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<document id="01EB3F3D384777A356B435420A457AF6" ID-CLB-Dataset="157773" ID-DOI="10.11646/zootaxa.5194.1.2" ID-GBIF-Dataset="9c76b088-9845-48f6-9561-d4c6e91c4343" ID-ISSN="1175-5326" ID-Zenodo-Dep="7141848" ID-ZooBank="7C18727C-3AF0-4BE6-AFBC-EA1AC2F2B926" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664862422389" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2022" docId="03EE7A4DC90DFD03FF5EFF41FC7FF823" docLanguage="en" docName="zootaxa.5194.1.2.pdf" docOrigin="Zootaxa 5194 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nodocepheus luxtoni Colloff 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="46" masterDocId="FFD70235C901FD0EFFC9FFD6FF9AFFDF" masterDocTitle="First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)" masterLastPageNumber="57" masterPageNumber="33" pageNumber="45" updateTime="1699387054636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FD6CA8D152A9298D60FF25DC6F99798C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="30B053AA8D8AA7F1D1D6F9C43AD00950">
<mods:title id="F292162658B169488FAA005C7BEDF5F8">First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)</mods:title>
</mods:titleInfo>
<mods:name id="C0836DEE84FFD5202C524F760CAC65A4" type="personal">
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<mods:namePart id="0CDFCC86ADF6D66020551FC633F79502">Colloff, Matthew J.</mods:namePart>
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<mods:title id="52D416EB2EAE1735AFB23557BB8CCD96">Zootaxa</mods:title>
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<mods:part id="1EF8A1D32A5B37AF725F4BCFDCA3746A">
<mods:date id="CF443DF7C244D57698539FEF65B59169">2022</mods:date>
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<mods:number id="EB90FD1BD6A0455BA3A0FF0A7E77F9E0">2022-10-04</mods:number>
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<mods:number id="8E02088741992D6498ED1351543BE5E1">5194</mods:number>
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<mods:identifier id="A5C6F0DE257CAEC3F974560A007B95C2" type="DOI">10.11646/zootaxa.5194.1.2</mods:identifier>
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<treatment id="03EE7A4DC90DFD03FF5EFF41FC7FF823" ID-DOI="http://doi.org/10.5281/zenodo.7141892" ID-GBIF-Taxon="202134808" ID-Zenodo-Dep="7141892" LSID="urn:lsid:plazi:treatment:03EE7A4DC90DFD03FF5EFF41FC7FF823" httpUri="http://treatment.plazi.org/id/03EE7A4DC90DFD03FF5EFF41FC7FF823" lastPageId="13" lastPageNumber="46" pageId="12" pageNumber="45">
<subSubSection id="C35D98D0C90DFD02FF5EFF41FE67FF6E" box="[151,509,151,178]" pageId="12" pageNumber="45" type="nomenclature">
<paragraph id="8BF8CB5BC90DFD02FF5EFF41FE67FF6E" blockId="12.[151,509,151,214]" box="[151,509,151,178]" pageId="12" pageNumber="45">
<heading id="D0B07C37C90DFD02FF5EFF41FE67FF6E" bold="true" box="[151,509,151,178]" fontSize="11" level="1" pageId="12" pageNumber="45" reason="1">
<emphasis id="B9331749C90DFD02FF5EFF41FE67FF6E" bold="true" box="[151,509,151,178]" pageId="12" pageNumber="45">
<taxonomicName id="4C47B0D8C90DFD02FF5EFF41FE0DFF6D" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[151,407,151,178]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
<emphasis id="B9331749C90DFD02FF5EFF41FE0DFF6D" bold="true" box="[151,407,151,178]" italics="true" pageId="12" pageNumber="45">Nodocepheus luxtoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200AA32C90DFD02FE56FF41FE67FF6E" box="[415,509,151,177]" pageId="12" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D98D0C90DFD03FF5EFF6DFD43FBFE" lastPageId="13" lastPageNumber="46" pageId="12" pageNumber="45" type="description">
<paragraph id="8BF8CB5BC90DFD02FF5EFF6DFEB5FF09" blockId="12.[151,509,151,214]" box="[151,303,187,214]" pageId="12" pageNumber="45">
(
<figureCitation id="137CD7DEC90DFD02FF56FF6DFE96FF0A" box="[159,268,187,213]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="12" pageNumber="45">Figures 4</figureCitation>
,
<figureCitation id="137CD7DEC90DFD02FED1FF6DFEBCFF09" box="[280,294,187,214]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="12" pageNumber="45">5</figureCitation>
)
</paragraph>
<paragraph id="8BF8CB5BC90DFD02FF5EFF2BFDECFEBF" blockId="12.[151,1437,253,568]" pageId="12" pageNumber="45">
<emphasis id="B9331749C90DFD02FF5EFF2BFEB3FEC8" bold="true" box="[151,297,253,279]" pageId="12" pageNumber="45">Dimensions.</emphasis>
<typeStatus id="54FC75F9C90DFD02FEF8FF28FE03FEC7" box="[305,409,254,280]" pageId="12" pageNumber="45" type="holotype">Holotype</typeStatus>
length 204 μm, breadth 117 μm.
<typeStatus id="54FC75F9C90DFD02FCDCFF28FCEDFEC7" box="[789,887,254,280]" pageId="12" pageNumber="45" type="paratype">Paratype</typeStatus>
male: length 192 μm, breadth 113 μm.
<typeStatus id="54FC75F9C90DFD02FAF3FF28FA06FEC7" box="[1338,1436,254,280]" pageId="12" pageNumber="45" type="paratype">Paratype</typeStatus>
females: mean length (
<emphasis id="B9331749C90DFD02FE53FEF5FE32FEE3" box="[410,424,291,316]" italics="true" pageId="12" pageNumber="45">n</emphasis>
= 11) 211 μm (range 199221 μm); mean breadth 125 μm (range 113132 μm). Ratio of prodorsum to total length: 0.41 (
<typeStatus id="54FC75F9C90DFD02FDCBFE90FDF1FEBF" box="[514,619,326,352]" pageId="12" pageNumber="45" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BF8CB5BC90DFD02FF0EFEBFFC15FDE7" blockId="12.[151,1437,253,568]" pageId="12" pageNumber="45">
<emphasis id="B9331749C90DFD02FF0EFEBFFE89FE5C" bold="true" box="[199,275,361,387]" pageId="12" pageNumber="45">Adult.</emphasis>
<emphasis id="B9331749C90DFD02FED2FEBCFE02FE5B" box="[283,408,362,388]" italics="true" pageId="12" pageNumber="45">Prodorsum</emphasis>
: rostrum bluntly point in dorsal view, with short rostral setae on short tubercles, linked by short, transverse ridge (
<figureCitation id="137CD7DEC90DFD02FE6DFE58FD8BFE77" box="[420,529,398,424]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="12" pageNumber="45">Figure 4i</figureCitation>
). Broad lamellae extending to just posterior of rostrum, with strongly incurved apical cusps at right angles to the rest of the lamellae, bearing short, smooth, setiform lamellar setae (
<emphasis id="B9331749C90DFD02FADCFE64FAB3FE13" box="[1301,1321,434,460]" italics="true" pageId="12" pageNumber="45">le</emphasis>
) and with well-developed translamella (
<figureCitation id="137CD7DEC90DFD02FE2CFE00FDCCFE2E" box="[485,598,470,497]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="12" pageNumber="45">Figure 5a</figureCitation>
). Short, smooth interlamellar setae (
<emphasis id="B9331749C90DFD02FC3BFE01FB92FE2F" box="[1010,1032,471,496]" italics="true" pageId="12" pageNumber="45">in</emphasis>
) positioned on centrodorsal region. Bothridium well-developed, cylindrical, with broad sclerotized rim and internal circular thickening. Bothridial setae elongated, club-shaped, with strong spinose ornamentation apically.
</paragraph>
<caption id="DF389BD3C90DFD02FF51F800FAA6F831" ID-DOI="http://doi.org/10.5281/zenodo.7141866" ID-Zenodo-Dep="7141866" box="[152,1340,2006,2030]" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="12" pageNumber="45" startId="12.[152,255,2006,2030]" targetBox="[269,1320,604,1966]" targetPageId="12">
<paragraph id="8BF8CB5BC90DFD02FF51F800FAA6F831" blockId="12.[152,1340,2006,2030]" box="[152,1340,2006,2030]" pageId="12" pageNumber="45">
<emphasis id="B9331749C90DFD02FF51F800FE82F831" bold="true" box="[152,280,2006,2030]" pageId="12" pageNumber="45">FIGURE 5.</emphasis>
<taxonomicName id="4C47B0D8C90DFD02FED6F800FE6EF831" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[287,500,2006,2030]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
<emphasis id="B9331749C90DFD02FED6F800FE6EF831" box="[287,500,2006,2030]" italics="true" pageId="12" pageNumber="45">Nodocepheus luxtoni</emphasis>
</taxonomicName>
<emphasis id="B9331749C90DFD02FE33F800FDD1F831" bold="true" box="[506,587,2006,2030]" pageId="12" pageNumber="45">
<taxonomicNameLabel id="A200AA32C90DFD02FE33F800FDD1F831" box="[506,587,2006,2030]" pageId="12" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm
</paragraph>
</caption>
<paragraph id="8BF8CB5BC90CFD03FF0EFF41FEE5FEBE" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFF41FED9FF6F" box="[199,323,151,176]" italics="true" pageId="13" pageNumber="46">Notogaster</emphasis>
: humeral processes (h) well-developed, abutting bothridia, with large, elongated lateral lobe bearing short, setiform seta
<emphasis id="B9331749C90CFD03FEB9FF6DFEE6FF0B" box="[368,380,187,212]" italics="true" pageId="13" pageNumber="46">c</emphasis>
<subScript id="17C3C91EC90CFD03FEB4FF1AFE1FFF05" attach="left" box="[381,389,204,218]" fontSize="6" pageId="13" pageNumber="46">1</subScript>
and smaller medial lobe bearing seta
<emphasis id="B9331749C90CFD03FCEAFF6DFCB5FF0B" box="[803,815,187,212]" italics="true" pageId="13" pageNumber="46">c</emphasis>
<subScript id="17C3C91EC90CFD03FCF9FF1AFCA2FF05" attach="right" box="[816,824,204,218]" fontSize="6" pageId="13" pageNumber="46">2</subScript>
; lateral lobe extending to point posterior of base of seta
<emphasis id="B9331749C90CFD03FF5EFF0BFF37FF28" box="[151,173,221,247]" italics="true" pageId="13" pageNumber="46">la</emphasis>
(
<figureCitation id="137CD7DEC90CFD03FF72FF0BFEB1FF27" box="[187,299,221,248]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="13" pageNumber="46">Figure 5a</figureCitation>
). Humeral processes smooth, not ornamented or sculpted. Notogaster smooth, ovoid, longer (216 μm;
<typeStatus id="54FC75F9C90CFD03FF5EFED6FF61FEC5" box="[151,251,256,282]" pageId="13" pageNumber="46" type="holotype">holotype</typeStatus>
) than broad; with 11 pairs of smooth, setiform setae positioned marginally (
<emphasis id="B9331749C90CFD03FB8BFED7FBD4FEC5" box="[1090,1102,257,282]" italics="true" pageId="13" pageNumber="46">c</emphasis>
<subScript id="17C3C91EC90CFD03FB87FEC5FBCCFEFE" attach="left" box="[1102,1110,275,289]" fontSize="6" pageId="13" pageNumber="46">1</subScript>
and
<emphasis id="B9331749C90CFD03FB42FED7FB0DFEC5" box="[1163,1175,257,282]" italics="true" pageId="13" pageNumber="46">c</emphasis>
<subScript id="17C3C91EC90CFD03FB5EFEC5FB05FEFE" attach="left" box="[1175,1183,275,289]" fontSize="6" pageId="13" pageNumber="46">2</subScript>
plus
<emphasis id="B9331749C90CFD03FB13FED6FB78FEC5" box="[1242,1250,256,282]" italics="true" pageId="13" pageNumber="46">l</emphasis>
,
<emphasis id="B9331749C90CFD03FB26FED7FA99FEC5" box="[1263,1283,257,282]" italics="true" pageId="13" pageNumber="46">m</emphasis>
and
<emphasis id="B9331749C90CFD03FAFEFED7FADFFEC5" box="[1335,1349,257,282]" italics="true" pageId="13" pageNumber="46">p</emphasis>
series);
<emphasis id="B9331749C90CFD03FF5EFEF5FF05FEE2" box="[151,159,291,317]" italics="true" pageId="13" pageNumber="46">l</emphasis>
and
<emphasis id="B9331749C90CFD03FF1AFEF5FF7BFEE2" box="[211,225,291,317]" italics="true" pageId="13" pageNumber="46">h</emphasis>
series on short, squat tubercles. Dorsosejugal furrow transverse; with faint, incurved ridges on anteriomedian region of notogaster.
</paragraph>
<paragraph id="8BF8CB5BC90CFD03FF0EFEBCFB61FD88" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFEBCFEFFFE5B" box="[199,357,362,388]" italics="true" pageId="13" pageNumber="46">Ventral aspect</emphasis>
: subcapitulum secondarily anarthric, with tectum derived from genae, bearing median infrabuccal cleft (ibc) on its ventral surface (
<figureCitation id="137CD7DEC90CFD03FE36FE5BFDEAFE77" box="[511,624,397,424]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="13" pageNumber="46">Figure 5b</figureCitation>
); short, setiform setae
<emphasis id="B9331749C90CFD03FCA3FE58FCE2FE78" box="[874,888,398,423]" italics="true" pageId="13" pageNumber="46">a</emphasis>
and
<emphasis id="B9331749C90CFD03FC64FE58FC5BFE78" box="[941,961,398,423]" italics="true" pageId="13" pageNumber="46">m</emphasis>
with bases not hidden by tectum in ventral orientation; setae
<emphasis id="B9331749C90CFD03FEAAFE66FEEBFE15" box="[355,369,432,458]" italics="true" pageId="13" pageNumber="46">h</emphasis>
short, spiniform, positioned just posteriomedially of setae
<emphasis id="B9331749C90CFD03FBD0FE67FBB7FE15" box="[1049,1069,433,458]" italics="true" pageId="13" pageNumber="46">m</emphasis>
. Labiogenal articulation absent. Rutellum Y-shaped (
<figureCitation id="137CD7DEC90CFD03FE4DFE05FE63FE31" box="[388,505,467,494]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="13" pageNumber="46">
Figure
<quantity id="4CBF66BEC90CFD03FE13FE02FE63FE31" box="[474,505,468,494]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="13" pageNumber="46" unit="g" value="4.0">4g</quantity>
</figureCitation>
). Chelicerae with digitus fixus pointed, smooth; digitus mobilis comb-like with sparse, fine teeth (
<figureCitation id="137CD7DEC90CFD03FEA9FE21FE55FDCE" box="[352,463,503,529]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="13" pageNumber="46">Figure 4e</figureCitation>
). Palp setal formula 0-2-1-3-8; all setae smooth. Eupathidia not on prominent tubercles; eupathidium
<emphasis id="B9331749C90CFD03FEE3FDCDFEC3FDEB" box="[298,345,539,564]" italics="true" pageId="13" pageNumber="46">acm</emphasis>
not fused with solendidum ω which is positioned posteriorly on palptarsus (
<figureCitation id="137CD7DEC90CFD03FB7AFDCCFABAFDEB" box="[1203,1312,538,564]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="13" pageNumber="46">Figure 4f</figureCitation>
). Tutorium well-developed, with a long apical tooth and 4 short teeth; sub-tutorium with blunt cusp (
<figureCitation id="137CD7DEC90CFD03FBB4FDEBFB74FD88" box="[1149,1262,573,599]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="13" pageNumber="46">Figure 4h</figureCitation>
).
</paragraph>
<paragraph id="8BF8CB5BC90CFD03FF0EFDB6FDF5FCAE" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
Epimeral plates I fused in mid-line, others discrete, broadly separated in midline and longitudinally; plates I and II sub-rectangular, III and IV oval; epimeral setation 3-1-3-2 (
<figureCitation id="137CD7DEC90CFD03FCBFFD55FC7DFD41" box="[886,999,643,670]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="13" pageNumber="46">Figure 5b</figureCitation>
). Pedotectum I and II large, prominent, with pointed anterior and posterior lobes; discidium (
<emphasis id="B9331749C90CFD03FD23FD71FC9AFD1E" box="[746,768,679,705]" italics="true" pageId="13" pageNumber="46">di</emphasis>
) broad, well-developed, pointed. Perigenital carina present, bearing enantiophysis E4. Ventral plate ovoid, markedly longer than broad. Genital and anal plates each surrounded by ring of sclerotized cuticle; genital plates 33 µm long (
<typeStatus id="54FC75F9C90CFD03FCD5FD3BFC1EFCD8" box="[796,900,749,775]" pageId="13" pageNumber="46" type="holotype">holotype</typeStatus>
), with 5 pairs of setae; 1 pair of aggenital setae positioned just posterior of genital plates; 3 pairs of adanal setae, 2 pairs of anal setae; lyrifissure
<emphasis id="B9331749C90CFD03FB22FCC6FA95FCF5" box="[1259,1295,784,810]" italics="true" pageId="13" pageNumber="46">iad</emphasis>
in para-anal position (
<figureCitation id="137CD7DEC90CFD03FECAFCE5FEE2FC91" box="[259,376,819,846]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="13" pageNumber="46">Figure 5b</figureCitation>
). Anal plates lozenge-shaped, 41 µm long (
<typeStatus id="54FC75F9C90CFD03FCBBFCE5FC40FC92" box="[882,986,819,845]" pageId="13" pageNumber="46" type="holotype">holotype</typeStatus>
); separated some distance from genital plates. Preanal organ broad, flat T-shaped.
</paragraph>
<paragraph id="8BF8CB5BC90CFD03FF0EFCACFE31FC22" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFCACFEFDFC4B" box="[199,359,890,916]" italics="true" pageId="13" pageNumber="46">Lateral aspect</emphasis>
: Rostrum straight, vertical; tutorium (
<emphasis id="B9331749C90CFD03FCC5FCADFCB8FC4B" box="[780,802,891,916]" italics="true" pageId="13" pageNumber="46">tu</emphasis>
) and sub-tutorium (
<emphasis id="B9331749C90CFD03FC37FCADFB85FC4B" box="[1022,1055,891,916]" italics="true" pageId="13" pageNumber="46">stu</emphasis>
) each with acute point,
<emphasis id="B9331749C90CFD03FAECFCADFAA1FC4B" box="[1317,1339,891,916]" italics="true" pageId="13" pageNumber="46">tu</emphasis>
strongly serrated dorsally; pedoctectum I and II pointed rounded apically; discidium leaf-shaped, bluntly pointed (
<figureCitation id="137CD7DEC90CFD03FAE8FC4BFA14FC67" box="[1313,1422,925,952]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="13" pageNumber="46">Figure 5c</figureCitation>
). Lamellae projecting as far as dorsal apex of rostrum. Exobothridial seta (
<emphasis id="B9331749C90CFD03FC11FC17FC6BFC05" box="[984,1009,961,986]" italics="true" pageId="13" pageNumber="46">ex</emphasis>
) present. Anterior notogastral margin overhanging prodorsum.
</paragraph>
<paragraph id="8BF8CB5BC90CFD03FF0EFBD1FD43FBFE" blockId="13.[151,1437,151,2044]" box="[199,729,1031,1057]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFBD1FF66FBFF" box="[199,252,1031,1056]" italics="true" pageId="13" pageNumber="46">Legs</emphasis>
: monodactylous; claw with a dorsal tooth.
</paragraph>
</subSubSection>
<subSubSection id="C35D98D0C90CFD03FF0EFBFCFE76FAE5" pageId="13" pageNumber="46" type="materials_examined">
<paragraph id="8BF8CB5BC90CFD03FF0EFBFCFE76FAE5" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFBFCFC11FB9B" bold="true" box="[199,907,1066,1092]" pageId="13" pageNumber="46">Type designation, material Examined and Locality Data.</emphasis>
<materialsCitation id="3B2FC106C90CFD03FC5EFBFCFBA6FB55" ID-GBIF-Occurrence="3920727305" collectingDate="2009-09-29" collectorName="M. J. Colloff" county="Sassafras" elevation="1077" latitude="-37.316666" location="Errinundra National Park" longLatPrecision="1182" longitude="148.85" municipality="Errinundra Saddle" pageId="13" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Victoria" typeStatus="holotype">
<typeStatus id="54FC75F9C90CFD03FC5EFBFCFC65FB9B" box="[919,1023,1066,1092]" pageId="13" pageNumber="46" type="holotype">Holotype</typeStatus>
female and
<specimenCount id="9D4100D2C90CFD03FB50FBFCFA02FB9B" box="[1177,1432,1066,1092]" pageId="13" pageNumber="46" type="female" typeStatus="paratype">five paratype females</specimenCount>
, moss on
<collectingCounty id="6299B3D7C90CFD03FF35FB9BFEFEFBB8" box="[252,356,1101,1127]" pageId="13" pageNumber="46">Sassafras</collectingCounty>
(
<taxonomicName id="4C47B0D8C90CFD03FEBFFB9BFD01FBB8" box="[374,667,1101,1127]" class="Magnoliopsida" family="Atherospermataceae" genus="Atherosperma" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="46" phylum="Tracheophyta" rank="species" species="moschatum">
<emphasis id="B9331749C90CFD03FEBFFB9BFD01FBB8" box="[374,667,1101,1127]" italics="true" pageId="13" pageNumber="46">Atherosperma moschatum</emphasis>
</taxonomicName>
), cool temperate rainforest,
<quantity id="4CBF66BEC90CFD03FC12FB9BFBACFBB7" box="[987,1078,1101,1128]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="13" pageNumber="46" unit="m" value="1077.0">
<elevation id="006A2C68C90CFD03FC12FB9BFBACFBB7" box="[987,1078,1101,1128]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="13" pageNumber="46" unit="m" value="1077.0">1077 m</elevation>
</quantity>
.,
<collectingMunicipality id="6B9C5121C90CFD03FB8EFB9BFA8DFBB8" box="[1095,1303,1101,1127]" pageId="13" pageNumber="46">Errinundra Saddle</collectingMunicipality>
,
<location id="8E989D80C90CFD03FAEAFB9BFEAFFB55" LSID="urn:lsid:plazi:treatment:03EE7A4DC90DFD03FF5EFF41FC7FF823:8E989D80C90CFD03FAEAFB9BFEAFFB55" county="Sassafras" latitude="-37.316666" longLatPrecision="1182" longitude="148.85" municipality="Errinundra Saddle" name="Errinundra National Park" pageId="13" pageNumber="46" stateProvince="Victoria">Errinundra National Park</location>
,
<collectingRegion id="498305B9C90CFD03FE89FBA6FE07FB55" box="[320,413,1136,1162]" country="Australia" name="Victoria" pageId="13" pageNumber="46">Victoria</collectingRegion>
,
<geoCoordinate id="EE73AD9CC90CFD03FE60FBA6FD9FFB55" box="[425,517,1136,1162]" degrees="37" direction="south" minutes="19" orientation="latitude" pageId="13" pageNumber="46" precision="925" value="-37.316666">37°19S</geoCoordinate>
<geoCoordinate id="EE73AD9CC90CFD03FDC5FBA6FDE0FB54" box="[524,634,1136,1163]" degrees="148" direction="east" minutes="51" orientation="longitude" pageId="13" pageNumber="46" precision="925" value="148.85">148°51E</geoCoordinate>
,
<quantity id="4CBF66BEC90CFD03FD4FFBA6FD7BFB54" box="[646,737,1136,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="13" pageNumber="46" unit="m" value="1030.0">
<elevation id="006A2C68C90CFD03FD4FFBA6FD7BFB54" box="[646,737,1136,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="13" pageNumber="46" unit="m" value="1030.0">1030 m</elevation>
</quantity>
., coll.
<collectorName id="26B2AE8DC90CFD03FCE0FBA7FC2FFB55" box="[809,949,1136,1163]" pageId="13" pageNumber="46">M.J. Colloff</collectorName>
,
<date id="FFF9ED9BC90CFD03FC08FBA6FBA6FB55" box="[961,1084,1136,1162]" pageId="13" pageNumber="46" value="2009-09-29">
<collectingDate id="EFBD1473C90CFD03FC08FBA6FBA6FB55" box="[961,1084,1136,1162]" pageId="13" pageNumber="46" value="2009-09-29">29.ix.2009</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B2FC106C90CFD03FB8EFBA7FC3CFB0E" ID-GBIF-Occurrence="3920727303" collectingDate="2008-03-14" collectionCode="T" collectorName="Weir &amp; C. Lemann" elevation="45" latitude="-41.65" location="Corinna" longLatPrecision="1155" longitude="145.1" municipality="State Forest" pageId="13" pageNumber="46" specimenCode="ANIC 3757" specimenCount="45" stateProvince="Tasmania" typeStatus="paratype">
<typeStatus id="54FC75F9C90CFD03FB8EFBA7FB37FB54" box="[1095,1197,1137,1163]" pageId="13" pageNumber="46" type="paratype">Paratype</typeStatus>
: female,
<specimenCode id="DBE16320C90CFD03FAD8FBA6FA02FB54" box="[1297,1432,1136,1163]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="13" pageNumber="46">ANIC 3757</specimenCode>
, sifted litter and mossy logs, rainforest with
<taxonomicName id="4C47B0D8C90CFD03FD41FB45FCB4FB71" box="[648,814,1171,1198]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="46" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9331749C90CFD03FD41FB45FC9EFB72" box="[648,772,1171,1197]" italics="true" pageId="13" pageNumber="46">Eucalyptus</emphasis>
sp.
</taxonomicName>
,
<collectingMunicipality id="6B9C5121C90CFD03FCF5FB42FC5FFB71" box="[828,965,1172,1198]" pageId="13" pageNumber="46">State Forest</collectingMunicipality>
, 2.2 km north-east of
<location id="8E989D80C90CFD03FB08FB45FA84FB72" LSID="urn:lsid:plazi:treatment:03EE7A4DC90DFD03FF5EFF41FC7FF823:8E989D80C90CFD03FB08FB45FA84FB72" box="[1217,1310,1171,1197]" latitude="-41.65" longLatPrecision="1155" longitude="145.1" municipality="State Forest" name="Corinna" pageId="13" pageNumber="46" stateProvince="Tasmania">Corinna</location>
,
<collectingRegion id="498305B9C90CFD03FAE3FB45FA02FB72" box="[1322,1432,1171,1197]" country="Australia" name="Tasmania" pageId="13" pageNumber="46">Tasmania</collectingRegion>
,
<geoCoordinate id="EE73AD9CC90CFD03FF5EFB61FF69FB0E" box="[151,243,1207,1233]" degrees="41" direction="south" minutes="39" orientation="latitude" pageId="13" pageNumber="46" precision="925" value="-41.65">41°39S</geoCoordinate>
<geoCoordinate id="EE73AD9CC90CFD03FF35FB60FEF7FB0E" box="[252,365,1206,1233]" degrees="145" direction="east" minutes="06" orientation="longitude" pageId="13" pageNumber="46" precision="925" value="145.1">145°06E</geoCoordinate>
.,
<specimenCount id="9D4100D2C90CFD03FEB6FB60FE59FB0E" box="[383,451,1206,1233]" pageId="13" pageNumber="46" type="generic">
<quantity id="4CBF66BEC90CFD03FEB6FB60FE25FB0E" box="[383,447,1206,1233]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="46" unit="m" value="45.0">
<elevation id="006A2C68C90CFD03FEB6FB60FE25FB0E" box="[383,447,1206,1233]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="46" unit="m" value="45.0">45 m</elevation>
</quantity>
.
</specimenCount>
, coll.
<collectionCode id="ED56539EC90CFD03FDC2FB61FD87FB0E" box="[523,541,1207,1233]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="46">T</collectionCode>
.
<collectorName id="26B2AE8DC90CFD03FDE3FB61FDFAFB0E" box="[554,608,1207,1233]" pageId="13" pageNumber="46">Weir</collectorName>
and
<collectorName id="26B2AE8DC90CFD03FD53FB61FC86FB0E" box="[666,796,1207,1233]" pageId="13" pageNumber="46">C. Lemann</collectorName>
,
<date id="FFF9ED9BC90CFD03FCE0FB61FC3CFB0E" box="[809,934,1207,1233]" pageId="13" pageNumber="46" value="2008-03-14">
<collectingDate id="EFBD1473C90CFD03FCE0FB61FC3CFB0E" box="[809,934,1207,1233]" pageId="13" pageNumber="46" value="2008-03-14">14.iii.2008</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B2FC106C90CFD03FC7BFB61FA84FB2B" ID-GBIF-Occurrence="3920727304" collectingDate="1989-06-20" collectorName="P. Greenslade" latitude="-42.9" location="Wellington, Tasmania" longLatPrecision="1147" longitude="147.23334" pageId="13" pageNumber="46" specimenCount="1" stateProvince="Wellington, Tasmania" typeStatus="paratype">
<typeStatus id="54FC75F9C90CFD03FC7BFB61FB82FB0E" box="[946,1048,1207,1233]" pageId="13" pageNumber="46" type="paratype">Paratype</typeStatus>
: female, TAS-183
<collectorName id="26B2AE8DC90CFD03FB39FB61FAAAFB0E" box="[1264,1328,1207,1233]" pageId="13" pageNumber="46">Litter</collectorName>
, eucalypt forest,
<collectorName id="26B2AE8DC90CFD03FF2DFB0CFE09FB2B" box="[228,403,1242,1268]" pageId="13" pageNumber="46">Old Farm Road</collectorName>
,
<collectorName id="26B2AE8DC90CFD03FE54FB0CFE72FB2B" box="[413,488,1242,1268]" pageId="13" pageNumber="46">Mount</collectorName>
<collectingRegion id="498305B9C90CFD03FE24FB0CFD79FB2B" box="[493,739,1242,1268]" isEnumeration="true" pageId="13" pageNumber="46">Wellington, Tasmania</collectingRegion>
,
<geoCoordinate id="EE73AD9CC90CFD03FD24FB0CFCD3FB2A" box="[749,841,1242,1269]" degrees="42" direction="south" minutes="54" orientation="latitude" pageId="13" pageNumber="46" precision="925" value="-42.9">42°54S</geoCoordinate>
<geoCoordinate id="EE73AD9CC90CFD03FC86FB0CFC27FB2B" box="[847,957,1242,1268]" degrees="147" direction="east" minutes="14" orientation="longitude" pageId="13" pageNumber="46" precision="925" value="147.23334">147°14E</geoCoordinate>
, coll.
<collectorName id="26B2AE8DC90CFD03FC36FB0CFB03FB2B" box="[1023,1177,1242,1268]" pageId="13" pageNumber="46">P. Greenslade</collectorName>
,
<date id="FFF9ED9BC90CFD03FB6AFB0CFA84FB2B" box="[1187,1310,1242,1268]" pageId="13" pageNumber="46" value="1989-06-20">
<collectingDate id="EFBD1473C90CFD03FB6AFB0CFA84FB2B" box="[1187,1310,1242,1268]" pageId="13" pageNumber="46" value="1989-06-20">20.vi.1989</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B2FC106C90CFD03FAE1FB0CFE73FAE5" ID-GBIF-Occurrence="3920727302" collectingDate="2011-07-19" collectorName="M. J. Colloff" location="Otway National Park" pageId="13" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Victoria" typeStatus="paratype">
<typeStatus id="54FC75F9C90CFD03FAE1FB0CFA03FB2B" box="[1320,1433,1242,1268]" pageId="13" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4100D2C90CFD03FF5EFB2BFEBCFAC8" box="[151,294,1277,1303]" pageId="13" pageNumber="46" type="female">four females</specimenCount>
, moss,
<taxonomicName id="4C47B0D8C90CFD03FEBFFB2BFD15FAC8" box="[374,655,1277,1303]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="46" phylum="Tracheophyta" rank="species" species="cunninghami">
<emphasis id="B9331749C90CFD03FEBFFB2BFD15FAC8" box="[374,655,1277,1303]" italics="true" pageId="13" pageNumber="46">Nothofagus cunninghami</emphasis>
</taxonomicName>
rainforest, rainforest walk, Maits Rest,
<location id="8E989D80C90CFD03FB8CFB2BFAA8FAC8" LSID="urn:lsid:plazi:treatment:03EE7A4DC90DFD03FF5EFF41FC7FF823:8E989D80C90CFD03FB8CFB2BFAA8FAC8" box="[1093,1330,1277,1303]" name="Otway National Park" pageId="13" pageNumber="46" stateProvince="Victoria">Otway National Park</location>
,
<collectingRegion id="498305B9C90CFD03FAF2FB2BFA02FAC8" box="[1339,1432,1277,1303]" country="Australia" name="Victoria" pageId="13" pageNumber="46">Victoria</collectingRegion>
, coll.
<collectorName id="26B2AE8DC90CFD03FF06FAF7FEC1FAE5" box="[207,347,1312,1339]" pageId="13" pageNumber="46">M.J. Colloff</collectorName>
,
<date id="FFF9ED9BC90CFD03FEAFFAF6FE73FAE5" box="[358,489,1312,1338]" pageId="13" pageNumber="46" value="2011-07-19">
<collectingDate id="EFBD1473C90CFD03FEAFFAF6FE73FAE5" box="[358,489,1312,1338]" pageId="13" pageNumber="46" value="2011-07-19">19.vii.2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D98D0C90CFD03FF0EFA92FD99FA7B" pageId="13" pageNumber="46" type="etymology">
<paragraph id="8BF8CB5BC90CFD03FF0EFA92FD99FA7B" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFA92FED4FA81" bold="true" box="[199,334,1348,1374]" pageId="13" pageNumber="46">Etymology.</emphasis>
This species is named in honour and memory of my friend and colleague, the late Dr Malcolm Luxton, in recognition of his contribution to our understanding of the taxonomy and biogeography of the oribatid mites of the Australasian region.
</paragraph>
</subSubSection>
<subSubSection id="C35D98D0C90CFD03FF0EFA7BFAD8F961" pageId="13" pageNumber="46" type="diagnosis">
<paragraph id="8BF8CB5BC90CFD03FF0EFA7BFAD8F961" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EFA7BFED8FA18" bold="true" box="[199,322,1453,1479]" pageId="13" pageNumber="46">Diagnosis.</emphasis>
<taxonomicName id="4C47B0D8C90CFD03FE84FA7BFDA7FA18" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[333,573,1453,1479]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
<emphasis id="B9331749C90CFD03FE84FA7BFDA7FA18" box="[333,573,1453,1479]" italics="true" pageId="13" pageNumber="46">Nodocepheus luxtoni</emphasis>
</taxonomicName>
<emphasis id="B9331749C90CFD03FD80FA7BFD3DFA18" bold="true" box="[585,679,1453,1479]" pageId="13" pageNumber="46">
<taxonomicNameLabel id="A200AA32C90CFD03FD80FA7BFD3DFA18" box="[585,679,1453,1479]" pageId="13" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other species in the genus by the following combination of characters: (1) most of the notogastral setae are on squat tubercles; (2) with the epimeral setal formula 3-1-3-2; (3) the very large, pedotecta I and II with an indented apex, appearing shallowly bifurcate; (4) the prominent, smooth, elongated lateral lobe of the humeral process, extending to a point posterior of base of seta
<emphasis id="B9331749C90CFD03FAB6F9C1FA0FF9EE" box="[1407,1429,1559,1585]" italics="true" pageId="13" pageNumber="46">la</emphasis>
; (5) the broad, discidium, bluntly pointed; (6) the rostral setae on short tubercles, linked by a short, transverse ridge; (7) the strongly incurved and opposing lamellar cusps (8) the smooth, short setae
<emphasis id="B9331749C90CFD03FBE7F98BFBD8F9A8" box="[1070,1090,1629,1655]" italics="true" pageId="13" pageNumber="46">le</emphasis>
positioned sub-apically on the lamellar cusps; (8) the bothridial seta with a long stalk, the head club-shaped and covered in long, well-developed spinules; (9) the tutorium with a long apical tooth and 4 short teeth and the sub-tutorium with a blunt cusp.
</paragraph>
</subSubSection>
<subSubSection id="C35D98D0C90CFD03FF0EF911FC7FF823" pageId="13" pageNumber="46" type="discussion">
<paragraph id="8BF8CB5BC90CFD03FF0EF911FC7FF823" blockId="13.[151,1437,151,2044]" pageId="13" pageNumber="46">
<emphasis id="B9331749C90CFD03FF0EF911FEAEF93E" bold="true" box="[199,308,1735,1761]" pageId="13" pageNumber="46">Remarks</emphasis>
.
<bibRefCitation id="EFD6B6AAC90CFD03FE8AF911FE63F93E" author="Mahunka, S." box="[323,505,1735,1761]" pageId="13" pageNumber="46" pagination="151 - 180" refId="ref15225" refString="Mahunka, S. (1983) Oribatids from the eastern part of the Ethiopian region. II. Acta Zoologica Academiae Scientiarum Hungaricae, 29, 151 - 180." type="journal article" year="1983">Mahunka (1983</bibRefCitation>
, p. 164) stated that of the six species described by that time, all of them except
<taxonomicName id="4C47B0D8C90CFD03FA43F911FE92F8DB" authorityName="Mahunka" authorityYear="1980" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="cerebralis">
<emphasis id="B9331749C90CFD03FA43F911FE92F8DB" italics="true" pageId="13" pageNumber="46">N cerebralis</emphasis>
</taxonomicName>
“stand very near to each other” and that, based on an examination of
<typeStatus id="54FC75F9C90CFD03FC3AF93CFBABF8DB" box="[1011,1073,1770,1796]" pageId="13" pageNumber="46">types</typeStatus>
, they could only be distinguished by the morphology of the humeral processes, tutorium and sub-tutorium, with chaetotaxy and the morphology of the sensillus considered as secondary characters.
<taxonomicName id="4C47B0D8C90CFD03FD5CF8E6FC1BF895" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[661,897,1840,1866]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
<emphasis id="B9331749C90CFD03FD5CF8E6FC1BF895" box="[661,897,1840,1866]" italics="true" pageId="13" pageNumber="46">Nodocepheus luxtoni</emphasis>
</taxonomicName>
<emphasis id="B9331749C90CFD03FC4EF8E6FC7BF895" bold="true" box="[903,993,1840,1866]" pageId="13" pageNumber="46">
<taxonomicNameLabel id="A200AA32C90CFD03FC4EF8E6FC7BF895" box="[903,993,1840,1866]" pageId="13" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to a group with
<taxonomicName id="4C47B0D8C90CFD03FB3EF8E7FAF7F895" authorityName="Mahunka" authorityYear="1983" box="[1271,1389,1840,1866]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="B9331749C90CFD03FB3EF8E7FAF7F895" box="[1271,1389,1840,1866]" italics="true" pageId="13" pageNumber="46">N. baloghi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C47B0D8C90CFD03FF5EF882FE8DF8B2" authorityName="Mahunka" authorityYear="1985" box="[151,279,1876,1901]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B9331749C90CFD03FF5EF882FE8DF8B2" box="[151,279,1876,1901]" italics="true" pageId="13" pageNumber="46">N. minimus</emphasis>
</taxonomicName>
that share the following character states: (1) the bothridial seta has a long stalk and a club-shaped head densely ornamented with well-developed spines and (2) the humeral process is large, expanded and extends from the bothridium to a position at least as far posteriorly as the level of the base of setae
<emphasis id="B9331749C90CFD03FB94F84CFBE9F86B" box="[1117,1139,1946,1972]" italics="true" pageId="13" pageNumber="46">la</emphasis>
. However,
<taxonomicName id="4C47B0D8C90CFD03FB3CF84DFAF7F86B" authorityName="Mahunka" authorityYear="1983" box="[1269,1389,1946,1972]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis id="B9331749C90CFD03FB3CF84DFAF7F86B" box="[1269,1389,1946,1972]" italics="true" pageId="13" pageNumber="46">N. baloghi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C47B0D8C90CFD03FF5EF868FE8CF808" authorityName="Mahunka" authorityYear="1985" box="[151,278,1982,2007]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="46" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B9331749C90CFD03FF5EF868FE8CF808" box="[151,278,1982,2007]" italics="true" pageId="13" pageNumber="46">N. minimus</emphasis>
</taxonomicName>
both have setae
<emphasis id="B9331749C90CFD03FE07F86BFE78F808" box="[462,482,1981,2007]" italics="true" pageId="13" pageNumber="46">le</emphasis>
with spinose ornamentation, rather than smooth, and are positioned apically on broad lamellar cusps, rather than sub-apically on narrow, strongly incurved cusps.
</paragraph>
</subSubSection>
</treatment>
</document>