326 lines
48 KiB
XML
326 lines
48 KiB
XML
<document id="4F81AE2B7B455AF525F19A3ADF5D3475" ID-CLB-Dataset="22330" ID-DOI="10.11646/zootaxa.4772.1.5" ID-GBIF-Dataset="b14b0d0d-cf67-4895-965c-12792c622b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3814079" ID-ZooBank="B497198A-08CD-4A21-AE04-14390499853B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834845818" checkinUser="plazi" docAuthor="Polhemus, Dan A." docDate="2020" docId="03FE00115B6FFFC1FF38F957FE1809FE" docLanguage="en" docName="zootaxa.4772.1.5.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enithares tagula Polhemus 2020, new species" docType="treatment" docVersion="7" lastPageNumber="151" masterDocId="FFC778695B7DFFD2FFAFFFC8FFF90E21" masterDocTitle="Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist" masterLastPageNumber="182" masterPageNumber="132" pageNumber="150" updateTime="1698838866946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B204B48DE65FC65FCF855D6D8A2BD030">
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<mods:title id="00E53407947E2CC025692B9BB61F2D1B">Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist</mods:title>
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<mods:name id="2DB682E1C58D66B338367F4C732CCE07" type="personal">
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<mods:roleTerm id="35329F7FB144B973FEC1284B16BD9963">Author</mods:roleTerm>
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<mods:namePart id="6F8481933996270467BE60384E97D1EF">Polhemus, Dan A.</mods:namePart>
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<mods:typeOfResource id="A7DEDEDB2C9C49129439E8D08D535633">text</mods:typeOfResource>
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<mods:title id="4F551E9D3389BB03EE0BB44638BB5DB0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="21906101D10854241D23B514BFB2D09F">
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<mods:date id="C03388896516E9DED584B3310C98FDFE">2020</mods:date>
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<mods:detail id="EAE4394EEC7D91BFD048F0E7EC25D81A" type="pubDate">
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<mods:number id="D0F0DC8F5BCB2ACCAFC18D6BC0C0D8E8">2020-05-07</mods:number>
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<mods:detail id="32A5CFEB4CE0674DC4DB78EA9782ADD1" type="volume">
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<mods:number id="2E8524407C1DC5A42656267E56EA0822">4772</mods:number>
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<mods:start id="F5A44D927FB59AB0EC3046883F2BE702">132</mods:start>
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<mods:classification id="940C371D9EB0F17CBA7C964F0BC6AD44">journal article</mods:classification>
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<mods:identifier id="8CDB53DCE98C7FEECAC318730590A41F" type="DOI">10.11646/zootaxa.4772.1.5</mods:identifier>
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<mods:identifier id="1C2760B5915C49E327BF5762571B75EA" type="GBIF-Dataset">b14b0d0d-cf67-4895-965c-12792c622b21</mods:identifier>
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<mods:identifier id="4D76A3466224CA648325E9EED1D1B79E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="47A863C1008B76342C5556B52C0EE625" type="Zenodo-Dep">3814079</mods:identifier>
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</mods:mods>
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<treatment id="03FE00115B6FFFC1FF38F957FE1809FE" ID-DOI="http://doi.org/10.5281/zenodo.3815476" ID-GBIF-Taxon="163975555" ID-Zenodo-Dep="3815476" LSID="urn:lsid:plazi:treatment:03FE00115B6FFFC1FF38F957FE1809FE" httpUri="http://treatment.plazi.org/id/03FE00115B6FFFC1FF38F957FE1809FE" lastPageId="19" lastPageNumber="151" pageId="18" pageNumber="150">
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<subSubSection id="C34DE28C5B6FFFC0FF38F957FE000898" box="[151,505,1695,1722]" pageId="18" pageNumber="150" type="nomenclature">
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<paragraph id="8BE8B1075B6FFFC0FF38F957FE000898" blockId="18.[151,505,1695,1758]" box="[151,505,1695,1722]" pageId="18" pageNumber="150">
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<heading id="D0A0066B5B6FFFC0FF38F957FE000898" bold="true" box="[151,505,1695,1722]" fontSize="11" level="1" pageId="18" pageNumber="150" reason="1">
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<emphasis id="B9236D155B6FFFC0FF38F957FE000898" bold="true" box="[151,505,1695,1722]" pageId="18" pageNumber="150">
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<taxonomicName id="4C57CA845B6FFFC0FF38F957FE9A089B" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[151,355,1695,1722]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="150" phylum="Arthropoda" rank="species" species="tagula" status="sp. nov.">
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<emphasis id="B9236D155B6FFFC0FF38F957FE9A089B" bold="true" box="[151,355,1695,1722]" italics="true" pageId="18" pageNumber="150">Enithares tagula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A210D06E5B6FFFC0FEC6F968FE000898" box="[361,505,1695,1722]" pageId="18" pageNumber="150" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34DE28C5B6FFFC0FF38F90BFE7B08FF" box="[151,386,1731,1758]" pageId="18" pageNumber="150" type="description">
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<paragraph id="8BE8B1075B6FFFC0FF38F90BFE7B08FF" blockId="18.[151,505,1695,1758]" box="[151,386,1731,1758]" pageId="18" pageNumber="150">
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(
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<figureCitation id="136CAD825B6FFFC0FF30F90BFEDE08FF" box="[159,295,1731,1758]" captionStart="FIGURES 31–34" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 31–34. Enithares species in the E. atra group, dorsal habitus photographs. 31–32. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 33–34. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="18" pageNumber="150">Figs. 33, 34</figureCitation>
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,
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<figureCitation id="136CAD825B6FFFC0FE9CF90CFEA808FF" box="[307,337,1732,1758]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="18" pageNumber="150">38</figureCitation>
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,
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<figureCitation id="136CAD825B6FFFC0FEF2F90BFE8308FF" box="[349,378,1731,1758]" captionStart="FIGURE 55" captionStartId="32.[151,250,1007,1032]" captionTargetBox="[157,1425,189,973]" captionTargetId="figure@32.[151,1436,181,982]" captionTargetPageId="32" captionText="FIGURE 55. Distribution of species in the E. atra group on New Guinea. Square = E. digitata Lansbury; circles = E. elongata Lansbury; triangles = E. insularis n. sp.; pentagon = E. tagula n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814105" httpUri="https://zenodo.org/record/3814105/files/figure.png" pageId="18" pageNumber="150">55</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34DE28C5B6FFFC0FF38F8CEFD3809D8" pageId="18" pageNumber="150" type="materials_examined">
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<paragraph id="8BE8B1075B6FFFC0FF38F8CEFD3809D8" blockId="18.[151,1436,1797,2041]" pageId="18" pageNumber="150">
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<emphasis id="B9236D155B6FFFC0FF38F8CEFE3B0901" bold="true" box="[151,450,1797,1824]" pageId="18" pageNumber="150">Type material examined</emphasis>
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.
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<materialsCitation id="3B3FBB5A5B6FFFC0FE7AF8CEFECA09AD" ID-GBIF-Occurrence="2610672481" collectingDate="2002-08-29" collectionCode="A, USNM" country="Papua New Guinea" county="Louisiade Archipelago" elevation="15" latitude="-11.364445" location="Island" longLatPrecision="21" longitude="153.24055" municipality="Tagula" pageId="18" pageNumber="150" specimenCount="1" stateProvince="Milne Bay" typeStatus="holotype">
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<typeStatus id="54EC0FA55B6FFFC0FE7AF8CEFDB90901" box="[469,576,1798,1824]" pageId="18" pageNumber="150" type="holotype">Holotype</typeStatus>
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, male (dissected):
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<collectingCountry id="F340F1975B6FFFC0FC8FF8CEFBC40901" box="[800,1085,1798,1825]" name="Papua New Guinea" pageId="18" pageNumber="150">PAPUA NEW GUINEA</collectingCountry>
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,
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<collectingRegion id="49937FE55B6FFFC0FBE1F8CDFAE30900" box="[1102,1306,1797,1825]" country="Papua New Guinea" name="Milne Bay" pageId="18" pageNumber="150">
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<emphasis id="B9236D155B6FFFC0FBE1F8CDFAED0901" bold="true" box="[1102,1300,1797,1824]" pageId="18" pageNumber="150">Milne Bay Prov</emphasis>
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.
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</collectingRegion>
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,
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<collectingCounty id="6289C98B5B6FFFC0FA81F8CEFEDD0965" pageId="18" pageNumber="150">Louisiade Archipelago</collectingCounty>
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,
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<collectingMunicipality id="6B8C2B7D5B6FFFC0FE9DF8E2FE870965" box="[306,382,1834,1860]" pageId="18" pageNumber="150">Tagula</collectingMunicipality>
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[Sudest]
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<location id="8E88E7DC5B6FFFC0FE40F8E2FDCF0965" LSID="urn:lsid:plazi:treatment:03FE00115B6FFFC1FF38F957FE1809FE:8E88E7DC5B6FFFC0FE40F8E2FDCF0965" box="[495,566,1834,1860]" country="Papua New Guinea" county="Louisiade Archipelago" latitude="-11.364445" longLatPrecision="21" longitude="153.24055" municipality="Tagula" name="Island" pageId="18" pageNumber="150" stateProvince="Milne Bay">Island</location>
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,
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<location id="8E88E7DC5B6FFFC0FDEBF8E2FD1E0965" LSID="urn:lsid:plazi:treatment:03FE00115B6FFFC1FF38F957FE1809FE:8E88E7DC5B6FFFC0FDEBF8E2FD1E0965" box="[580,743,1834,1860]" country="Papua New Guinea" county="Louisiade Archipelago" latitude="-11.364445" longLatPrecision="21" longitude="153.24055" municipality="Tagula" name="Kalitau Creek" pageId="18" pageNumber="150" stateProvince="Milne Bay">Kalitau Creek</location>
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,
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<quantity id="4CAF1CE25B6FFFC0FD5BF8E2FCCA0965" box="[756,819,1834,1860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="150" unit="km" value="1.0">1 km</quantity>
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. upstream from head of estuary,
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<quantity id="4CAF1CE25B6FFFC0FB1DF8E2FAF40964" box="[1202,1293,1834,1861]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="18" pageNumber="150" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">
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<elevation id="007A56345B6FFFC0FB1DF8E2FAF40964" box="[1202,1293,1834,1861]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="18" pageNumber="150" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">0–30 m</elevation>
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</quantity>
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. [
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<quantity id="4CAF1CE25B6FFFC0FA8BF8E2FA760965" box="[1316,1423,1833,1860]" metricMagnitude="1" metricUnit="m" metricValue="1.524" metricValueMax="3.048" metricValueMin="0.0" pageId="18" pageNumber="150" unit="ft" value="50.0" valueMax="100.0" valueMin="0.0">
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<elevation id="007A56345B6FFFC0FA8BF8E2FA760965" box="[1316,1423,1833,1860]" metricMagnitude="1" metricUnit="m" metricValue="1.524" metricValueMax="3.048" metricValueMin="0.0" pageId="18" pageNumber="150" unit="ft" value="50.0" valueMax="100.0" valueMin="0.0">0–100 ft.</elevation>
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</quantity>
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],
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<geoCoordinate id="EE63D7C05B6FFFC0FF38F885FEE30949" box="[151,282,1869,1896]" degrees="11" direction="south" minutes="21" orientation="latitude" pageId="18" pageNumber="150" precision="15" seconds="52" value="-11.364445">11°21′52′′S</geoCoordinate>
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,
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<geoCoordinate id="EE63D7C05B6FFFC0FE8AF885FE410949" box="[293,440,1869,1896]" degrees="153" direction="east" minutes="14" orientation="longitude" pageId="18" pageNumber="150" precision="15" seconds="26" value="153.24055">153°14′26′′E</geoCoordinate>
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, water temp. 24.4° C.,
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<date id="FFE997C75B6FFFC0FD18F886FC930949" box="[695,874,1870,1896]" pageId="18" pageNumber="150" value="2002-08-29">
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<collectingDate id="EFAD6E2F5B6FFFC0FD18F886FC930949" box="[695,874,1870,1896]" pageId="18" pageNumber="150" value="2002-08-29">29 August 2002</collectingDate>
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</date>
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, 09:30–14:30 hrs., CL 7190, D.
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<collectionCode id="ED4629C25B6FFFC0FB61F886FB1C0949" box="[1230,1253,1870,1896]" pageId="18" pageNumber="150">A</collectionCode>
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. and J. T. Polhe- mus (
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<collectionCode id="ED4629C25B6FFFC0FF7BF8BAFED209AD" box="[212,299,1906,1932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="150" type="Museum">USNM</collectionCode>
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)
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</materialsCitation>
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||
.
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<materialsCitation id="3B3FBB5A5B6FFFC0FE92F8BBFB3809AD" ID-GBIF-Occurrence="2610672596" box="[317,1217,1906,1933]" collectingDate="2002-08-29" collectionCode="A" country="Papua New Guinea" county="Louisiade Archipelago" elevation="15" latitude="-11.364445" location="Island" longLatPrecision="21" longitude="153.24055" municipality="Tagula" pageId="18" pageNumber="150" specimenCount="4" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="2" stateProvince="Milne Bay" typeStatus="paratype">
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<emphasis id="B9236D155B6FFFC0FE92F8BBFE5609AD" box="[317,431,1907,1932]" italics="true" pageId="18" pageNumber="150">
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||
<typeStatus id="54EC0FA55B6FFFC0FE92F8BBFE5609AD" box="[317,431,1907,1932]" pageId="18" pageNumber="150" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
:
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||
<specimenCount id="9D517A8E5B6FFFC0FE12F8BAFDEC09AD" box="[445,533,1906,1932]" pageId="18" pageNumber="150" type="male">2 males</specimenCount>
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||
,
|
||
<specimenCount id="9D517A8E5B6FFFC0FDB0F8BAFD7A09AD" box="[543,643,1906,1932]" pageId="18" pageNumber="150" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="9D517A8E5B6FFFC0FD22F8BAFCF609AD" box="[653,783,1906,1932]" pageId="18" pageNumber="150" type="immature">1 immature</specimenCount>
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||
, same data as holotype, CL 7190, D.
|
||
<collectionCode id="ED4629C25B6FFFC0FB05F8BAFB3809AD" box="[1194,1217,1906,1932]" pageId="18" pageNumber="150">A</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3FBB5A5B6FFFC0FB64F8BAFEA30990" ID-GBIF-Occurrence="2610672679" collectionCode="USNM, BPBM" collectorName="Polhemus" country="Papua New Guinea" pageId="18" pageNumber="150" specimenCount="1" stateProvince="Milne Bay" typeStatus="paratype">
|
||
and J. T.
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||
<collectorName id="26A2D4D15B6FFFC0FA81F8BAFA6509AD" box="[1326,1436,1906,1932]" pageId="18" pageNumber="150">Polhemus</collectorName>
|
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(
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||
<collectionCode id="ED4629C25B6FFFC0FF30F85EFF0D0991" box="[159,244,1942,1968]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="150" type="Museum">USNM</collectionCode>
|
||
,
|
||
<collectionCode id="ED4629C25B6FFFC0FEAFF85FFEAA0990" box="[256,339,1943,1969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="150" type="Museum">BPBM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3FBB5A5B6FFFC0FEC9F85DFE8309F4" ID-GBIF-Occurrence="2610672410" collectorName="Louisiade Archipelago & Tagula & Sudest" country="Papua New Guinea" elevation="45" location="Island" municipality="Kalitau Creek" pageId="18" pageNumber="150" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Milne Bay" typeStatus="paratype">
|
||
<specimenCount id="9D517A8E5B6FFFC0FEC9F85DFE460991" box="[358,447,1941,1968]" pageId="18" pageNumber="150" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D517A8E5B6FFFC0FE64F85EFDC30991" box="[459,570,1942,1968]" pageId="18" pageNumber="150" type="female">2 females</specimenCount>
|
||
,
|
||
<collectorName id="26A2D4D15B6FFFC0FDEAF85EFCBF0991" box="[581,838,1942,1968]" pageId="18" pageNumber="150">Louisiade Archipelago</collectorName>
|
||
,
|
||
<collectorName id="26A2D4D15B6FFFC0FCFDF85EFC670991" box="[850,926,1942,1968]" pageId="18" pageNumber="150">Tagula</collectorName>
|
||
[
|
||
<collectorName id="26A2D4D15B6FFFC0FC04F85DFC020991" box="[939,1019,1941,1968]" pageId="18" pageNumber="150">Sudest</collectorName>
|
||
]
|
||
<location id="8E88E7DC5B6FFFC0FBA6F85EFBA90991" LSID="urn:lsid:plazi:treatment:03FE00115B6FFFC1FF38F957FE1809FE:8E88E7DC5B6FFFC0FBA6F85EFBA90991" box="[1033,1104,1942,1968]" country="Papua New Guinea" municipality="Kalitau Creek" name="Island" pageId="18" pageNumber="150" stateProvince="Milne Bay">Island</location>
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, small spring-fed stream near
|
||
<collectingMunicipality id="6B8C2B7D5B6FFFC0FF38F872FECF09F5" box="[151,310,1978,2004]" pageId="18" pageNumber="150">Kalitau Creek</collectingMunicipality>
|
||
,
|
||
<quantity id="4CAF1CE25B6FFFC0FE90F871FE8309F4" box="[319,378,1977,2005]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="150" unit="m" value="45.0">
|
||
<specimenCount id="9D517A8E5B6FFFC0FE90F871FE8309F4" box="[319,378,1977,2005]" pageId="18" pageNumber="150" type="generic">
|
||
<elevation id="007A56345B6FFFC0FE90F871FE8309F4" box="[319,378,1977,2005]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="150" unit="m" value="45.0">45 m</elevation>
|
||
</specimenCount>
|
||
</quantity>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B3FBB5A5B6FFFC0FED5F873FEEC09D9" ID-GBIF-Occurrence="2610672364" collectingDate="2002-08-29" collectionCode="A" country="Papua New Guinea" elevation="46" latitude="-11.358334" longLatPrecision="21" longitude="153.24445" pageId="18" pageNumber="150" specimenCount="1" stateProvince="Milne Bay" typeStatus="paratype">
|
||
. [
|
||
<quantity id="4CAF1CE25B6FFFC0FE24F871FE2009F5" box="[395,473,1977,2004]" metricMagnitude="1" metricUnit="m" metricValue="4.572" pageId="18" pageNumber="150" unit="ft" value="150.0">
|
||
<elevation id="007A56345B6FFFC0FE24F871FE2009F5" box="[395,473,1977,2004]" metricMagnitude="1" metricUnit="m" metricValue="4.572" pageId="18" pageNumber="150" unit="ft" value="150.0">150 ft.</elevation>
|
||
</quantity>
|
||
], water temp. 26° C.,
|
||
<date id="FFE997C75B6FFFC0FD66F872FC8209F5" box="[713,891,1978,2004]" pageId="18" pageNumber="150" value="2002-08-29">
|
||
<collectingDate id="EFAD6E2F5B6FFFC0FD66F872FC8209F5" box="[713,891,1978,2004]" pageId="18" pageNumber="150" value="2002-08-29">29 August 2002</collectingDate>
|
||
</date>
|
||
, 10:00–12:00 hrs.,
|
||
<geoCoordinate id="EE63D7C05B6FFFC0FBE0F871FB2809F5" box="[1103,1233,1977,2004]" degrees="11" direction="south" minutes="21" orientation="latitude" pageId="18" pageNumber="150" precision="15" seconds="30" value="-11.358334">11°21′30′′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE63D7C05B6FFFC0FB73F871FA9609F5" box="[1244,1391,1977,2004]" degrees="153" direction="east" minutes="14" orientation="longitude" pageId="18" pageNumber="150" precision="15" seconds="40" value="153.24445">153°14′40′′E</geoCoordinate>
|
||
, CL 7192, D.
|
||
<collectionCode id="ED4629C25B6FFFC0FF51F816FEEC09D9" box="[254,277,2014,2040]" pageId="18" pageNumber="150">A</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3FBB5A5B6FFFC0FE8FF816FD4409D8" ID-GBIF-Occurrence="2610672648" box="[288,701,2014,2041]" collectionCode="USNM, BPBM" collectorName="Polhemus" country="Papua New Guinea" pageId="18" pageNumber="150" specimenCount="1" stateProvince="Milne Bay" typeStatus="paratype">
|
||
and J. T.
|
||
<collectorName id="26A2D4D15B6FFFC0FE2AF816FE0A09D9" box="[389,499,2014,2040]" pageId="18" pageNumber="150">Polhemus</collectorName>
|
||
(
|
||
<collectionCode id="ED4629C25B6FFFC0FDADF816FDAE09D9" box="[514,599,2014,2040]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="150" type="Museum">USNM</collectionCode>
|
||
,
|
||
<collectionCode id="ED4629C25B6FFFC0FDCCF817FD4F09D8" box="[611,694,2015,2041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="150" type="Museum">BPBM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B6EFFC1FF68FF50FD430ACA" pageId="19" pageNumber="151" type="description">
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FF50FC1F0E92" blockId="19.[151,1437,152,2015]" box="[199,998,152,179]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FF50FEA80E92" bold="true" box="[199,337,152,179]" pageId="19" pageNumber="151">Description</emphasis>
|
||
.
|
||
<emphasis id="B9236D155B6EFFC1FECFFF51FE640E92" bold="true" box="[352,413,153,179]" pageId="19" pageNumber="151">Male</emphasis>
|
||
: Length
|
||
<quantity id="4CAF1CE25B6EFFC1FDAEFF51FD9E0E92" box="[513,615,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="19" pageNumber="151" unit="mm" value="8.9">8.90 mm</quantity>
|
||
, width across pronotum
|
||
<quantity id="4CAF1CE25B6EFFC1FCD4FF51FC1B0E92" box="[891,994,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="19" pageNumber="151" unit="mm" value="3.4">3.40 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FF74FDAF0FD7" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FF74FEBB0EF7" box="[199,322,188,214]" italics="true" pageId="19" pageNumber="151">Coloration</emphasis>
|
||
: Ground color dark brown to black, with scattered pale brown markings on pronotum and hemelytra (
|
||
<figureCitation id="136CAD825B6EFFC1FF0FFF28FF0D0EDA" box="[160,244,224,251]" captionStart="FIGURES 31–34" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 31–34. Enithares species in the E. atra group, dorsal habitus photographs. 31–32. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 33–34. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="19" pageNumber="151">Fig. 33</figureCitation>
|
||
). Vertex and frons orange-brown, with an irregular dark brown patch at apex of head flanked by reddish; irregular dark brown, transverse, inverted V-shaped fascia present on lower frons, isolating a roughly triangular dark yellow patch above base of labrum. Pronotum yellowish-brown on anterior two-thirds between and surrounding pronotal fovea, greyish-brown along posterior margin. Scutellum dark brown to black, with central section embrowned. Hemelytra greyish-brown, with inner margins of clavus, outer base of clavus, outer and posterior corium with large, irregular tan patches; wing membrane black. Legs with coxae dark yellowish, all other segments medium brown, anterior faces of fore femora broadly suffused with black; ventral surfaces of trochanters and femora marked with dark reddish brown. Venter brown.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FDC8FDD90CEE" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FDC8FE4E0C3B" box="[199,439,512,538]" italics="true" pageId="19" pageNumber="151">Structural characters</emphasis>
|
||
: Head broadly rounded anteriorly when viewed dorsally. Head length 0.75; greatest width 2.75, equal to 0.81 pronotal width; anterior width of vertex 0.90, equal to 1.20 head length. Synthlipsis 0.60, about 0.67 anterior width of vertex and clearly shorter than pronotum. Pronotal length along midline 1.00, humeral width 3.40, lateral margins convex, posterior margin weakly sinuate. Dorsal margin of pronotal fovea directed caudad behind eyes (
|
||
<figureCitation id="136CAD825B6EFFC1FEBDFD58FE9F0C8A" box="[274,358,656,683]" captionStart="FIGURES 31–34" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 31–34. Enithares species in the E. atra group, dorsal habitus photographs. 31–32. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 33–34. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="19" pageNumber="151">Fig. 33</figureCitation>
|
||
). Nodal furrow nearly straight, removed by 1.6× its length from membranal suture, length 0.50, distance to membranal suture 0.80.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FD11FE680D86" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">Fore tibia with thick, dense pad of short golden setae along anteroventral margin, covering anterior half of ventral face of tibia. Middle trochanter rounded. Middle femur with single moderately large, elongate, sharp subapical tooth, bordered basally by about 15 small black pegs, distally by about 3 similar pegs. Hind legs typical for genus, lacking unusual modifications. Lengths of leg segments as follows: fore femur–tibia–tarsal 1–tarsal 2 = 1.10/1.20/0.60/0.30; middle femur–tibia–tarsal 1–tarsal 2 = 2.30/2.00/0.70/0.40; hind femur–tibia–tarsal 1–tarsal 2 = 3.50/3.00/1.35/0.70.</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FC78FBCB0DEB" blockId="19.[151,1437,152,2015]" box="[199,1074,944,971]" pageId="19" pageNumber="151">Ventral abdomen with metaxyphus triangular, slightly concave, tip acuminate.</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FC1CFC240A83" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FC1CFE940DCF" box="[199,365,980,1006]" italics="true" pageId="19" pageNumber="151">Male genitalia</emphasis>
|
||
when viewed laterally (
|
||
<figureCitation id="136CAD825B6EFFC1FDD1FC1CFD280DCE" box="[638,721,980,1007]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="19" pageNumber="151">Fig. 38</figureCitation>
|
||
) with ventral margin of proctiger broadly rounded, apex acute; posterior lobe of pygophore with posteroventral angle slightly obtuse, sharp, bearing a small setal tuft, posterior lobe of pygophore erect, apex truncate, posteriorly setiferous, not notched anteriorly on dorsal margin; paramere elongate, slightly tapering and curved anteriorly, posterior margin straight, apex rounded, reaching to dorsal margin of posterior lobe; lateral arm of basal plate broadly sinuate, of relatively even width throughout, apex rounded (
|
||
<figureCitation id="136CAD825B6EFFC1FAC0FBACFF4F0A82" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="19" pageNumber="151">Fig. 38</figureCitation>
|
||
); aedeagus semicircular in lateral view, with numerous transverse folds.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FB65FD430ACA" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FB65FEE70AE6" bold="true" box="[199,286,1197,1223]" pageId="19" pageNumber="151">Female</emphasis>
|
||
: Length 9.50, width across pronotum
|
||
<quantity id="4CAF1CE25B6EFFC1FD67FB65FCD60AE6" box="[712,815,1197,1223]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="151" unit="mm" value="3.7">3.70 mm</quantity>
|
||
. Similar to male in general structure and coloration, but slightly larger and more robust in form (
|
||
<figureCitation id="136CAD825B6EFFC1FDF6FB18FD570ACA" box="[601,686,1232,1259]" captionStart="FIGURES 31–34" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 31–34. Enithares species in the E. atra group, dorsal habitus photographs. 31–32. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 33–34. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="19" pageNumber="151">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B6EFFC1FF68FB3DFE640B13" pageId="19" pageNumber="151" type="etymology">
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FB3DFE640B13" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FB3DFEB10B2E" bold="true" box="[199,328,1269,1295]" pageId="19" pageNumber="151">Etymology</emphasis>
|
||
. The name “tagula” is a noun in apposition, and refers to the
|
||
<typeStatus id="54EC0FA55B6EFFC1FBACFB3DFBCA0B2E" box="[1027,1075,1269,1295]" pageId="19" pageNumber="151">type</typeStatus>
|
||
locality of Tagula Island, in the Louisiade Archipelago.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B6EFFC1FF68FAF4FE1809FE" pageId="19" pageNumber="151" type="discussion">
|
||
<paragraph id="8BE8B1075B6EFFC1FF68FAF4FED808BB" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68FAF4FEBC0B76" bold="true" box="[199,325,1340,1367]" pageId="19" pageNumber="151">Discussion</emphasis>
|
||
.
|
||
<taxonomicName id="4C57CA845B6EFFC1FEF9FAF4FDEA0B77" authorityName="Polhemus" authorityYear="2020" box="[342,531,1340,1366]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FEF9FAF4FDEA0B77" box="[342,531,1340,1366]" italics="true" pageId="19" pageNumber="151">Enithares tagula</emphasis>
|
||
</taxonomicName>
|
||
is similar in general size and coloration to
|
||
<taxonomicName id="4C57CA845B6EFFC1FBA2FAF5FBA70B77" box="[1037,1118,1341,1366]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9236D155B6EFFC1FBA2FAF5FBA70B77" box="[1037,1118,1341,1366]" italics="true" pageId="19" pageNumber="151">E. atra</emphasis>
|
||
</taxonomicName>
|
||
from the Papuan Peninsula and
|
||
<taxonomicName id="4C57CA845B6EFFC1FF68FAA9FEB00B5B" authorityName="Polhemus" authorityYear="2020" box="[199,329,1376,1402]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B9236D155B6EFFC1FF68FAA9FEB00B5B" box="[199,329,1376,1402]" italics="true" pageId="19" pageNumber="151">E. insularis</emphasis>
|
||
</taxonomicName>
|
||
from the D’Entrecasteaux Islands, but may be differentiated by aspects of the male genitalia. The posteroventral angle of the pygophore is far more sharply defined in
|
||
<taxonomicName id="4C57CA845B6EFFC1FC3EFA4DFC0E0BBF" authorityName="Polhemus" authorityYear="2020" box="[913,1015,1412,1438]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FC3EFA4DFC0E0BBF" box="[913,1015,1412,1438]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
than in either of the other two species (compare
|
||
<figureCitation id="136CAD825B6EFFC1FEA7FA60FE690BE2" box="[264,400,1448,1475]" captionStart="FIGURES 16–21" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 16–21. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="19" pageNumber="151">Figs. 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="136CAD825B6EFFC1FE33FA61FE420BE2" box="[412,443,1449,1475]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="19" pageNumber="151">38</figureCitation>
|
||
), and the dorsal margin of the ventral lobe of the proctiger is more transversely truncate (compare
|
||
<figureCitation id="136CAD825B6EFFC1FEA6FA04FE6B0BC6" box="[265,402,1484,1511]" captionStart="FIGURES 16–21" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 16–21. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="19" pageNumber="151">Figs. 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="136CAD825B6EFFC1FE30FA05FE460BC6" box="[415,447,1485,1511]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="19" pageNumber="151">38</figureCitation>
|
||
). The most pronounced difference lies in the structure of the LABP, which in
|
||
<taxonomicName id="4C57CA845B6EFFC1FA9BFA05FA640BC7" authorityName="Polhemus" authorityYear="2020" box="[1332,1437,1484,1510]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FA9BFA05FA640BC7" box="[1332,1437,1484,1510]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
is gently sinuate in form when viewed laterally and of relatively even width throughout (
|
||
<figureCitation id="136CAD825B6EFFC1FBD8FA38FB33082A" box="[1143,1226,1520,1547]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="19" pageNumber="151">Fig. 38</figureCitation>
|
||
), rather than being broadly L-shaped and with a tapering distal arm that widens slightly at the apex, as seen in
|
||
<taxonomicName id="4C57CA845B6EFFC1FB38F9DDFB1C080F" box="[1175,1253,1557,1582]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9236D155B6EFFC1FB38F9DDFB1C080F" box="[1175,1253,1557,1582]" italics="true" pageId="19" pageNumber="151">E. atra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57CA845B6EFFC1FAB4F9DDFA65080F" authorityName="Polhemus" authorityYear="2020" box="[1307,1436,1556,1582]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B9236D155B6EFFC1FAB4F9DDFA65080F" box="[1307,1436,1556,1582]" italics="true" pageId="19" pageNumber="151">E. insularis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136CAD825B6EFFC1FF30F9F0FED10872" box="[159,296,1592,1619]" captionStart="FIGURES 16–21" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 16–21. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="19" pageNumber="151">Figs. 17, 18</figureCitation>
|
||
). In addition to these genitalic characters,
|
||
<taxonomicName id="4C57CA845B6EFFC1FCAEF9F1FC910873" authorityName="Polhemus" authorityYear="2020" box="[769,872,1592,1618]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FCAEF9F1FC910873" box="[769,872,1592,1618]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
has a broad vertex whose anterior width is greater than the length of the head when viewed dorsally, whereas in the other two species the vertex width is less than the head length.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68F96CFBB008C2" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
Both
|
||
<taxonomicName id="4C57CA845B6EFFC1FEABF96DFE92089F" authorityName="Polhemus" authorityYear="2020" box="[260,363,1700,1726]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FEABF96DFE92089F" box="[260,363,1700,1726]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57CA845B6EFFC1FE0EF96DFDD8089F" authorityName="Polhemus" authorityYear="2020" box="[417,545,1700,1726]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B9236D155B6EFFC1FE0EF96DFDD8089F" box="[417,545,1700,1726]" italics="true" pageId="19" pageNumber="151">E. insularis</emphasis>
|
||
</taxonomicName>
|
||
possess well-developed pads of thick, dense setae along the anterior margins of their fore tibiae, although these setae are darker and more promiment in
|
||
<taxonomicName id="4C57CA845B6EFFC1FC6EF901FBBB08C3" authorityName="Polhemus" authorityYear="2020" box="[961,1090,1736,1762]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B9236D155B6EFFC1FC6EF901FBBB08C3" box="[961,1090,1736,1762]" italics="true" pageId="19" pageNumber="151">E. insularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68F924FB0C096F" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<emphasis id="B9236D155B6EFFC1FF68F924FE7F0926" bold="true" box="[199,390,1772,1799]" pageId="19" pageNumber="151">Ecological notes</emphasis>
|
||
. At the Kalitau Creek
|
||
<typeStatus id="54EC0FA55B6EFFC1FDD0F925FD560926" box="[639,687,1773,1799]" pageId="19" pageNumber="151">type</typeStatus>
|
||
locality (CL 7190),
|
||
<taxonomicName id="4C57CA845B6EFFC1FC3FF925FC0E0927" authorityName="Polhemus" authorityYear="2020" box="[912,1015,1772,1798]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FC3FF925FC0E0927" box="[912,1015,1772,1798]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
was taken in side pools amid bedrock along the main stream channel, and from pools along a rocky tributary. At a small, spring-fed stream nearby (CL 7192), this species was collected from scattered standing pools amid boulders in a bedrock channel.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B6EFFC1FF68F890FE1809FE" blockId="19.[151,1437,152,2015]" pageId="19" pageNumber="151">
|
||
<taxonomicName id="4C57CA845B6EFFC1FF68F890FE620953" authorityName="Polhemus" authorityYear="2020" box="[199,411,1880,1906]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B9236D155B6EFFC1FF68F890FE620953" box="[199,411,1880,1906]" italics="true" pageId="19" pageNumber="151">Enithares insularis</emphasis>
|
||
</taxonomicName>
|
||
is apparently endemic to Tagula Island in the Louisiade Archipelago (
|
||
<figureCitation id="136CAD825B6EFFC1FB0AF890FB010952" box="[1189,1272,1880,1907]" captionStart="FIGURE 55" captionStartId="32.[151,250,1007,1032]" captionTargetBox="[157,1425,189,973]" captionTargetId="figure@32.[151,1436,181,982]" captionTargetPageId="32" captionText="FIGURE 55. Distribution of species in the E. atra group on New Guinea. Square = E. digitata Lansbury; circles = E. elongata Lansbury; triangles = E. insularis n. sp.; pentagon = E. tagula n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814105" httpUri="https://zenodo.org/record/3814105/files/figure.png" pageId="19" pageNumber="151">Fig. 55</figureCitation>
|
||
), although it is plausible that it might also occur on the other adjacent high islands in the Louisiades such as Panatinane and Rossel. Based on current records,
|
||
<taxonomicName id="4C57CA845B6EFFC1FE17F869FDE6099B" authorityName="Polhemus" authorityYear="2020" box="[440,543,1952,1978]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="151" phylum="Arthropoda" rank="species" species="tagula">
|
||
<emphasis id="B9236D155B6EFFC1FE17F869FDE6099B" box="[440,543,1952,1978]" italics="true" pageId="19" pageNumber="151">E. tagula</emphasis>
|
||
</taxonomicName>
|
||
occupies the Tagula Island area of freshwater endemism (Area 38) as defined by D.
|
||
<bibRefCitation id="EFC6CCF65B6EFFC1FF16F80CFE2409FE" author="Polhemus, D. A. & Allen, G. A." box="[185,477,1988,2015]" pageId="19" pageNumber="151" pagination="207 - 245" refId="ref39028" refString="Polhemus, D. A. & Allen, G. A. (2007) Freshwater biogeography of Papua. In: Marshall, A. J. & Beehler, B. M. (Eds.), Ecology of Papua. Part 1. Periplus Editions (HK) Ltd., Singapore, pp. 207 - 245." type="book chapter" year="2007">Polhemus & Allen (2007)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |