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<document id="CB64DF735530634C16F5FC893F79776C" ID-DOI="10.1080/00222930802002535" ID-ISSN="1464-5262" ID-Zenodo-Dep="5219397" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629340601867" checkinUser="felipe" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Scholtz, Clarke H." docDate="2008" docId="1F77BC72FFC23062201CFE77FE04FAA1" docLanguage="en" docName="JNatHist.42.21-24.1477-1508.pdf" docOrigin="Journal of Natural History 42 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222930802002535" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Coptorhina angolensis Arrow" docType="treatment" docVersion="3" lastPageNumber="1498" masterDocId="E34EC40AFFD430752211FF81FFF5FF8C" masterDocTitle="Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1508" masterPageNumber="1477" pageNumber="1497" updateTime="1699068592934" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="916D8ACF68A9F50375A643AE3E3A939F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="88DCA81ACB133042F5D17CE8D755BA51">
<mods:title id="58219489D17A91F5B2BA89781FD4C74D">Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="441069F91CA797A6B978564139F2911B" type="personal">
<mods:role id="5F5EBAF1C5EEFD8302044E3D03A9701A">
<mods:roleTerm id="206CD39CD4ED659C6D1D1B1EF170F13C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E539CCC29403122EC1628F51EE9AE93D">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="5C779C087AB10506C55716B1122509A4" type="personal">
<mods:role id="E212B2E26E26464EDF34959D53D64CEF">
<mods:roleTerm id="9CB9AC2109A94D6116B4304832F90B95">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9128FDF079A08668F0440E686D264FE7">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:name id="EF49A4AD356CB7DF5C42D1A878DA0AF1" type="personal">
<mods:role id="946A40FF8D559B4EE0301F7D74CA20CC">
<mods:roleTerm id="1D1B3200933B54C7330F5143A23FDF1E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E87ACA715DD59503C13F17BB4DE354B1">Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1DF8DE47079C1A1EEF99A15A3FC42D5B">text</mods:typeOfResource>
<mods:relatedItem id="1DE18BFA13C4F35F80A99D01687F2999" type="host">
<mods:titleInfo id="C42F537E14CC6487323144FF86D805A7">
<mods:title id="8C7A42BF8995857E845C7888EE2AA7F3">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="F4FC3E04E703C3799F58CBFC6B7CBB24">
<mods:date id="761130A2AD9938E7772B629C841D6C29">2008</mods:date>
<mods:detail id="029D40D43B9069AAA10A569FEE1BE100" type="pubDate">
<mods:number id="4D38F7ACC5605186C8786EB62FFC2126">2008-06-30</mods:number>
</mods:detail>
<mods:detail id="FD21AC667FBFDAC45600E26ACA7794E5" type="volume">
<mods:number id="9546689F8F73C360A44DF8FD2C2DF4EB">42</mods:number>
</mods:detail>
<mods:detail id="0EE6E92EC4A1AF5F13E660A1F49E7FE9" type="issue">
<mods:number id="109955A8805BA70C68EC362FB8BD901C">21 - 24</mods:number>
</mods:detail>
<mods:extent id="39CEC6C5649E883E2183D7C791053702" unit="page">
<mods:start id="31FFBA39980401F3DED5C6E735DC599E">1477</mods:start>
<mods:end id="0A2A20B077DB89CD8DD710F654A8E405">1508</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="065CC9028C6A5B52B839EF4A14EE0F2D">
<mods:url id="145289D0747EF8E9492C921F7A48601C">http://dx.doi.org/10.1080/00222930802002535</mods:url>
</mods:location>
<mods:classification id="0B7EB11746A7D7021CE854196D2DFC00">journal article</mods:classification>
<mods:identifier id="51F8EF7597B5D1E4E51CE62EB9B528CE" type="DOI">10.1080/00222930802002535</mods:identifier>
<mods:identifier id="A9282B2BEF49CF0047B05E2640E4566A" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="B42C6510F4A472D6A089740CEE31D881" type="Zenodo-Dep">5219397</mods:identifier>
</mods:mods>
<treatment id="1F77BC72FFC23062201CFE77FE04FAA1" LSID="urn:lsid:plazi:treatment:1F77BC72FFC23062201CFE77FE04FAA1" httpUri="http://treatment.plazi.org/id/1F77BC72FFC23062201CFE77FE04FAA1" lastPageId="23" lastPageNumber="1498" pageId="22" pageNumber="1497">
<subSubSection id="DFC45EEFFFC23063201CFE77FCADFD82" box="[525,856,502,526]" pageId="22" pageNumber="1497" type="nomenclature">
<paragraph id="97610D64FFC23063201CFE77FCADFD82" blockId="22.[213,1169,502,623]" box="[525,856,502,526]" pageId="22" pageNumber="1497">
<heading id="CC29BA08FFC23063201CFE77FCADFD82" box="[525,856,502,526]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="1497" reason="3">
<taxonomicName id="50DE76E7FFC23063201CFE77FCADFD82" ID-CoL="5ZY87" authority="Arrow" authorityName="Arrow" box="[525,856,502,526]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="A5AAD176FFC23063201CFE77FCF1FD82" bold="true" box="[525,772,502,526]" italics="true" pageId="22" pageNumber="1497">Coptorhina angolensis</emphasis>
Arrow
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFC230632043FD97FDD4FC57" pageId="22" pageNumber="1497" type="description">
<paragraph id="97610D64FFC230632043FD97FEB4FDE3" blockId="22.[213,1169,502,623]" pageId="22" pageNumber="1497">
(
<figureCitation id="0FE511E1FFC23063204BFD97FD14FDA2" box="[602,737,534,558]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="22" pageNumber="1497">Figures 7, 8</figureCitation>
,
<figureCitation id="0FE511E1FFC2306320E1FD97FCFEFDA2" box="[752,779,534,558]" captionStart="Figure 19" captionStartId="17.[146,213,1510,1532]" captionTargetBox="[146,1104,157,1473]" captionTargetId="figure-41@17.[146,1104,157,1473]" captionTargetPageId="17" captionText="Figure 19. Coptorhina spp., distribution map." figureDoi="http://doi.org/10.5281/zenodo.5219407" httpUri="https://zenodo.org/record/5219407/files/figure.png" pageId="22" pageNumber="1497">19</figureCitation>
)
<taxonomicName id="50DE76E7FFC2306322C4FDB6FD7CFDC3" authority="Arrow 1906: 129" authorityName="Arrow" authorityPageNumber="129" authorityYear="1906" box="[213,649,567,591]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="A5AAD176FFC2306322C4FDB6FE3FFDC3" box="[213,458,567,591]" italics="true" pageId="22" pageNumber="1497">Coptorhina angolensis</emphasis>
<bibRefCitation id="F34F7095FFC2306323C1FDB6FD7CFDC3" author="Arrow GJ" box="[464,649,567,591]" pageId="22" pageNumber="1497" pagination="127 - 136" refId="ref18906" refString="Arrow GJ. 1906. On lamellicorn Coleoptera from Portuguese West Africa, with descriptions of new species. Ann Mag Nat Hist. 18: 127 - 136." type="journal article" year="1906">Arrow 1906:129</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="F34F7095FFC230632085FDB6FCA0FDC3" author="Janssens A" box="[660,853,567,591]" pageId="22" pageNumber="1497" pagination="1 - 104" refId="ref19398" refString="Janssens A. 1939. Coprini. Exploration du Parc National Albert, Mission G. F. de Witte 29: 1 - 104." type="journal article" year="1939">Janssens 1939:35</bibRefCitation>
, 38;
<bibRefCitation id="F34F7095FFC230632198FDB6FBDFFDC3" author="Gillet JJE" box="[905,1066,567,591]" pageId="22" pageNumber="1497" pagination="1 - 100" refId="ref19275" refString="Gillet JJE. 1911. Scarabaeidae: Coprinae I. Coleopterorum Catalogus. W. Junk, Berlin 19: 1 - 100." type="journal article" year="1911">Gillet 1911:47</bibRefCitation>
;
<bibRefCitation id="F34F7095FFC230632624FDB6FEC8FDE3" author="Ferreira M" pageId="22" pageNumber="1497" pagination="5 - 1088" refId="ref19177" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Entomol Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972:356</bibRefCitation>
.
</paragraph>
<paragraph id="97610D64FFC2306322C4FD29FE10FD4C" blockId="22.[213,485,680,704]" box="[213,485,680,704]" pageId="22" pageNumber="1497">
<emphasis id="A5AAD176FFC2306322C4FD29FE10FD4C" box="[213,485,680,704]" italics="true" pageId="22" pageNumber="1497">
<typeStatus id="4865B3C6FFC2306322C4FD29FEF8FD4C" box="[213,269,680,704]" pageId="22" pageNumber="1497">Type</typeStatus>
material examined
</emphasis>
</paragraph>
<paragraph id="97610D64FFC2306322C4FD50FBE3FC06" blockId="22.[213,1169,721,906]" pageId="22" pageNumber="1497">
<taxonomicName id="50DE76E7FFC2306322C4FD50FE2DFD65" box="[213,472,721,745]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="A5AAD176FFC2306322C4FD50FE2DFD65" box="[213,472,721,745]" italics="true" pageId="22" pageNumber="1497">Coptorhina angolensis</emphasis>
</taxonomicName>
.
<typeStatus id="4865B3C6FFC2306323E4FD50FD93FD65" box="[501,614,721,745]" pageId="22" pageNumber="1497" type="lectotype">Lectotype</typeStatus>
with labels ‘‘
<collectingCountry id="EFC94DF4FFC230632132FD50FC8BFD65" box="[803,894,721,745]" name="Angola" pageId="22" pageNumber="1497">Angola</collectingCountry>
. 76.28’’, ‘‘
<taxonomicName id="50DE76E7FFC230632618FD50FE59FC85" authority="Arrow." authorityName="Arrow." class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">Coptorhina angolensis Arrow.</taxonomicName>
Type
<emphasis id="A5AAD176FFC230632012FD75FDE0FC84" box="[515,533,756,776]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
a. m. 1906’’, ‘‘
<taxonomicName id="50DE76E7FFC2306320C8FD70FC94FC85" box="[729,865,753,777]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="genus">Coptorhina</taxonomicName>
<emphasis id="A5AAD176FFC230632160FD75FC76FC84" box="[881,899,756,776]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
<taxonomicName id="50DE76E7FFC230632183FD70FEC6FCA5" authority="Arr. M. E. Bacchus" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">angolensis Arr. M.E. Bacchus</taxonomicName>
det 1975. Lectotype’’, ‘‘Lectotype’’ (BNMH).
<typeStatus id="4865B3C6FFC230632179FC90FBF6FCA5" box="[872,1027,785,809]" pageId="22" pageNumber="1497" type="paralectotype">Paralectotype</typeStatus>
with labels ‘‘
<collectingCountry id="EFC94DF4FFC2306322CEFCB0FECFFCC5" box="[223,314,817,841]" name="Angola" pageId="22" pageNumber="1497">Angola</collectingCountry>
. 76.28’’, ‘‘Paralectotype’’, ‘‘
<taxonomicName id="50DE76E7FFC23063207FFCB0FD03FCC5" box="[622,758,817,841]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="genus">Coptorhina</taxonomicName>
<emphasis id="A5AAD176FFC2306320ECFCB4FCFAFCC5" box="[765,783,821,841]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
<taxonomicName id="50DE76E7FFC230632104FCB0FB92FCC5" authority="Arr. M. E. Bacchus" box="[789,1127,817,841]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">angolensis Arr. M.E. Bacchus</taxonomicName>
det 1975. Paralectotype’’ and with labels ‘‘
<collectingCountry id="EFC94DF4FFC23063208BFCD3FD00FCE6" box="[666,757,850,874]" name="Angola" pageId="22" pageNumber="1497">Angola</collectingCountry>
. 76.28’’, ‘‘
<taxonomicName id="50DE76E7FFC23063217AFCD3FC06FCE6" box="[875,1011,850,874]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="genus">Coptorhina</taxonomicName>
<emphasis id="A5AAD176FFC2306321EEFCD4FBE4FCE5" box="[1023,1041,853,873]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
<taxonomicName id="50DE76E7FFC23063260DFCD3FE5AFC06" authority="Arr. M. E. Bacchus" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="angolensis">angolensis Arr. M.E. Bacchus</taxonomicName>
det 1975. Paralectotype’’, ‘‘Paralectotype’’ (BNMH).
</paragraph>
<paragraph id="97610D64FFC2306322C4FC42FDD4FC57" blockId="22.[213,545,963,987]" box="[213,545,963,987]" pageId="22" pageNumber="1497">
<emphasis id="A5AAD176FFC2306322C4FC42FDD4FC57" box="[213,545,963,987]" italics="true" pageId="22" pageNumber="1497">Additional material examined</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFC2306222C4FC6DFE04FAA1" lastPageId="23" lastPageNumber="1498" pageId="22" pageNumber="1497" type="materials_examined">
<paragraph id="97610D64FFC2306322C4FC6DFCEDFBE8" blockId="22.[213,1169,1004,1124]" pageId="22" pageNumber="1497">
<materialsCitation id="27B60739FFC2306322C4FC6DFC35FBA8" collectingDate="1954-08" collectionCode="BMSA, MHNG, SAMC" collectorName="Chassot" country="Angola" latitude="-15.15" location="Angola" longLatPrecision="1286" longitude="15.15" municipality="Bailundo" pageId="22" pageNumber="1497" specimenCount="1" stateProvince="Huila">
<emphasis id="A5AAD176FFC2306322C4FC6DFED1FB88" bold="true" box="[213,292,1004,1028]" pageId="22" pageNumber="1497">
<collectingCountry id="EFC94DF4FFC2306322C4FC6DFED1FB88" box="[213,292,1004,1028]" name="Angola" pageId="22" pageNumber="1497">Angola</collectingCountry>
</emphasis>
:
<collectingMunicipality id="7705971EFFC230632324FC6DFE68FB88" box="[309,413,1004,1028]" pageId="22" pageNumber="1497">Bailundo</collectingMunicipality>
[not traced], 1
<emphasis id="A5AAD176FFC230632058FC6EFDAEFB8F" box="[585,603,1007,1027]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
(
<collectionCode id="F1CF95A1FFC23063207DFC6DFD35FB88" box="[620,704,1004,1028]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="22" pageNumber="1497">BMSA</collectionCode>
);
<location id="92015BBFFFC2306320C7FC6DFCD8FB88" LSID="urn:lsid:plazi:treatment:1F77BC72FFC23062201CFE77FE04FAA1:92015BBFFFC2306320C7FC6DFCD8FB88" box="[726,813,1004,1028]" country="Angola" latitude="-15.15" longLatPrecision="1286" longitude="15.15" municipality="Bailundo" name="Angola" pageId="22" pageNumber="1497" stateProvince="Huila">Angola</location>
,
<date id="E3602BA4FFC23063212AFC6DFC3BFB88" box="[827,974,1004,1028]" pageId="22" pageNumber="1497" value="1954-08">
<collectingDate id="F324D24CFFC23063212AFC6DFC3BFB88" box="[827,974,1004,1028]" pageId="22" pageNumber="1497" value="1954-08">August 1954</collectingDate>
</date>
,
<collectorName id="3A2B68B2FFC2306321CDFC6DFBC2FB88" box="[988,1079,1004,1028]" pageId="22" pageNumber="1497">Chassot</collectorName>
leg. 1
<emphasis id="A5AAD176FFC23063266EFC6EFB64FB8F" box="[1151,1169,1007,1027]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
(
<collectionCode id="F1CF95A1FFC2306322CCFB8DFEC8FBA8" box="[221,317,1036,1060]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="22" pageNumber="1497" type="Museum">MHNG</collectionCode>
); 1
<emphasis id="A5AAD176FFC230632371FB8EFE87FBAF" box="[352,370,1039,1059]" italics="true" pageId="22" pageNumber="1497">„</emphasis>
(
<collectionCode id="F1CF95A1FFC230632393FB8DFE23FBA8" box="[386,470,1036,1060]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="22" pageNumber="1497">BMSA</collectionCode>
);
<collectingRegion id="551AC386FFC2306323FAFB8DFDDEFBA8" box="[491,555,1036,1060]" country="Angola" name="Huila" pageId="22" pageNumber="1497">Huila</collectingRegion>
<geoCoordinate id="F2EA6BA3FFC230632022FB8DFD7AFBA8" box="[563,655,1036,1060]" degrees="15" direction="south" minutes="009" orientation="latitude" pageId="22" pageNumber="1497" precision="925" value="-15.15">
15
<emphasis id="A5AAD176FFC23063205FFB8DFDADFBA8" bold="true" box="[590,600,1036,1060]" italics="true" pageId="22" pageNumber="1497">°</emphasis>
009 S
</geoCoordinate>
<geoCoordinate id="F2EA6BA3FFC230632089FB8DFD0DFBA8" box="[664,760,1036,1060]" degrees="15" direction="east" minutes="009" orientation="longitude" pageId="22" pageNumber="1497" precision="925" value="15.15">
15
<emphasis id="A5AAD176FFC2306320A3FB8DFD49FBA8" bold="true" box="[690,700,1036,1060]" italics="true" pageId="22" pageNumber="1497">°</emphasis>
009 E
</geoCoordinate>
, 2 spm. (
<collectionCode id="F1CF95A1FFC230632173FB8DFC4DFBA8" box="[866,952,1036,1060]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="22" pageNumber="1497" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFC2306321DCFB8DFCE1FBE8" collectingDate="1999-12-19" collectionCode="UMO" collectorName="Mann &amp; Marais &amp; Newman" country="Namibia" latitude="-18.166666" location="River" municipality="Park Nova" pageId="22" pageNumber="1497" specimenCount="1" stateProvince="Zambezi">
<emphasis id="A5AAD176FFC2306321DCFB8DFBD9FBA8" bold="true" box="[973,1068,1036,1060]" pageId="22" pageNumber="1497">
<collectingCountry id="EFC94DF4FFC2306321DCFB8DFBD9FBA8" box="[973,1068,1036,1060]" name="Namibia" pageId="22" pageNumber="1497">Namibia</collectingCountry>
</emphasis>
:
<collectingRegion id="551AC386FFC23063262DFB8DFB64FBA8" box="[1084,1169,1036,1060]" country="Namibia" name="Zambezi" pageId="22" pageNumber="1497">Caprivi</collectingRegion>
<collectingMunicipality id="7705971EFFC2306322C4FBADFEAFFBC8" box="[213,346,1068,1092]" pageId="22" pageNumber="1497">Park Nova</collectingMunicipality>
,
<quantity id="5026A081FFC23063237CFBADFE56FBC8" box="[365,419,1068,1092]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="1497" unit="km" value="5.0">5 km</quantity>
N of
<collectingRegion id="551AC386FFC2306323E8FBADFD9AFBC8" box="[505,623,1068,1092]" country="Namibia" name="Kavango West" pageId="22" pageNumber="1497">Okavango</collectingRegion>
<location id="92015BBFFFC23063206CFBADFD49FBC8" LSID="urn:lsid:plazi:treatment:1F77BC72FFC23062201CFE77FE04FAA1:92015BBFFFC23063206CFBADFD49FBC8" box="[637,700,1068,1092]" country="Namibia" latitude="-18.166666" municipality="Park Nova" name="River" pageId="22" pageNumber="1497" stateProvince="Zambezi">River</location>
[
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18
<emphasis id="A5AAD176FFC2306320FAFBADFD00FBC8" bold="true" box="[747,757,1068,1092]" italics="true" pageId="22" pageNumber="1497">°</emphasis>
10 S
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21
<emphasis id="A5AAD176FFC23063215AFBADFCA0FBC8" bold="true" box="[843,853,1068,1092]" italics="true" pageId="22" pageNumber="1497">°</emphasis>
449 E],
<date id="E3602BA4FFC2306321BAFBADFB79FBC8" box="[939,1164,1068,1092]" pageId="22" pageNumber="1497" value="1999-12-19">
<collectingDate id="F324D24CFFC2306321BAFBADFB79FBC8" box="[939,1164,1068,1092]" pageId="22" pageNumber="1497" value="1999-12-19">19 December 1999</collectingDate>
</date>
,
<collectorName id="3A2B68B2FFC2306322C4FBCDFEE8FBE8" box="[213,285,1100,1124]" pageId="22" pageNumber="1497">Mann</collectorName>
,
<collectorName id="3A2B68B2FFC23063233BFBCDFE8BFBE8" box="[298,382,1100,1124]" pageId="22" pageNumber="1497">Marais</collectorName>
, and
<collectorName id="3A2B68B2FFC2306323AFFBCDFDD1FBE8" box="[446,548,1100,1124]" pageId="22" pageNumber="1497">Newman</collectorName>
leg., 9 spm. (
<collectionCode id="F1CF95A1FFC2306320D3FBCDFCF9FBE8" box="[706,780,1100,1124]" country="USA" httpUri="http://grbio.org/cool/asxa-80fx" name="University of Maine" pageId="22" pageNumber="1497">UMO</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="97610D64FFC2306322C4FB1CFEB7FB39" blockId="22.[213,322,1181,1205]" box="[213,322,1181,1205]" pageId="22" pageNumber="1497">
<emphasis id="A5AAD176FFC2306322C4FB1CFEB7FB39" box="[213,322,1181,1205]" italics="true" pageId="22" pageNumber="1497">Diagnosis</emphasis>
</paragraph>
<paragraph id="97610D64FFC2306322C4FB47FBF8FAD3" blockId="22.[213,1169,1222,1375]" pageId="22" pageNumber="1497">
This species is most similar to
<taxonomicName id="50DE76E7FFC230632021FB47FD65FB52" authorityName="Hope. The" authorityYear="2008" box="[560,656,1222,1246]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="klugii">
<emphasis id="A5AAD176FFC230632021FB47FD65FB52" box="[560,656,1222,1246]" italics="true" pageId="22" pageNumber="1497">C. klugii</emphasis>
</taxonomicName>
but can be separated from it by the shape of the transverse carina of the pronotum which has four distinct tubercles in larger specimens (
<figureCitation id="0FE511E1FFC230632346FA87FE42FA92" box="[343,439,1286,1310]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="22" pageNumber="1497">Figure 7</figureCitation>
). From other species it differs in having narrow lateral borders of the pronotum and evenly rounded lateral sides of the pronotum. From
<taxonomicName id="50DE76E7FFC230632666FAA6FEC3FAD3" authorityName="Gillet" authorityYear="1932" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1497" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFC230632666FAA6FEC3FAD3" italics="true" pageId="22" pageNumber="1497">C. nitefacta</emphasis>
</taxonomicName>
it also differs in having densely punctured upper side of body.
</paragraph>
<paragraph id="97610D64FFC2306322C4FA19FEA1FA3C" blockId="22.[213,340,1432,1456]" box="[213,340,1432,1456]" pageId="22" pageNumber="1497">
<emphasis id="A5AAD176FFC2306322C4FA19FEA1FA3C" box="[213,340,1432,1456]" italics="true" pageId="22" pageNumber="1497">Description</emphasis>
</paragraph>
<paragraph id="97610D64FFC2306322C4FA41FCA8FA75" blockId="22.[213,1169,1472,1529]" pageId="22" pageNumber="1497">Strongly convex, black to dark brown, subopaque beetle. Body length 11.6–18.5 mm, width 7.5–12.0 mm. Dorsal surface without visible setae.</paragraph>
<paragraph id="97610D64FFC2306222C4F9B3FED7FE93" blockId="22.[213,1169,1585,1706]" lastBlockId="23.[146,1103,168,287]" lastPageId="23" lastPageNumber="1498" pageId="22" pageNumber="1497">
<emphasis id="A5AAD176FFC2306322C4F9B3FEE4F9C6" box="[213,273,1586,1610]" italics="true" pageId="22" pageNumber="1497">Head</emphasis>
. Clypeus with protruding anterior angles separated by deep sinuation (
<figureCitation id="0FE511E1FFC2306322CDF9D3FECBF9E6" box="[220,318,1618,1642]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="22" pageNumber="1497">Figure 6</figureCitation>
). Sinuation right-angled to broadly rounded. Protruding angles acute to narrowly rounded apically, with short longitudinal carinae on the upper side. Genae obtusely rounded. Genal sutures feebly distinct. Lateral margins of clypeus sinuate near genal sutures in some individuals. Frontal suture feebly visible, broadly interrupted in the middle. Head very densely and evenly punctuate, punctures sometimes adjacent and sculpture appears rugose. Antennal club dark-brown.
</paragraph>
<paragraph id="97610D64FFC330622283FED9FD9DFDA3" blockId="23.[146,1103,343,559]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FED9FEF4FEFC" box="[146,257,344,368]" italics="true" pageId="23" pageNumber="1498">Pronotum</emphasis>
. Trapezoidal, two times wider than long. Anterior margin and base not bordered. Sides more or less evenly rounded. Lateral border relatively narrow, evenly tapering basally and apically. Pronotum divided into larger anterior part and smaller posterior part by a transverse carina (
<figureCitation id="0FE511E1FFC330622081FE36FD04FE43" box="[656,753,439,463]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="23" pageNumber="1498">Figure 7</figureCitation>
). The carina is more developed and has four tubercles in larger specimens. Anterior part of pronotum rugose to granulate, excavated laterally in larger specimens. Posterior part in specimens with well developed carina densely punctuate.
</paragraph>
<paragraph id="97610D64FFC330622283FDE6FE5CFD52" blockId="23.[146,1102,615,734]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FDE6FF2CFDF3" box="[146,217,615,639]" italics="true" pageId="23" pageNumber="1498">Elytra</emphasis>
. Striae fine but distinct, punctate (punctures separated by 3–4 puncture diameter). Intervals flat, very densely punctuate, punctures separated by about 1 puncture diameter, sometimes almost adjacent; their margins sometimes indistinct especially in apical part.
</paragraph>
<paragraph id="97610D64FFC330622283FC97FD9EFCE2" blockId="23.[146,1103,790,878]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FC97FEF4FCA2" box="[146,257,790,814]" italics="true" pageId="23" pageNumber="1498">Underside</emphasis>
. Pygidium with fine border, its disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of metasternum which is very sparsely to almost indistinctly punctuate.
</paragraph>
<paragraph id="97610D64FFC330622283FC27FC8CFC32" blockId="23.[146,889,934,958]" box="[146,889,934,958]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FC27FF09FC32" box="[146,252,934,958]" italics="true" pageId="23" pageNumber="1498">Aedeagus</emphasis>
. Parameres with widely rounded apices in lateral view.
</paragraph>
<paragraph id="97610D64FFC330622283FC77FEFFFB82" blockId="23.[146,266,1014,1038]" box="[146,266,1014,1038]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FC77FEFFFB82" box="[146,266,1014,1038]" italics="true" pageId="23" pageNumber="1498">Variability</emphasis>
</paragraph>
<paragraph id="97610D64FFC330622283FB9FFD4FFBFA" blockId="23.[146,1103,1054,1142]" pageId="23" pageNumber="1498">Except for the body size variability the beetles differ in shape and sculpture of pronotum. Smaller specimens have less developed transverse pronotal carina and little excavated anterolateral sides of pronotum.</paragraph>
<paragraph id="97610D64FFC330622283FB2FFEEDFB4A" blockId="23.[146,280,1198,1222]" box="[146,280,1198,1222]" pageId="23" pageNumber="1498">
<emphasis id="A5AAD176FFC330622283FB2FFEEDFB4A" box="[146,280,1198,1222]" italics="true" pageId="23" pageNumber="1498">Distribution</emphasis>
</paragraph>
<paragraph id="97610D64FFC330622283FB54FE04FAA1" blockId="23.[146,1103,1237,1325]" pageId="23" pageNumber="1498">
The range of this species is not quite clear because of limited material available. It is known from a few localities in
<collectingCountry id="EFC94DF4FFC330622011FB74FDA1FA81" box="[512,596,1269,1293]" name="Angola" pageId="23" pageNumber="1498">Angola</collectingCountry>
and was recently collected in north-eastern
<collectingCountry id="EFC94DF4FFC330622283FA94FF00FAA1" box="[146,245,1301,1325]" name="Namibia" pageId="23" pageNumber="1498">Namibia</collectingCountry>
(
<collectingRegion id="551AC386FFC330622317FA94FEA8FAA1" box="[262,349,1301,1325]" country="Namibia" name="Zambezi" pageId="23" pageNumber="1498">Caprivi</collectingRegion>
) (
<figureCitation id="0FE511E1FFC330622367FA94FE11FAA1" box="[374,484,1301,1325]" captionStart="Figure 19" captionStartId="17.[146,213,1510,1532]" captionTargetBox="[146,1104,157,1473]" captionTargetId="figure-41@17.[146,1104,157,1473]" captionTargetPageId="17" captionText="Figure 19. Coptorhina spp., distribution map." figureDoi="http://doi.org/10.5281/zenodo.5219407" httpUri="https://zenodo.org/record/5219407/files/figure.png" pageId="23" pageNumber="1498">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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