 0.58-0.64 (0.62); PSL: 0.14-0.17 (0.15); MTL: 0.35-0.38 (0.37); PNW: 0.28-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.1-0.13 (0.11); CI: 85.2-93.0 (88.9); SI: 105.1-119.7 (111.8); PSLI: 29.2-34.3 (31.5); PPI: 60.0-73.2 (65.8); PNI: 67.5-74.1 (70.7); MTI: 79.8-88.6 (84.0).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin straight or indistinctly concave; occipital carina indistinct (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37A</figureCitation>
|
||
). Pilosity moderately dense, long, suberect. Head foveolate; genae with fading sculpture. Clypeus foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove present; propodeal spines long, triangular, with acute apex and narrow base (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C</figureCitation>
|
||
). Sculpture foveolate. Pilosity moderately sparse, short, and erect (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle moderately short and thin; with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Short, low, and convex; with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dark yellow (Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole manantenensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923218) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0068456)." figureDoi="10.3897/zookeys.905.39592.figure37" httpUri="https://binary.pensoft.net/fig/373084" pageId="0" pageNumber="1">37C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The species was collected between 225-775 m in elevation, in rainforest. Nest was located in the petiole of
|
||
<taxonomicName class="Insecta" family="Melastomataceae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="1" rank="family">Melastomataceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole manantenensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manantenensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole manantenensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="P. vohemarensis" pageId="0" pageNumber="1" rank="species" species="vohemarensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. vohemarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Majors workers.</emphasis>
|
||
It differs from
|
||
<taxonomicName lsidName="P. vohemarensis" pageId="0" pageNumber="1" rank="species" species="vohemarensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. vohemarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in presence of irregular rugae on antennal scrobes, frons with foveolae, lower promesonotum, and petiolar peduncle with short horizontal lobes on its basal part.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
It differs from
|
||
<taxonomicName lsidName="P. vohemarensis" pageId="0" pageNumber="1" rank="species" species="vohemarensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. vohemarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in promesonotum with posterior declivity smoothly declining towards propodeum, and foveolate katepisternum and mesonotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |