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<document id="9C44EA51768896C5ED076B561EDECB81" ID-DOI="10.11646/zootaxa.3682.2.3" ID-GBIF-Dataset="3ca59aa7-bc2b-4dd5-99bd-337a33215dbb" ID-ISSN="1175-5326" ID-Zenodo-Dep="222599" ID-ZooBank="0B7C8F41-3843-48C2-9D2D-77C17DC4B83B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460642359980" checkinUser="plazi" docAuthor="Pérez-González, Sergio &amp; Zaballos, Juan P." docDate="2013" docId="03E92035F969FFA6FF555D8F5877A3D4" docLanguage="en" docName="zt03682p269.pdf" docOrigin="Zootaxa 3682 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhlocharis scrofa Perez-Gonzalez &amp; Zaballos, new species" docType="treatment" docVersion="7" lastPageNumber="258" masterDocId="FFD0584DF96FFFAFFFC25C705A50A13B" masterDocTitle="Tarsal tetramery and extreme size reduction in Anillini (Coleoptera, Carabidae, Trechinae): the case of Typhlocharis Dieck, 1869; description of three new species and definition of a new intra-generic species group" masterLastPageNumber="269" masterPageNumber="249" pageNumber="255" updateTime="1698336104760" updateUser="plazi">
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<mods:title id="888B087779EBC9E1B8F2D6FBE8487220">Tarsal tetramery and extreme size reduction in Anillini (Coleoptera, Carabidae, Trechinae): the case of Typhlocharis Dieck, 1869; description of three new species and definition of a new intra-generic species group</mods:title>
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<mods:namePart id="A3C888D7D085E847DC33D8C0C386F87E">Pérez-González, Sergio</mods:namePart>
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<mods:namePart id="0203184580DC9001DDEC221714656C68">Zaballos, Juan P.</mods:namePart>
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<mods:date id="3F9925480564F24299F166701470F894">2013</mods:date>
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<mods:number id="92F9A6BF514D7EA40188F7215E6EB7BB">3682</mods:number>
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<treatment id="03E92035F969FFA6FF555D8F5877A3D4" ID-DOI="http://doi.org/10.5281/zenodo.6159488" ID-GBIF-Taxon="119571405" ID-Zenodo-Dep="6159488" LSID="urn:lsid:plazi:treatment:03E92035F969FFA6FF555D8F5877A3D4" httpUri="http://treatment.plazi.org/id/03E92035F969FFA6FF555D8F5877A3D4" lastPageId="9" lastPageNumber="258" pageId="6" pageNumber="255">
<subSubSection id="C35AC2A8F969FFA9FF555D8F5B5AA301" pageId="6" pageNumber="255" type="nomenclature">
<paragraph id="8BFF9123F969FFA9FF555D8F592BA322" blockId="6.[151,891,511,570]" box="[151,891,511,537]" pageId="6" pageNumber="255">
<heading id="D0B7264FF969FFA9FF555D8F592BA322" bold="true" box="[151,891,511,537]" fontSize="11" level="1" pageId="6" pageNumber="255" reason="1">
<emphasis id="B9344D31F969FFA9FF555D8F592BA322" bold="true" box="[151,891,511,537]" pageId="6" pageNumber="255">
<taxonomicName id="4C40EAA0F969FFA9FF555D8F58B4A322" ID-CoL="59VKQ" authority="Perez-Gonzalez &amp; Zaballos" authorityName="Perez-Gonzalez &amp; Zaballos" box="[151,740,511,537]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F969FFA9FF555D8F5BD7A322" bold="true" box="[151,391,511,537]" italics="true" pageId="6" pageNumber="255">Typhlocharis scrofa</emphasis>
Pérez-González &amp; Zaballos
</taxonomicName>
<taxonomicNameLabel id="A207F04AF969FFA9FD295D8F592BA322" box="[747,891,511,537]" pageId="6" pageNumber="255" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF9123F969FFA9FF555E525B5AA301" blockId="6.[151,891,511,570]" box="[151,266,546,570]" pageId="6" pageNumber="255">
(
<figureCitation id="137B8DA6F969FFA9FF5D5E525B53A301" box="[159,259,546,570]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="7.[151,250,1949,1972]" captionStartId-1="8.[151,250,907,930]" captionTargetBox-0="[176,1406,188,1906]" captionTargetBox-1="[165,1406,187,861]" captionTargetId-0="figure@7.[155,1432,169,1928]" captionTargetId-1="figure@8.[151,1436,169,886]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 3. Typhlocharis scrofa sp. n. habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." captionText-1="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri-0="https://zenodo.org/record/222602/files/figure.png" httpUri-1="https://zenodo.org/record/222603/files/figure.png" pageId="6" pageNumber="255">Figs 3–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F969FFA9FF555E1B5B0CA229" pageId="6" pageNumber="255" type="materials_examined">
<paragraph id="8BFF9123F969FFA9FF555E1B5B0CA229" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
<emphasis id="B9344D31F969FFA9FF555E1B5B74A3B8" bold="true" box="[151,292,618,643]" pageId="6" pageNumber="255">
<typeStatus id="54FB2F81F969FFA9FF555E1B5A82A3B8" box="[151,210,619,643]" pageId="6" pageNumber="255">Type</typeStatus>
series.
</emphasis>
<typeStatus id="54FB2F81F969FFA9FEEE5E1A5BC7A3B9" box="[300,407,618,642]" pageId="6" pageNumber="255" type="holotype">Holotype</typeStatus>
, 3,
<collectingCountry id="F357D1B3F969FFA9FE0B5E1A584CA3B9" box="[457,540,618,642]" name="Spain" pageId="6" pageNumber="255">SPAIN</collectingCountry>
: Huelva, Cumbres de San Bartolomé (
<quantity id="4CB83CC6F969FFA9FC105E1A5E74A3B9" box="[978,1060,618,642]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="255" unit="km" value="3.4">3.4 km</quantity>
SW), Arroyo La Dehesa,
<date id="FFFEB7E3F969FFA9FA935E1A5A82A39D" pageId="6" pageNumber="255" value="2012-03-21">
21-03-
<quantity id="4CB83CC6F969FFA9FF555EFE5A82A39D" box="[151,210,654,678]" metricMagnitude="6" metricUnit="m" metricValue="2.012358" pageId="6" pageNumber="255" unit="m" value="2012358.0">2012</quantity>
</date>
, 358 m (
<geoCoordinate id="EE74F7E4F969FFA9FEF35EFE5BCFA39D" box="[305,415,654,678]" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="925" value="38.066666">N 38º 04’</geoCoordinate>
,
<geoCoordinate id="EE74F7E4F969FFA9FE695EFF584EA39D" box="[427,542,654,678]" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="925" value="-6.7666664">W 06º 46’</geoCoordinate>
), J.P. Zaballos &amp; S. Pérez leg. (Coll. J.P. Zaballos,
<collectionCode id="ED5109E6F969FFA9FB965EFE5EC9A39D" LSID="urn:lsid:biocol.org:col:34854" box="[1108,1177,654,678]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="6" pageNumber="255">UCM</collectionCode>
).
<typeStatus id="54FB2F81F969FFA9FB695EFF5F4CA39C" box="[1195,1308,655,679]" pageId="6" pageNumber="255" type="paratype">Paratypes</typeStatus>
: 62 3, 45 ƤƤ same data as
<typeStatus id="54FB2F81F969FFA9FE9D5EC25B94A3F1" box="[351,452,690,714]" pageId="6" pageNumber="255" type="holotype">holotype</typeStatus>
; 1 Ƥ
<collectingCountry id="F357D1B3F969FFA9FDC05EC25805A3F1" box="[514,597,690,714]" name="Spain" pageId="6" pageNumber="255">SPAIN</collectingCountry>
: Huelva, Cumbres de San Bartolomé (
<quantity id="4CB83CC6F969FFA9FBC75EC25E05A3F1" box="[1029,1109,690,714]" metricMagnitude="3" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="255" unit="km" value="7.4">7.4 km</quantity>
SW), Rio Sillo,
<date id="FFFEB7E3F969FFA9FAD05EC25FC8A3F1" box="[1298,1432,690,714]" pageId="6" pageNumber="255" value="2012-03-21">
21-03-
<quantity id="4CB83CC6F969FFA9FA985EC25FC8A3F1" box="[1370,1432,690,714]" metricMagnitude="6" metricUnit="m" metricValue="2.012275" pageId="6" pageNumber="255" unit="m" value="2012275.0">2012</quantity>
</date>
, 275 m (
<geoCoordinate id="EE74F7E4F969FFA9FF325EA65B0FA3D5" box="[240,351,726,750]" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="925" value="38.05">N 38º 03'</geoCoordinate>
,
<geoCoordinate id="EE74F7E4F969FFA9FEA95EA75BB0A3D5" box="[363,480,726,750]" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="925" value="-6.8166666">W 06º 49'</geoCoordinate>
), J.P. Zaballos &amp; S. Pérez leg. (106 ex. Coll. J.P. Zaballos,
<collectionCode id="ED5109E6F969FFA9FB445EA65E99A3D5" LSID="urn:lsid:biocol.org:col:34854" box="[1158,1225,726,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="6" pageNumber="255">UCM</collectionCode>
, 2 ex. nº ref. 2375
<collectionCode id="ED5109E6F969FFA9FF555E8A5ABEA229" LSID="urn:lsid:biocol.org:col:35275" box="[151,238,762,786]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="6" pageNumber="255">MNCN</collectionCode>
, Madrid).
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F969FFA9FF055F6E5E63A2D1" pageId="6" pageNumber="255" type="diagnosis">
<paragraph id="8BFF9123F969FFA9FF055F6E5E63A2D1" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
<emphasis id="B9344D31F969FFA9FF055F6E5B11A20C" bold="true" box="[199,321,798,823]" pageId="6" pageNumber="255">Diagnosis.</emphasis>
Very small endogean beetle, anophthalmous, with narrow and subrectangular body covered by microreticulate integument and scattered pubescence. Vertex with
<emphasis id="B9344D31F969FFA9FC4D5F345E73A260" box="[911,1059,835,859]" italics="true" pageId="6" pageNumber="255">pars stridens</emphasis>
. Gula partially fused to cephalic capsule. Elytra subtly serrated in lateral margins, almost imperceptible in posterior region. Apical margin with two pairs of denticles (a sutural pair and a pair associated to 7th stria). One pair of anterior discal setae. Umbilicate series with five setae (4+1). Tarsal formula: 4-4-4. Aedeagus as in
<figureCitation id="137B8DA6F969FFA9FC225FDE5E7BA2FD" box="[992,1067,942,966]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="6" pageNumber="255">Fig. 4</figureCitation>
A,B, with &quot;parenthesis-shaped&quot; endophallic sclerite. Female genitalia with club-shaped gonocoxites, as in
<figureCitation id="137B8DA6F969FFA9FC1A5FA25E4AA2D1" box="[984,1050,978,1002]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="6" pageNumber="255">Fig. 4</figureCitation>
C.
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F969FFA7FF055F865881A77E" lastPageId="8" lastPageNumber="257" pageId="6" pageNumber="255" type="description">
<paragraph id="8BFF9123F969FFA9FF055F865B32A509" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
<emphasis id="B9344D31F969FFA9FF055F865B09A534" bold="true" box="[199,345,1014,1039]" pageId="6" pageNumber="255">Description.</emphasis>
Length of
<typeStatus id="54FB2F81F969FFA9FE245F86581CA535" box="[486,588,1014,1038]" pageId="6" pageNumber="255" type="holotype">holotype</typeStatus>
:
<quantity id="4CB83CC6F969FFA9FD9F5F86589AA535" box="[605,714,1014,1038]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="255" unit="mm" value="0.86">0.86 mm</quantity>
; length of
<typeStatus id="54FB2F81F969FFA9FC965F875994A534" box="[852,964,1015,1039]" pageId="6" pageNumber="255" type="paratype">paratypes</typeStatus>
:
<quantity id="4CB83CC6F969FFA9FC165F865E2DA535" box="[980,1149,1014,1038]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="10.5" metricValueMin="8.6" pageId="6" pageNumber="255" unit="mm" value="0.955" valueMax="1.05" valueMin="0.86">0.86–1.05 mm</quantity>
(males),
<quantity id="4CB83CC6F969FFA9FB315F865FCCA535" box="[1267,1436,1014,1038]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.7" metricValueMin="8.8" pageId="6" pageNumber="255" unit="mm" value="0.975" valueMax="1.07" valueMin="0.88">0.88–1.07 mm</quantity>
(females) (
<figureCitation id="137B8DA6F969FFA9FED2586A5B04A509" box="[272,340,1050,1074]" captionStart="FIGURE 3" captionStartId="7.[151,250,1949,1972]" captionTargetBox="[176,1406,188,1906]" captionTargetId="figure@7.[155,1432,169,1928]" captionTargetPageId="7" captionText="FIGURE 3. Typhlocharis scrofa sp. n. habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222602/files/figure.png" pageId="6" pageNumber="255">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BFF9123F969FFA9FF05584E59BFA414" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
Head: almost as wide (
<quantity id="4CB83CC6F969FFA9FE22584E58DDA56D" box="[480,653,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="6" pageNumber="255" unit="mm" value="0.215" valueMax="0.24" valueMin="0.19">0.19–0.24 mm</quantity>
) as long (
<quantity id="4CB83CC6F969FFA9FCCC584E59EDA56D" box="[782,957,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="6" pageNumber="255" unit="mm" value="0.2" valueMax="0.23" valueMin="0.17">0.17–0.23 mm</quantity>
). Clypeus with straight anterior margin. Moniliform antennae with 11 rounded antennomeres progressively more quadrate (
<emphasis id="B9344D31F969FFA9FBA258135E84A540" bold="true" box="[1120,1236,1122,1147]" pageId="6" pageNumber="255">morph 1)</emphasis>
, except last one, pyriform. The last antennomere has a pattern of two antero-dorsal and one postero-dorsal
<emphasis id="B9344D31F969FFA9FB6C58F65FC5A5A5" box="[1198,1429,1158,1182]" italics="true" pageId="6" pageNumber="255">sensilla coeloconica</emphasis>
. Ventral
<emphasis id="B9344D31F969FFA9FF2C58DB5B81A5F8" box="[238,465,1195,1219]" italics="true" pageId="6" pageNumber="255">sensilla coeloconica</emphasis>
in antennomeres 5th and 6th. Stem of third antennomere short. Labium without special characters, but epilobes tend to be pointy. Ligula with prominent middle lobe and short, thick paraglossae. Gula partially fused to cephalic capsule (diffuse suture, only visible in proximal and distal ends) (
<figureCitation id="137B8DA6F969FFA9FB0458825F5CA431" box="[1222,1292,1266,1290]" captionStart="FIGURE 7" captionStartId="13.[151,250,1886,1909]" captionTargetBox="[155,1432,580,1885]" captionTargetId="figure@13.[155,1432,561,1891]" captionTargetPageId="13" captionText="FIGURE 7. Different types of gula in the genus Typhlocharis. Light microscopy photographs and schematic diagrams of: A) T. pacensis Zaballos &amp; Jeanne, 1987; wide gula, approx. twice as long as wide, lateral sutures clearly visible. B) T. elenae Serrano &amp; Aguiar, 2002; narrow gula, approx. four of five times as long as wide, lateral sutures clearly visible, subparallel. C) T. scrofa sp. n.; diffuse gula, same proportions as wide gula, but lateral sutures only visible in the proximal and distal ends, partially fused to the cephalic capsule." httpUri="https://zenodo.org/record/222606/files/figure.png" pageId="6" pageNumber="255">Fig. 7</figureCitation>
C). Cephalic chaetotaxy and rest of cephalic characters as described for
<taxonomicName id="4C40EAA0F969FFA9FCEB596859FEA415" box="[809,942,1302,1327]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F969FFA9FCEB596859FEA415" box="[809,942,1302,1327]" italics="true" pageId="6" pageNumber="255">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF969FFA9FC75596759BFA414" box="[951,1007,1303,1327]" pageId="6" pageNumber="255" rank="species">
<emphasis id="B9344D31F969FFA9FC75596759BFA414" bold="true" box="[951,1007,1303,1327]" pageId="6" pageNumber="255">sp.n.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFF9123F969FFA9FF05594B592CA774" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
Pronotum: subtrapezoidal, slightly longer (
<quantity id="4CB83CC6F969FFA9FD71594A591EA469" box="[691,846,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="6" pageNumber="255" unit="mm" value="0.265" valueMax="0.3" valueMin="0.23">0.23–0.3 mm</quantity>
) than wide (
<quantity id="4CB83CC6F969FFA9FC20594A5EDEA469" box="[994,1166,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.6" metricValueMin="2.1" pageId="6" pageNumber="255" unit="mm" value="0.235" valueMax="0.26" valueMin="0.21">0.21–0.26 mm</quantity>
), narrowed in posterior region. Anterior margin virtually straight, slightly crenulate; medial hiatus approx. as wide as two or three adjacent intersetae spaces. Posterior margin smoothly curved. Denticles of lateral margins near posterior angles absent or barely insinuated; the teeth in posterior angles are also blunt and smooth. Rest of characters and chaetotaxy as described for
<taxonomicName id="4C40EAA0F969FFA9FEF259BC5BE5A4D8" box="[304,437,1483,1507]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F969FFA9FEF259BC5BE5A4D8" box="[304,437,1483,1507]" italics="true" pageId="6" pageNumber="255">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF969FFA9FE7F59BB5BA4A4D8" box="[445,500,1483,1507]" pageId="6" pageNumber="255" rank="species">
<emphasis id="B9344D31F969FFA9FE7F59BB5BA4A4D8" bold="true" box="[445,500,1483,1507]" pageId="6" pageNumber="255">sp.n.</emphasis>
</taxonomicNameLabel>
, except the row of setae parallel to anterior margin, with five to six pairs of setae [
<emphasis id="B9344D31F969FFA9FA4959BB5B2BA73C" bold="true" pageId="6" pageNumber="255">l- (l)-l-l-l-l / l-l-l-l-(l)-l</emphasis>
], three pairs of setae parallel to posterior margin
<emphasis id="B9344D31F969FFA9FC65599F5E74A73C" bold="true" box="[935,1060,1518,1543]" pageId="6" pageNumber="255">[l-l-l / l-l-l]</emphasis>
, a row of small, filiform and thin setae regularly placed in anterior and posterior margins, and four to five pairs of longitudinal rows of short pubescence in disc. Prosternum as described for
<taxonomicName id="4C40EAA0F969FFA9FD755A48596CA775" box="[695,828,1590,1615]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F969FFA9FD755A48596CA775" box="[695,828,1590,1615]" italics="true" pageId="6" pageNumber="255">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF969FFA9FC875A47592CA774" box="[837,892,1591,1615]" pageId="6" pageNumber="255" rank="species">
<emphasis id="B9344D31F969FFA9FC875A47592CA774" bold="true" box="[837,892,1591,1615]" pageId="6" pageNumber="255">sp.n.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFF9123F969FFA9FF055A2A5BA6A68C" blockId="6.[151,1437,618,1975]" pageId="6" pageNumber="255">
Elytra: approximately two times longer (
<quantity id="4CB83CC6F969FFA9FD6E5A2A5908A749" box="[684,856,1626,1650]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.6" metricValueMin="4.4" pageId="6" pageNumber="255" unit="mm" value="0.5" valueMax="0.56" valueMin="0.44">0.44–0.56 mm</quantity>
) than wide (
<quantity id="4CB83CC6F969FFA9FC345A2A5EF5A749" box="[1014,1189,1626,1650]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="6" pageNumber="255" unit="mm" value="0.245" valueMax="0.27" valueMin="0.22">0.22–0.27 mm</quantity>
), subparallel. Lateral margins with 17–20 sub-triangular denticles, very subtly marked and progressively smoother, practically inexistent beyond the first third. Apical margin with two pairs of denticles, a pair associated to the 7th stria and a pair associated to the suture, the latter pointing outwards and defining a characteristic &quot;V-shaped” end of the suture. Small and scarce elytral pores, present along 7th stria, scutellar region and disc. Transverse scutellar organ (straight to irregular) and pair of &quot;buttonholes&quot; present near base of elytra. Chaetotaxy: umbilicate series with anterior group of four setae and posterior group of one seta (
<figureCitation id="137B8DA6F969FFA9FD695B4258A1A671" box="[683,753,1842,1866]" captionStart="FIGURE 3" captionStartId="7.[151,250,1949,1972]" captionTargetBox="[176,1406,188,1906]" captionTargetId="figure@7.[155,1432,169,1928]" captionTargetPageId="7" captionText="FIGURE 3. Typhlocharis scrofa sp. n. habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222602/files/figure.png" pageId="6" pageNumber="255">Fig. 3</figureCitation>
). One pair of scutellar setae and one pair of anterior discal setae. Discal pubescence arranged in five pairs of longitudinal rows, progressively longer towards posterior region. Apical pair of pubescent setae present. Lateral margins with a short seta for every denticle. Other characters as described for
<taxonomicName id="4C40EAA0F969FFA9FEF35BD05BE7A68D" box="[305,439,1950,1975]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F969FFA9FEF35BD05BE7A68D" box="[305,439,1950,1975]" italics="true" pageId="6" pageNumber="255">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF969FFA9FE7C5BEF5BA6A68C" box="[446,502,1951,1975]" pageId="6" pageNumber="255" rank="species">
<emphasis id="B9344D31F969FFA9FE7C5BEF5BA6A68C" bold="true" box="[446,502,1951,1975]" pageId="6" pageNumber="255">sp.n.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="DF3FC1ABF968FFA8FF555BED58BDA6E9" httpUri="https://zenodo.org/record/222602/files/figure.png" pageId="7" pageNumber="256" targetBox="[176,1406,188,1906]" targetPageId="7">
<paragraph id="8BFF9123F968FFA8FF555BED58BDA6E9" blockId="7.[151,1436,1949,2002]" pageId="7" pageNumber="256">
<emphasis id="B9344D31F968FFA8FF555BED5B44A689" bold="true" box="[151,276,1949,1972]" pageId="7" pageNumber="256">FIGURE 3.</emphasis>
<taxonomicName id="4C40EAA0F968FFA8FED95BED5BB4A688" box="[283,484,1949,1971]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F968FFA8FED95BED5BB4A688" box="[283,484,1949,1971]" italics="true" pageId="7" pageNumber="256">Typhlocharis scrofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF968FFA8FE2E5BEE584FA688" box="[492,543,1950,1971]" pageId="7" pageNumber="256" rank="species">
<emphasis id="B9344D31F968FFA8FE2E5BEE584FA688" bold="true" box="[492,543,1950,1971]" pageId="7" pageNumber="256">sp.n.</emphasis>
</taxonomicNameLabel>
habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex.
</paragraph>
</caption>
<caption id="DF3FC1ABF967FFA7FF555FFB5811A2FB" httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="8" pageNumber="257" targetBox="[165,1406,187,861]" targetPageId="8">
<paragraph id="8BFF9123F967FFA7FF555FFB5811A2FB" blockId="8.[151,1435,907,961]" pageId="8" pageNumber="257">
<emphasis id="B9344D31F967FFA7FF555FFB5B43A29A" bold="true" box="[151,275,907,930]" pageId="8" pageNumber="257">FIGURE 4.</emphasis>
<taxonomicName id="4C40EAA0F967FFA7FEDB5FFC5BB1A299" box="[281,481,908,930]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F967FFA7FEDB5FFC5BB1A299" box="[281,481,908,930]" italics="true" pageId="8" pageNumber="257">Typhlocharis scrofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF967FFA7FE2A5FFC5871A29A" box="[488,545,908,929]" pageId="8" pageNumber="257" rank="species">
<emphasis id="B9344D31F967FFA7FE2A5FFC5871A29A" bold="true" box="[488,545,908,929]" pageId="8" pageNumber="257">sp. n.</emphasis>
</taxonomicNameLabel>
(A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view).
</paragraph>
</caption>
<paragraph id="8BFF9123F967FFA7FF055F9E5817A576" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">Legs: similar in both sexes. Intermetacoxal space slightly widened. Metacoxae with blunt triangular &quot;flap&quot;. Rounded metatrochanters, slightly angular pro- and metafemora, metatibiae with dilated distal region. Tarsi with four tarsomeres (tarsal formula 4-4-4).</paragraph>
<paragraph id="8BFF9123F967FFA7FF0558295FD9A582" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
Abdomen: microsculture and belt as in
<taxonomicName id="4C40EAA0F967FFA7FDBC582A5953A54A" box="[638,771,1113,1137]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F967FFA7FDBC582A5953A54A" box="[638,771,1113,1137]" italics="true" pageId="8" pageNumber="257">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF967FFA7FCCE58295913A54A" box="[780,835,1113,1137]" pageId="8" pageNumber="257" rank="species">
<emphasis id="B9344D31F967FFA7FCCE58295913A54A" bold="true" box="[780,835,1113,1137]" pageId="8" pageNumber="257">sp.n.</emphasis>
</taxonomicNameLabel>
Ventral foveae very faint to absent. Last ventrite with lateral teeth, pattern of six to seven pairs of setae in posterior margin, sexually dimorphic:
<emphasis id="B9344D31F967FFA7FB5D580D5FCCA5AE" bold="true" box="[1183,1436,1148,1173]" pageId="8" pageNumber="257">l-s-s-l-s-s / s-l-s-l-s-s-l</emphasis>
(
<figureCitation id="137B8DA6F967FFA7FF5D58D15AB4A582" box="[159,228,1185,1209]" captionStart="FIGURE 3" captionStartId="7.[151,250,1949,1972]" captionTargetBox="[176,1406,188,1906]" captionTargetId="figure@7.[155,1432,169,1928]" captionTargetPageId="7" captionText="FIGURE 3. Typhlocharis scrofa sp. n. habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222602/files/figure.png" pageId="8" pageNumber="257">Fig. 3</figureCitation>
). In females, it is characteristic that the second (
<emphasis id="B9344D31F967FFA7FCC358D15959A582" bold="true" box="[769,777,1185,1209]" pageId="8" pageNumber="257">l</emphasis>
) and third (
<emphasis id="B9344D31F967FFA7FC4F58D159C8A582" bold="true" box="[909,920,1185,1209]" pageId="8" pageNumber="257">s</emphasis>
) innermost setae are practically overlapped.
</paragraph>
<paragraph id="8BFF9123F967FFA7FF0558B55865A41E" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
Aedeagus: sickle-shaped median lobe (length:
<quantity id="4CB83CC6F967FFA7FD2358B55919A5E6" box="[737,841,1221,1245]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="257" unit="mm" value="0.14">0.14 mm</quantity>
) and broad, rounded apex, not bend in dorsal view (
<figureCitation id="137B8DA6F967FFA7FF5D58995AB7A43A" box="[159,231,1257,1281]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="8" pageNumber="257">Fig. 4</figureCitation>
A,B). Endophallus with short and thin &quot;parenthesis-shaped&quot; sclerites. Parameres subtriangular, with two medium-sized apical setae (
<figureCitation id="137B8DA6F967FFA7FE0F597D5841A41E" box="[461,529,1293,1317]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="8" pageNumber="257">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BFF9123F967FFA7FF0559415F01A48E" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
Female genitalia: club-shaped, robust gonocoxites with two apical setae: one large, one very small (
<figureCitation id="137B8DA6F967FFA7FAF759415F2BA472" box="[1333,1403,1329,1353]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="8" pageNumber="257">Fig. 4</figureCitation>
C). Two short lateral setae, pointing inwards, and scattered pores. Gonoduct with a thinner proximal region (diameter:
<quantity id="4CB83CC6F967FFA7FF5559095B42A4AA" box="[151,274,1401,1425]" metricMagnitude="-6" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="257" unit="mm" value="0.002">0.002 mm</quantity>
) and thicker distal region (diameter:
<quantity id="4CB83CC6F967FFA7FD235909590FA4AA" box="[737,863,1401,1425]" metricMagnitude="-6" metricUnit="m" metricValue="7" pageId="8" pageNumber="257" unit="mm" value="0.007">0.007 mm</quantity>
). Subsphaeric spermatheca (length:
<quantity id="4CB83CC6F967FFA7FAD059095FC0A4AA" box="[1298,1424,1401,1425]" metricMagnitude="-5" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="257" unit="mm" value="0.016">0.016 mm</quantity>
). Spermathecal gland tubular to conical (length:
<quantity id="4CB83CC6F967FFA7FD6659ED594AA48E" box="[676,794,1437,1461]" metricMagnitude="-5" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="257" unit="mm" value="0.028">0.028 mm</quantity>
), slender and sclerotized in distal region (
<figureCitation id="137B8DA6F967FFA7FB2E59ED5F60A48E" box="[1260,1328,1437,1461]" captionStart="FIGURE 4" captionStartId="8.[151,250,907,930]" captionTargetBox="[165,1406,187,861]" captionTargetId="figure@8.[151,1436,169,886]" captionTargetPageId="8" captionText="FIGURE 4. Typhlocharis scrofa sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222603/files/figure.png" pageId="8" pageNumber="257">Fig. 4</figureCitation>
C).
</paragraph>
<paragraph id="8BFF9123F967FFA7FF0559B05881A77E" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
<emphasis id="B9344D31F967FFA7FF0559B05B7AA4E2" bold="true" box="[199,298,1472,1497]" pageId="8" pageNumber="257">Habitat.</emphasis>
<taxonomicName id="4C40EAA0F967FFA7FEF659B25BC5A4E2" box="[308,405,1473,1497]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F967FFA7FEF659B25BC5A4E2" box="[308,405,1473,1497]" italics="true" pageId="8" pageNumber="257">T. scrofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF967FFA7FE6359B15B89A4E2" box="[417,473,1473,1497]" pageId="8" pageNumber="257" rank="species">
<emphasis id="B9344D31F967FFA7FE6359B15B89A4E2" bold="true" box="[417,473,1473,1497]" pageId="8" pageNumber="257">sp.n.</emphasis>
</taxonomicNameLabel>
was found in two localities separated
<quantity id="4CB83CC6F967FFA7FC5859B1598AA4E2" box="[922,986,1473,1497]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="257" unit="km" value="4.0">4 km</quantity>
: 1) Under embedded boulders of shale (Upper Cambrian) in open land with some holm oaks (
<taxonomicName id="4C40EAA0F967FFA7FCD6599659F4A4C6" box="[788,932,1509,1533]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="257" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B9344D31F967FFA7FCD6599659F4A4C6" box="[788,932,1509,1533]" italics="true" pageId="8" pageNumber="257">Quercus ilex</emphasis>
</taxonomicName>
L.), laudanum shrubs (
<taxonomicName id="4C40EAA0F967FFA7FB7659965F3AA4C6" box="[1204,1386,1509,1533]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="257" phylum="Tracheophyta" rank="species" species="ladanifer">
<emphasis id="B9344D31F967FFA7FB7659965F3AA4C6" box="[1204,1386,1509,1533]" italics="true" pageId="8" pageNumber="257">Cistus ladanifer</emphasis>
</taxonomicName>
L.), broom (
<emphasis id="B9344D31F967FFA7FF325A7A5B35A71A" box="[240,357,1546,1569]" italics="true" pageId="8" pageNumber="257">
<taxonomicName id="4C40EAA0F967FFA7FF325A7A5B16A71A" box="[240,326,1546,1569]" class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="257" phylum="Tracheophyta" rank="genus">Retama</taxonomicName>
sp
</emphasis>
. Raf.) and asphodels (
<emphasis id="B9344D31F967FFA7FDA35A795959A71A" box="[609,777,1545,1569]" italics="true" pageId="8" pageNumber="257">
<taxonomicName id="4C40EAA0F967FFA7FDA35A7958B2A71A" box="[609,738,1545,1569]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Asphodelus" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="257" phylum="Tracheophyta" rank="genus">Asphodelus</taxonomicName>
sp.
</emphasis>
L.). Brownish clayey soil. 2) In the slopes of a small river, with the same kind of environment and vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F967FFA6FF055A215877A3D4" lastPageId="9" lastPageNumber="258" pageId="8" pageNumber="257" type="etymology">
<paragraph id="8BFF9123F967FFA7FF055A215EB2A7B6" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
<emphasis id="B9344D31F967FFA7FF055A215B1EA752" bold="true" box="[199,334,1617,1641]" pageId="8" pageNumber="257">Etymology.</emphasis>
The new species is named after the black Iberian pig (
<emphasis id="B9344D31F967FFA7FC305A225E3AA752" box="[1010,1130,1617,1641]" italics="true" pageId="8" pageNumber="257">
Sus
<taxonomicName id="4C40EAA0F967FFA7FBE75A215E3AA752" box="[1061,1130,1617,1641]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="scrofa">scrofa</taxonomicName>
</emphasis>
L.), a common and very important component of the landscapes, culture and economy in the surrounds of the
<typeStatus id="54FB2F81F967FFA7FB935A065ED1A7B5" box="[1105,1153,1654,1678]" pageId="8" pageNumber="257">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BFF9123F967FFA7FF055AE859E0A65E" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
<emphasis id="B9344D31F967FFA7FF055AE85B1DA78A" bold="true" box="[199,333,1688,1713]" pageId="8" pageNumber="257">Variability.</emphasis>
The large series studied (
<specimenCount id="9D465AAAF967FFA7FDBB5AE95974A78A" box="[633,804,1689,1713]" pageId="8" pageNumber="257" type="generic">109 specimens</specimenCount>
) allowed a good perception of intraspecific variability. There are the usual minor alterations: chaetotaxy in labium and basilar; some variability in clypeus, usually straight but slightly arcuate in some specimens; development of pronotum denticles, more pronounced in some males than in the rest; or in the chaetotaxy of the last ventrite. Some individuals have a narrow hiatus. Proportions of pronotum are variable between shorter, almost square, and longer, more rectangular shapes. General body proportions vary, especially within males, from more robust to more slender individuals.
</paragraph>
<paragraph id="8BFF9123F967FFA7FF055B0158F8A696" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
Three specimens show asymmetric alteration in the pattern of
<emphasis id="B9344D31F967FFA7FC475B015E39A6B2" box="[901,1129,1905,1929]" italics="true" pageId="8" pageNumber="257">sensilla coeloconica</emphasis>
in last antennomere. Some individuals have longer paraglossae than usual.
</paragraph>
<paragraph id="8BFF9123F967FFA7FF055BC95F75A6CE" blockId="8.[151,1437,1005,2037]" pageId="8" pageNumber="257">
One male
<typeStatus id="54FB2F81F967FFA7FEF85BCA5BCAA6E9" box="[314,410,1978,2002]" pageId="8" pageNumber="257" type="paratype">paratype</typeStatus>
shows asymmetry in the sutural denticles of elytra (lacking one of the denticles). These are also slightly variable within the series, with some specimens lacking the characteristic &quot;V-shaped&quot; apex.
</paragraph>
<paragraph id="8BFF9123F966FFA6FF055CE75B19A1CC" blockId="9.[151,1436,151,752]" pageId="9" pageNumber="258">
There is some variation in the marking of the &quot;flap&quot; of metacoxae, but never reaching the degree observed in
<taxonomicName id="4C40EAA0F966FFA6FA455CE95B52A1EF" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FA455CE95B52A1EF" italics="true" pageId="9" pageNumber="258">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FECB5CCC5B11A1EF" box="[265,321,188,212]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FECB5CCC5B11A1EF" bold="true" box="[265,321,188,212]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
In contrast with the observed in
<taxonomicName id="4C40EAA0F966FFA6FD745CCD596DA1EF" box="[694,829,188,212]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FD745CCD596DA1EF" box="[694,829,188,212]" italics="true" pageId="9" pageNumber="258">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FC865CCC592BA1EF" box="[836,891,188,212]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FC865CCC592BA1EF" bold="true" box="[836,891,188,212]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
, the shape of apodemal ring is virtually constant in all the males.
</paragraph>
<paragraph id="8BFF9123F966FFA6FF055D745B02A07B" blockId="9.[151,1436,151,752]" pageId="9" pageNumber="258">
One of the male
<typeStatus id="54FB2F81F966FFA6FEBD5D755BBAA026" box="[383,490,261,285]" pageId="9" pageNumber="258" type="paratype">paratypes</typeStatus>
presents an abnormal three-setae paramere. The apex of the elytra is damaged in one female
<typeStatus id="54FB2F81F966FFA6FF285D585B1EA07B" box="[234,334,296,320]" pageId="9" pageNumber="258" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8BFF9123F966FFA6FF055D3B591EA307" blockId="9.[151,1436,151,752]" pageId="9" pageNumber="258">
<emphasis id="B9344D31F966FFA6FF055D3B5B6CA05F" bold="true" box="[199,316,331,356]" pageId="9" pageNumber="258">Affinities.</emphasis>
<taxonomicName id="4C40EAA0F966FFA6FE855D3D5BFAA05F" box="[327,426,332,356]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FE855D3D5BFAA05F" box="[327,426,332,356]" italics="true" pageId="9" pageNumber="258">T. scrofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FE755D3C5BBFA05F" box="[439,495,332,356]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FE755D3C5BBFA05F" bold="true" box="[439,495,332,356]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
is the fourth species known to have a diffuse gula along with
<taxonomicName id="4C40EAA0F966FFA6FB1F5D3D5B1DA0BC" authority="Coiffait, 1969" authorityName="Coiffait" authorityYear="1969" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F966FFA6FB1F5D3D5FCCA05F" box="[1245,1436,332,356]" italics="true" pageId="9" pageNumber="258">T. quadridentata</emphasis>
(
<bibRefCitation id="EFD1ECD2F966FFA6FF5C5D1F5B15A0BC" author="Coiffait" box="[158,325,367,391]" pageId="9" pageNumber="258" refString="Coiffait, H. (1969) Nouveaux Anillini du Maroc et Sud de la peninsule iberique. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 48, 55 - 56." type="journal article" year="1969">Coiffait, 1969</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C40EAA0F966FFA6FE9F5D0158A4A0BC" authority="Serrano &amp; Aguiar, 2008" authorityName="Serrano &amp; Aguiar" authorityYear="2008" box="[349,756,367,392]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F966FFA6FE9F5D015B9EA0B3" box="[349,462,369,392]" italics="true" pageId="9" pageNumber="258">T. crespoi</emphasis>
<bibRefCitation id="EFD1ECD2F966FFA6FE185D1F58A4A0BC" author="Serrano" box="[474,756,367,391]" pageId="9" pageNumber="258" refString="Serrano, A. R. M. &amp; Aguiar C. A. S. (2008) Two new species of Typhlocharis Dieck, 1869 (Coleoptera, Carabidae) from Portugal: Description and notes on related species. G raellsia, 64, 281 - 293. http: // dx. doi. org / 10.3989 / graellsia. 2008. v 64. i 2.38" type="journal article" year="2008">Serrano &amp; Aguiar, 2008</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C40EAA0F966FFA6FCF65D0159EDA0B3" box="[820,957,368,392]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FCF65D0159EDA0B3" box="[820,957,368,392]" italics="true" pageId="9" pageNumber="258">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FC085D005E52A0B3" box="[970,1026,368,392]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FC085D005E52A0B3" bold="true" box="[970,1026,368,392]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
, and the third with four tarsomere condition. It is also the smallest species known to date, with individuals of
<quantity id="4CB83CC6F966FFA6FC215DE45E19A097" box="[995,1097,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="258" unit="mm" value="0.86">0.86 mm</quantity>
, even more than the minute
<taxonomicName id="4C40EAA0F966FFA6FA455DE55B66A0EB" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F966FFA6FA455DE55B66A0EB" italics="true" pageId="9" pageNumber="258">T. quadridentata</emphasis>
</taxonomicName>
. It is very similar to
<taxonomicName id="4C40EAA0F966FFA6FDE45DC958FEA0EB" box="[550,686,440,464]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FDE45DC958FEA0EB" box="[550,686,440,464]" italics="true" pageId="9" pageNumber="258">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FD745DC858BEA0EB" box="[694,750,440,464]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FD745DC858BEA0EB" bold="true" box="[694,750,440,464]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
but immediately recognizable from it by the 4+1 umbilicate series (which would include it in the
<taxonomicName id="4C40EAA0F966FFA6FD825DAC58F4A0CF" box="[576,676,476,500]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="outereloi">
<emphasis id="B9344D31F966FFA6FD825DAC58F4A0CF" box="[576,676,476,500]" italics="true" pageId="9" pageNumber="258">outereloi</emphasis>
</taxonomicName>
group
<emphasis id="B9344D31F966FFA6FD355DAD5966A0CF" box="[759,822,477,500]" italics="true" pageId="9" pageNumber="258">sensu</emphasis>
<bibRefCitation id="EFD1ECD2F966FFA6FCFC5DAC5EE0A0CF" author="Zaballos" box="[830,1200,476,500]" pageId="9" pageNumber="258" refString="Zaballos, J. P. &amp; Ruiz-Tapiador, I. (1997) Nuevos Typhlocharis Dieck (Coleoptera: Caraboidea: Trechidae) de Espana. Graellsia, 52, 95 - 106. http: // dx. doi. org / 10.3989 / graellsia. 1996. v 52. i 0.381" type="journal article" year="1997">Zaballos &amp; Ruiz-Tapiador, 1997</bibRefCitation>
), instead of 4+3; the extremely faint lateral denticles of elytra, the presence of anterior discal setae (also present in
<taxonomicName id="4C40EAA0F966FFA6FB725E715F4DA323" box="[1200,1309,513,536]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F966FFA6FB725E715F4DA323" box="[1200,1309,513,536]" italics="true" pageId="9" pageNumber="258">T. crespoi</emphasis>
</taxonomicName>
) the shape of teeth of elytral apex, and the genitalia, among other details.
</paragraph>
<paragraph id="8BFF9123F966FFA6FF055E375877A3D4" blockId="9.[151,1436,151,752]" pageId="9" pageNumber="258">
Female genitalia is quite unique, and resembles an intermediate between the unguiform-like condition present in
<taxonomicName id="4C40EAA0F966FFA6FF755E1D5944A3BF" authority="Perez-Gonzalez &amp; Zaballos 2012" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2012" box="[183,788,620,644]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F966FFA6FF755E1D5B26A3BF" box="[183,374,620,644]" italics="true" pageId="9" pageNumber="258">T. quadridentata</emphasis>
(
<bibRefCitation id="EFD1ECD2F966FFA6FE4A5E1C595CA3BF" author="Perez-Gonzalez" box="[392,780,620,644]" pageId="9" pageNumber="258" refString="Perez-Gonzalez, S. &amp; Zaballos, J. P. (2012) Re-description of two species of Typhlocharis (Coleoptera: Carabidae: Anillini) and revision of the models of female genitalia within the genus. Zootaxa, 3279, 46 - 62." type="journal article" year="2012">Pérez-González &amp; Zaballos 2012</bibRefCitation>
)
</taxonomicName>
and the short tubular gonocoxites present in
<taxonomicName id="4C40EAA0F966FFA6FAE95E1D5BE9A39C" authority="Serrano &amp; Aguiar 2008" authorityName="Serrano &amp; Aguiar" authorityYear="2008" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F966FFA6FAE95E1D5FCBA3BF" box="[1323,1435,621,644]" italics="true" pageId="9" pageNumber="258">T. crespoi</emphasis>
(
<bibRefCitation id="EFD1ECD2F966FFA6FF5C5EFF5BE1A39C" author="Serrano" box="[158,433,655,679]" pageId="9" pageNumber="258" refString="Serrano, A. R. M. &amp; Aguiar C. A. S. (2008) Two new species of Typhlocharis Dieck, 1869 (Coleoptera, Carabidae) from Portugal: Description and notes on related species. G raellsia, 64, 281 - 293. http: // dx. doi. org / 10.3989 / graellsia. 2008. v 64. i 2.38" type="journal article" year="2008">Serrano &amp; Aguiar 2008</bibRefCitation>
)
</taxonomicName>
. Affinities with other species are the same as commented for
<taxonomicName id="4C40EAA0F966FFA6FBB85EE15F53A393" box="[1146,1283,656,680]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FBB85EE15F53A393" box="[1146,1283,656,680]" italics="true" pageId="9" pageNumber="258">T. baeturica</emphasis>
</taxonomicName>
<emphasis id="B9344D31F966FFA6FAC95EE05F1AA393" bold="true" box="[1291,1354,656,680]" pageId="9" pageNumber="258">
<taxonomicNameLabel id="A207F04AF966FFA6FAC95EE05F15A393" box="[1291,1349,656,680]" pageId="9" pageNumber="258" rank="species">sp.n.</taxonomicNameLabel>
,
</emphasis>
except the intermetacoxal space is narrower in
<taxonomicName id="4C40EAA0F966FFA6FD9E5EC558ECA3F7" box="[604,700,692,716]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="scrofa" status="sp. nov.">
<emphasis id="B9344D31F966FFA6FD9E5EC558ECA3F7" box="[604,700,692,716]" italics="true" pageId="9" pageNumber="258">T. scrofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF966FFA6FD045EC458AEA3F7" box="[710,766,692,716]" pageId="9" pageNumber="258" rank="species">
<emphasis id="B9344D31F966FFA6FD045EC458AEA3F7" bold="true" box="[710,766,692,716]" pageId="9" pageNumber="258">sp.n.</emphasis>
</taxonomicNameLabel>
and the aedeagus mostly closely resembles the broad apex present in
<taxonomicName id="4C40EAA0F966FFA6FECE5EA95BDBA3CB" box="[268,395,728,752]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="outereloi">
<emphasis id="B9344D31F966FFA6FECE5EA95BDBA3CB" box="[268,395,728,752]" italics="true" pageId="9" pageNumber="258">T. outereloi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C40EAA0F966FFA6FE035EA95870A3CB" box="[449,544,728,752]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="diecki">
<emphasis id="B9344D31F966FFA6FE035EA95870A3CB" box="[449,544,728,752]" italics="true" pageId="9" pageNumber="258">T. diecki</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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