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<document id="254FB21C730688105A2A11FEE23EC5EA" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D64256FFE382AA3CE14DA0FAF7" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diaulota orientalis Tasaku, Ono &amp; Maruyama 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="269" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="268" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E6713E474C4E7B4DE2B320495AA2B716">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="75CE5B9F161481B9C9A4552790346706">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="68F84F9B8511716340DD223BE7B34CA1">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="EBE927379E67063136AE622DF5B617BD">Ono, Hiroki</mods:namePart>
<mods:affiliation id="69B8D6426A6E2EE353E75C2B7E487B98">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
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<mods:namePart id="DF3B13DE6AD288FE348D31EE90DC7993">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="F36BE45C8F8B1B91A85C6CB782B7E40E">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<taxonomicName id="4C574D434256FFE082AA3CE14CCDF969" authority="Tasaku, Ono &amp; Maruyama" authorityName="Tasaku, Ono &amp; Maruyama" authorityYear="2023" box="[151,726,1713,1740]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B923EAD24256FFE082AA3CE14F61F96E" bold="true" box="[151,378,1713,1740]" italics="true" pageId="17" pageNumber="268">Diaulota orientalis</emphasis>
Tasaku, Ono &amp; Maruyama
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,
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<paragraph id="8BE836C04256FFE082AA3C864CF5F94D" blockId="17.[151,805,1713,1811]" box="[151,750,1749,1776]" pageId="17" pageNumber="268">[Japanese name: Higashi-shiributo-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082AA3CA94F5FF8B1" box="[151,324,1785,1811]" pageId="17" pageNumber="268" type="description">
<paragraph id="8BE836C04256FFE082AA3CA94F5FF8B1" blockId="17.[151,805,1713,1811]" box="[151,324,1785,1811]" pageId="17" pageNumber="268">
(
<figureCitation id="136C2A454256FFE082A23CA94EE0F8B1" box="[159,251,1785,1811]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="17" pageNumber="268">Figs. 12</figureCitation>
&amp;
<figureCitation id="136C2A454256FFE083223CA94F27F8B1" box="[287,316,1785,1811]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="17" pageNumber="268">24</figureCitation>
)
</paragraph>
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<emphasis id="B923EAD24256FFE082AA3D114F5EF8F9" bold="true" box="[151,325,1857,1883]" pageId="17" pageNumber="268">Type material.</emphasis>
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<typeStatus id="54EC88624256FFE083713D114FA0F8F9" box="[332,443,1857,1883]" pageId="17" pageNumber="268" type="holotype">Holotype</typeStatus>
.
</emphasis>
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, “
<collectingCountry id="F34076504256FFE080243D114C9EF8F9" box="[537,645,1857,1883]" name="Japan" pageId="17" pageNumber="268">NIPPON</collectingCountry>
: Kanagawa-ken, / Jôgashima,
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<collectingDate id="EFADE9E84256FFE081E43D114A47F8F9" box="[985,1116,1857,1884]" pageId="17" pageNumber="268" value="2003-06-16">16 VI 2003</collectingDate>
</date>
, / Maruyama M. /
<emphasis id="B923EAD24256FFE087113D124F63F8DF" bold="true" pageId="17" pageNumber="268">Dzâ川県 µヶ島(潮®帯)</emphasis>
” (
<figureCitation id="136C2A454256FFE083AE3D354FE2F8DD" box="[403,505,1893,1919]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="17" pageNumber="268">Fig. 12A</figureCitation>
) (
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)
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.
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<typeStatus id="54EC88624256FFE080563D354CFEF8DD" box="[619,741,1893,1919]" pageId="17" pageNumber="268" type="paratype">Paratypes</typeStatus>
.
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:
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:
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:
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,
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,
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,
<date id="FFE910004256FFE083663DD94FB8F806" box="[347,419,1929,1956]" pageId="17" pageNumber="268" value="2021-03-31">
31.
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. 2021, H.
<location id="8E88601B4256FFE0802E3DD94C58F801" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4256FFE0802E3DD94C58F801" box="[531,579,1929,1955]" country="Japan" name="Ono" pageId="17" pageNumber="268">Ono</location>
(
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;
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<specimenCount id="9D51FD494256FFE080963DD94D18F801" box="[683,771,1929,1955]" count="2" pageId="17" pageNumber="268" type="male">2 males</specimenCount>
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15.
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. 2008,
<collectorName id="26A253164256FFE086D73DDA4B20F801" box="[1258,1339,1929,1956]" pageId="17" pageNumber="268">H. Ono</collectorName>
(
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).
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<emphasis id="B923EAD24256FFE082AA3DFD4F13F865" bold="true" box="[151,264,1965,1991]" pageId="17" pageNumber="268">Tôkyô-to</emphasis>
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<specimenCount id="9D51FD494256FFE083293DFD4F79F865" box="[276,354,1965,1991]" count="1" pageId="17" pageNumber="268" type="male">1 male</specimenCount>
,
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,
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,
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,
<date id="FFE910004256FFE080DD3DFD4D28F86A" box="[736,819,1965,1992]" pageId="17" pageNumber="268" value="2022-07-15">
15.
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. 2022, Y. Tasaku (cYT);
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, ditto,
<date id="FFE910004256FFE087663DFD4B8DF86A" box="[1371,1430,1965,1992]" pageId="17" pageNumber="268" value="2022-05-17">
17.
<collectionCode id="ED46AE054256FFE087B83DFE4B8DF86A" box="[1413,1430,1966,1992]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="268" type="Museum">V</collectionCode>
</date>
</materialsCitation>
. 2022, H. Ono (cHO);
<materialsCitation id="3B3F3C9D4256FFE083B53D814C4CF84E" box="[392,599,2001,2028]" collectionCode="V" collectorName="Izu-oshima &amp; Okada" country="Japan" location="Noda-hama" pageId="17" pageNumber="268" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D51FD494256FFE083B53D814FCFF849" box="[392,468,2001,2027]" count="1" pageId="17" pageNumber="268" type="male">1 male</specimenCount>
, ditto,
<date id="FFE910004256FFE080233D814C4CF84E" box="[542,599,2001,2028]" pageId="17" pageNumber="268" value="2022-05-18">
18.
<collectionCode id="ED46AE054256FFE0807B3D824C4CF84E" box="[582,599,2002,2028]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="268" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B3F3C9D4256FFE0806A3D824A17F849" box="[599,1036,2001,2028]" collectorName="H. Ono" country="Japan" location="Kanagawa-ken" pageId="17" pageNumber="268" specimenCount="1" typeStatus="paratype">
. 2022,
<collectorName id="26A253164256FFE080953D824CE2F849" box="[680,761,2001,2028]" pageId="17" pageNumber="268">H. Ono</collectorName>
(cHO).
<location id="8E88601B4256FFE0816E3D814A17F849" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4256FFE0816E3D814A17F849" box="[851,1036,2001,2027]" country="Japan" name="Kanagawa-ken" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE0816E3D814A17F849" bold="true" box="[851,1036,2001,2027]" pageId="17" pageNumber="268">Kanagawa-ken</emphasis>
</location>
</materialsCitation>
<emphasis id="B923EAD24256FFE086313D814A0AF849" bold="true" box="[1036,1041,2001,2027]" pageId="17" pageNumber="268">:</emphasis>
<materialsCitation id="3B3F3C9D4256FFE0862A3D814B83F849" box="[1047,1432,2001,2027]" country="Japan" location="Yokosuka-shi" pageId="17" pageNumber="268" specimenCount="22" specimenCount-female="13" specimenCount-male="9" typeStatus="paratype">
<specimenCount id="9D51FD494256FFE0862A3D814A75F849" box="[1047,1134,2001,2027]" count="9" pageId="17" pageNumber="268" type="male">9 males</specimenCount>
,
<specimenCount id="9D51FD494256FFE086453D814AEFF849" box="[1144,1268,2001,2027]" count="13" pageId="17" pageNumber="268" type="female">13 females</specimenCount>
,
<location id="8E88601B4256FFE086C13D814B83F849" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4256FFE086C13D814B83F849" box="[1276,1432,2001,2027]" country="Japan" name="Yokosuka-shi" pageId="17" pageNumber="268">Yokosuka-shi</location>
</materialsCitation>
,
</paragraph>
<paragraph id="8BE836C04255FFE382AA3AC74DA4FEBF" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
Wada-nagahama (Nahama),
<date id="FFE910004255FFE383EA3AC64C4CFF13" box="[471,599,150,177]" pageId="18" pageNumber="269" value="2007-04-05">
5.
<collectionCode id="ED46AE054255FFE383CE3AC74C16FF13" box="[499,525,151,177]" pageId="18" pageNumber="269">IV</collectionCode>
<materialsCitation id="3B3F3C9D4255FFE380303AC74C4CFF13" box="[525,599,151,177]" collectionCode="KUM, OMNH, HIWA" collectorName="I. Kawashima" pageId="18" pageNumber="269" specimenCount="16" specimenCount-female="9" specimenCount-male="7">. 2007</materialsCitation>
</date>
,
<collectorName id="26A253164255FFE3805F3AC74CE5FF13" box="[610,766,151,177]" pageId="18" pageNumber="269">I. Kawashima</collectorName>
(
<collectionCode id="ED46AE054255FFE381333AC74D4FFF13" box="[782,852,151,177]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="18" pageNumber="269" type="Herbarium">KUM</collectionCode>
,
<collectionCode id="ED46AE054255FFE381623AC74DACFF13" box="[863,951,151,177]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="18" pageNumber="269" type="Museum">OMNH</collectionCode>
,
<collectionCode id="ED46AE054255FFE381FE3AC74A14FF13" box="[963,1039,151,177]" pageId="18" pageNumber="269">HIWA</collectionCode>
, cYT);
<specimenCount id="9D51FD494255FFE386583AC74AA4FF13" box="[1125,1215,151,177]" count="7" pageId="18" pageNumber="269" type="male">7 males</specimenCount>
,
<specimenCount id="9D51FD494255FFE386F73AC74B21FF13" box="[1226,1338,151,177]" count="9" pageId="18" pageNumber="269" type="female">9 females</specimenCount>
, ditto,
<date id="FFE910004255FFE387BA3AC64EE2FF77" pageId="18" pageNumber="269" value="2007-04-05">
5.
<collectionCode id="ED46AE054255FFE382AA3AEB4EAAFF77" box="[151,177,187,213]" pageId="18" pageNumber="269">IV</collectionCode>
<materialsCitation id="3B3F3C9D4255FFE3828C3AEB4EE2FF77" box="[177,249,187,213]" collectionCode="KUM" collectorName="M. Asano" county="Aburatsubo" location="Miura-shi" municipality="Koajiro" pageId="18" pageNumber="269" specimenCount="8" specimenCount-female="4" specimenCount-male="4">. 2007</materialsCitation>
</date>
,
<collectorName id="26A253164255FFE3833F3AEB4F75FF77" box="[258,366,187,213]" pageId="18" pageNumber="269">M.Asano</collectorName>
(
<collectionCode id="ED46AE054255FFE383413AEB4FD8FF77" box="[380,451,187,213]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="18" pageNumber="269" type="Herbarium">KUM</collectionCode>
);
<specimenCount id="9D51FD494255FFE383E83AEB4C36FF77" box="[469,557,187,213]" count="4" pageId="18" pageNumber="269" type="male">4 males</specimenCount>
,
<specimenCount id="9D51FD494255FFE3800A3AEB4CBFFF77" box="[567,676,187,213]" count="4" pageId="18" pageNumber="269" type="female">4 females</specimenCount>
,
<location id="8E88601B4255FFE380933AEB4D04FF77" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4255FFE380933AEB4D04FF77" box="[686,799,187,213]" county="Aburatsubo" municipality="Koajiro" name="Miura-shi" pageId="18" pageNumber="269">Miura-shi</location>
,
<location id="8E88601B4255FFE381143AEB4DD3FF77" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4255FFE381143AEB4DD3FF77" box="[809,968,187,213]" county="Aburatsubo" municipality="Koajiro" name="Misaki-machi" pageId="18" pageNumber="269">Misaki-machi</location>
,
<collectingMunicipality id="6B8CACBA4255FFE381EC3AEB4A31FF77" box="[977,1066,187,213]" pageId="18" pageNumber="269">Koajiro</collectingMunicipality>
,
<collectingCounty id="62894E4C4255FFE3860F3AEB4AA3FF77" box="[1074,1208,187,213]" pageId="18" pageNumber="269">Aburatsubo</collectingCounty>
,
<date id="FFE910004255FFE386FC3AEB4B5AFF77" box="[1217,1345,187,213]" pageId="18" pageNumber="269" value="2018-05-29">
29.
<collectionCode id="ED46AE054255FFE386D43AEB4AE1FF77" box="[1257,1274,187,213]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="269" type="Museum">V</collectionCode>
<materialsCitation id="3B3F3C9D4255FFE386C73AEB4B5AFF77" box="[1274,1345,187,213]" collectingDate="2003-06-16" collectionCode="KUM, OMNH" collectorName="H. Ono &amp; M. Maruyama &amp; Shizuoka-ken" country="Japan" location="Shimoda-shi" municipality="Tarai-misaki" pageId="18" pageNumber="269" specimenCount="18" specimenCount-female="12" specimenCount-male="6">. 2018</materialsCitation>
</date>
,
<collectorName id="26A253164255FFE387763AEB4B87FF77" box="[1355,1436,187,213]" pageId="18" pageNumber="269">H. Ono</collectorName>
(cHO);
<specimenCount id="9D51FD494255FFE382D03A8E4F5EFF5B" box="[237,325,222,249]" count="5" pageId="18" pageNumber="269" type="male">5 males</specimenCount>
,
<specimenCount id="9D51FD494255FFE3836D3A8F4FD7FF5B" box="[336,460,223,249]" count="11" pageId="18" pageNumber="269" type="female">11 females</specimenCount>
, same data as holotype,
<collectorName id="26A253164255FFE380E43A8F4D63FF5B" box="[729,888,223,249]" pageId="18" pageNumber="269">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE054255FFE381BA3A8F4DD6FF5B" box="[903,973,223,249]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="18" pageNumber="269" type="Herbarium">KUM</collectionCode>
,
<collectionCode id="ED46AE054255FFE381EA3A8F4A34FF5B" box="[983,1071,223,249]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="18" pageNumber="269" type="Museum">OMNH</collectionCode>
, cYT).
<emphasis id="B923EAD24255FFE386BE3A8E4B34FF5A" bold="true" box="[1155,1327,222,248]" pageId="18" pageNumber="269">
<collectorName id="26A253164255FFE386BE3A8E4B30FF5A" box="[1155,1323,222,248]" pageId="18" pageNumber="269">Shizuoka-ken</collectorName>
:
</emphasis>
<specimenCount id="9D51FD494255FFE3870B3A8F4B98FF5B" box="[1334,1411,223,249]" count="1" pageId="18" pageNumber="269" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD494255FFE387B33A8F4EFDFEBF" count="1" pageId="18" pageNumber="269" type="female">1 female</specimenCount>
,
<location id="8E88601B4255FFE382CC3B534F9AFEBF" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4255FFE382CC3B534F9AFEBF" box="[241,385,259,285]" municipality="Tarai-misaki" name="Shimoda-shi" pageId="18" pageNumber="269">Shimoda-shi</location>
,
<collectingMunicipality id="6B8CACBA4255FFE383B13B534C07FEBF" box="[396,540,259,285]" pageId="18" pageNumber="269">Tarai-misaki</collectingMunicipality>
,
<date id="FFE910004255FFE380153B534CAFFEBF" box="[552,692,259,285]" pageId="18" pageNumber="269" value="2003-04-21">
21.
<collectionCode id="ED46AE054255FFE3806F3B534C77FEBF" box="[594,620,259,285]" pageId="18" pageNumber="269">IV</collectionCode>
<materialsCitation id="3B3F3C9D4255FFE380563B534CAFFEBF" box="[619,692,259,285]" collectionCode="KUM" country="Japan" location="Moriguchi" pageId="18" pageNumber="269" specimenCount="1">. 2003</materialsCitation>
</date>
, M.
<location id="8E88601B4255FFE380DB3B534D47FEBF" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE382AA3CE14DA0FAF7:8E88601B4255FFE380DB3B534D47FEBF" box="[742,860,259,285]" country="Japan" name="Moriguchi" pageId="18" pageNumber="269">Moriguchi</location>
(
<collectionCode id="ED46AE054255FFE381513B534DA8FEBF" box="[876,947,259,285]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="18" pageNumber="269" type="Herbarium">KUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4255FFE382FA3B764D84FE0F" pageId="18" pageNumber="269" type="diagnosis">
<paragraph id="8BE836C04255FFE382FA3B764D84FE0F" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3B764F59FEE2" bold="true" box="[199,322,294,320]" pageId="18" pageNumber="269">Diagnosis.</emphasis>
This new species is very similar to
<taxonomicName id="4C574D434255FFE380DA3B774D51FEE2" authorityName="K. Sawada" authorityYear="1955" box="[743,842,295,321]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24255FFE380DA3B774CE0FEE2" box="[743,763,295,320]" italics="true" pageId="18" pageNumber="269">D</emphasis>
.
<emphasis id="B923EAD24255FFE381313B774D51FEE2" box="[780,842,295,320]" italics="true" pageId="18" pageNumber="269">uenoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434255FFE381BB3B774A38FEE3" authorityName="Ahn" authorityYear="2023" box="[902,1059,295,321]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24255FFE381BB3B774D81FEE2" box="[902,922,295,320]" italics="true" pageId="18" pageNumber="269">D</emphasis>
.
<emphasis id="B923EAD24255FFE381963B774A38FEE3" box="[939,1059,295,321]" italics="true" pageId="18" pageNumber="269">submarina</emphasis>
</taxonomicName>
but, can be distinguished by the combination of following characteristics: apex of median lobe truncate in ventral view; apical sclerite rounded in lateral view (
<figureCitation id="136C2A454255FFE383123B3F4F8DFE2B" box="[303,406,367,393]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12E</figureCitation>
), narrow toward apical margin in ventral view (
<figureCitation id="136C2A454255FFE381FF3B3F4A33FE2B" box="[962,1064,367,393]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12F</figureCitation>
); suspensonia spoon-shaped and broad in apical half (
<figureCitation id="136C2A454255FFE383423BC34FF8FE0F" box="[383,483,403,429]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12F</figureCitation>
); spermatheca twice curved (
<figureCitation id="136C2A454255FFE381173BC34D89FE0F" box="[810,914,403,429]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4255FFE382FA3BE64B95FBDF" pageId="18" pageNumber="269" type="description">
<paragraph id="8BE836C04255FFE382FA3BE64FD8FC9B" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3BE64F42FE72" bold="true" box="[199,345,438,464]" pageId="18" pageNumber="269">Description.</emphasis>
Body robust, flattened. Ground color reddish brown, abdominal segment VI black, mouth parts yellow (
<figureCitation id="136C2A454255FFE382C83B8B4F83FE57" box="[245,408,475,501]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12A &amp; B</figureCitation>
). Head quadrate, broader than pronotum in males, as broad as pronotum in females; eyes relatively small; antennae shorter than head and pronotum combined; segment I longest; segment II as long as XI; segment III shorter than II, about 1.5 times longer than IV; segments IVVIII as long as wide; segments IX and X slightly wider than long; segment XI about 1.5 times longer than wide. Labrum semicircular, with about 20 setae. Mandibles almost symmetric, with a small tooth. Maxillary palpi 4 segmented; segment I very short, as long as IV; segment II quadrate, 2 times longer than I; segment III oval, slightly longer than II; segment IV thin. Labium: prementum without medial seta; ligula unilobed, short; labial palpi 3 segmented, segment II as thin as I, shorter than twice length of I; segment III thin, shorter than I (
<figureCitation id="136C2A454255FFE380FD38874D3CFD53" box="[704,807,727,753]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12C</figureCitation>
). Mentum almost trapezoidal (
<figureCitation id="136C2A454255FFE3864138874AF8FD53" box="[1148,1251,727,753]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12D</figureCitation>
), emarginated at apical margin, with 6 setae. Pronotum flattened, trapezoidal. Elytra shorter than wide. Hind wings absent. Abdomen oval. Tarsal formula 4-4-5.
</paragraph>
<paragraph id="8BE836C04255FFE382FA39134C5BFC6B" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
Male; tergite VIII with apical margin truncate; sternite VIII with apical margin slightly prolonged posteriorly; apical lobe of median lobe short, slightly pointed in lateral view (
<figureCitation id="136C2A454255FFE3814439374DC4FC23" box="[889,991,871,897]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12E</figureCitation>
); internal structure of median lobe with apical sclerites rounded at apex (
<figureCitation id="136C2A454255FFE3802D39DB4C63FC07" box="[528,632,907,933]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12E</figureCitation>
); suspensonia spoon-shaped, broad in apical half in ventral view (
<figureCitation id="136C2A454255FFE3875239DB4EDDFC6B" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12F</figureCitation>
); apical lobe of paramere slender.
</paragraph>
<paragraph id="8BE836C04255FFE382FA39834C80FBB3" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
Female; tergite VIII apical margin truncate; sternite VIII apical margin rounded; spermatheca short, apical part circular, twice curved around base (
<figureCitation id="136C2A454255FFE3801B39A74C94FBB3" box="[550,655,1015,1041]" captionStart="FIGURE 12" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[189,1398,191,1909]" captionTargetId="figure-19@19.[180,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 12 Diaulota orientalis sp. n. A, habitus, male (holotype); B, ditto, female (paratype); C, labium; D, mentum; E, median lode of aedeagus, lateral view; F, internal structure of median lobe, ventral view; G, spermatheca. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for CG." figureDoi="http://doi.org/10.5281/zenodo.10361610" httpUri="https://zenodo.org/record/10361610/files/figure.png" pageId="18" pageNumber="269">Fig. 12G</figureCitation>
).
</paragraph>
<paragraph id="8BE836C04255FFE382FA3E4B4B95FBDF" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3E4B4F65FB97" bold="true" box="[199,382,1051,1077]" pageId="18" pageNumber="269">Measurements.</emphasis>
Male (N = 7): BL,
<quantity id="4CAF9B254255FFE3805C3E4A4D13FB97" box="[609,776,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.675" metricValueMax="2.8" metricValueMin="2.55" pageId="18" pageNumber="269" unit="mm" value="2.675" valueMax="2.8" valueMin="2.55">2.552.80 mm</quantity>
; FBL,
<quantity id="4CAF9B254255FFE381653E4B4DE4FB97" box="[856,1023,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.245" metricValueMax="1.31" metricValueMin="1.18" pageId="18" pageNumber="269" unit="mm" value="1.245" valueMax="1.31" valueMin="1.18">1.181.31 mm</quantity>
; HW,
<quantity id="4CAF9B254255FFE386753E4B4AF4FB97" box="[1096,1263,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.7" metricValueMin="6.0" pageId="18" pageNumber="269" unit="mm" value="0.635" valueMax="0.67" valueMin="0.6">0.600.67 mm</quantity>
; PL,
<quantity id="4CAF9B254255FFE387113E4A4EDDFBFB" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.4" metricValueMin="4.699999999999999" pageId="18" pageNumber="269" unit="mm" value="0.505" valueMax="0.54" valueMin="0.47">0.470.54 mm</quantity>
; PW,
<quantity id="4CAF9B254255FFE3833B3E6E4FB0FBFB" box="[262,427,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="18" pageNumber="269" unit="mm" value="0.535" valueMax="0.56" valueMin="0.51">0.510.56 mm</quantity>
; EW,
<quantity id="4CAF9B254255FFE383D13E6E4C8AFBFB" box="[492,657,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="18" pageNumber="269" unit="mm" value="0.535" valueMax="0.56" valueMin="0.51">0.510.56 mm</quantity>
; HW/PW, 1.131.21. Female (N = 7): BL,
<quantity id="4CAF9B254255FFE386533E6E4B08FBFB" box="[1134,1299,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="2.52" metricValueMax="2.75" metricValueMin="2.29" pageId="18" pageNumber="269" unit="mm" value="2.52" valueMax="2.75" valueMin="2.29">2.292.75 mm</quantity>
; FBL, 1.11 1.21. mm; HW,
<quantity id="4CAF9B254255FFE383763E324FEAFBDF" box="[331,497,1122,1149]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="18" pageNumber="269" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.450.48 mm</quantity>
; PL,
<quantity id="4CAF9B254255FFE380163E334CCAFBDF" box="[555,721,1123,1149]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="18" pageNumber="269" unit="mm" value="0.45999999999999996" valueMax="0.48" valueMin="0.44">0.440.48 mm</quantity>
; PW,
<quantity id="4CAF9B254255FFE3812C3E334DACFBDF" box="[785,951,1123,1149]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="18" pageNumber="269" unit="mm" value="0.47" valueMax="0.48" valueMin="0.46">0.460.48 mm</quantity>
; EW,
<quantity id="4CAF9B254255FFE381C73E324ABBFBDF" box="[1018,1184,1122,1149]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="18" pageNumber="269" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.450.47 mm</quantity>
; HW/PW, 0.941.03.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4255FFE382FA3ED74FD0FA93" pageId="18" pageNumber="269" type="etymology">
<paragraph id="8BE836C04255FFE382FA3ED74FA6FB67" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3ED74F55FB03" bold="true" box="[199,334,1159,1185]" pageId="18" pageNumber="269">Etymology.</emphasis>
The Latin
<taxonomicName id="4C574D434255FFE383F03ED74C5FFB03" authorityName="Tasaku, Ono &amp; Maruyama" authorityYear="2023" box="[461,580,1159,1185]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B923EAD24255FFE383E43ED74C5FFB03" box="[473,580,1159,1185]" italics="true" pageId="18" pageNumber="269">orientalis</emphasis>
</taxonomicName>
” means eastern in referring to its eastern distribution of the
<taxonomicName id="4C574D434255FFE386CF3ED74B87FB02" authorityName="K. Sawada" authorityYear="1955" box="[1266,1436,1159,1185]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24255FFE386CF3ED74B87FB02" box="[1266,1436,1159,1185]" italics="true" pageId="18" pageNumber="269">Diaulota uenoi</emphasis>
</taxonomicName>
species-complex in
<collectingCountry id="F34076504255FFE3834B3EFB4FA2FB67" box="[374,441,1195,1221]" name="Japan" pageId="18" pageNumber="269">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BE836C04255FFE382FA3E9E4FD0FA93" blockId="18.[151,1443,150,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3E9E4F31FB4A" bold="true" box="[199,298,1230,1256]" pageId="18" pageNumber="269">Habitat.</emphasis>
This species is found from the crevices of rocks and occasionally inside of the dead barnacles (
<taxonomicName id="4C574D434255FFE3870D3E9F4F71FAAF" authority="(Pilsbry))" authorityName="" baseAuthorityName="Pilsbry" baseAuthorityYear="1916" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclita" kingdom="Animalia" order="Sessilia" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B923EAD24255FFE3870D3E9F4EE2FAAE" italics="true" pageId="18" pageNumber="269">Tetraclita japonica</emphasis>
(Pilsbry))
</taxonomicName>
on the intertidal zones. They usually occur lower than where
<taxonomicName id="4C574D434255FFE386323EA34A6AFAAF" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[1039,1137,1267,1293]" class="Insecta" family="Staphylinidae" genus="Rotundicephala" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="koheii" status="sp. nov.">
<emphasis id="B923EAD24255FFE386323EA34A6AFAAF" box="[1039,1137,1267,1293]" italics="true" pageId="18" pageNumber="269">R. koheii</emphasis>
</taxonomicName>
<emphasis id="B923EAD24255FFE3864A3EA34AAEFAAF" bold="true" box="[1143,1205,1267,1293]" pageId="18" pageNumber="269">
<taxonomicNameLabel id="A21057A94255FFE3864A3EA34AAEFAAF" box="[1143,1205,1267,1293]" pageId="18" pageNumber="269" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are found, its habitat is exposed to strong waves.
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4255FFE382FA3F6A4DA0FAF7" box="[199,955,1338,1365]" pageId="18" pageNumber="269" type="distribution">
<paragraph id="8BE836C04255FFE382FA3F6A4DA0FAF7" blockId="18.[151,1443,150,1365]" box="[199,955,1338,1365]" pageId="18" pageNumber="269">
<emphasis id="B923EAD24255FFE382FA3F6A4F7AFAF6" bold="true" box="[199,353,1338,1364]" pageId="18" pageNumber="269">Distribution.</emphasis>
<collectingCountry id="F34076504255FFE3835A3F6B4FB0FAF7" box="[359,427,1339,1365]" name="Japan" pageId="18" pageNumber="269">Japan</collectingCountry>
: central Honshû (Pacific Ocean side) (
<figureCitation id="136C2A454255FFE381673F6B4DB5FAF7" box="[858,942,1339,1365]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="18" pageNumber="269">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>