140 lines
20 KiB
XML
140 lines
20 KiB
XML
<document id="F9DD171BA20D59F62DAA00B159CC175F" ID-CLB-Dataset="6167" ID-DOI="10.11646/zootaxa.4975.2.4" ID-GBIF-Dataset="3d8639f0-544a-4ca5-83a8-b729cf1c3e56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4925219" ID-ZooBank="4F22565A-7089-49FD-9313-63195491B3EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622022232820" checkinUser="plazi" docAuthor="Henry, R. & Bøggild, Esben" docDate="2021" docId="03E487AEFFCC7B0D78B7E28EFBFDADB1" docLanguage="en" docName="zootaxa.4975.2.4.pdf" docOrigin="Zootaxa 4975 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia culmenensis Henry & Bøggild 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="308" masterDocId="FFDDFFD6FFCE7B0F7820E251FFF9AE78" masterDocTitle="Nineteen new species of Megaselia Rondani (Diptera, Phoridae) from Denmark" masterLastPageNumber="342" masterPageNumber="306" pageNumber="308" updateTime="1698991867495" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="2A3E3E5FD52A5213D04807FA2265FD4E">
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<mods:title id="3026C0CC9DB68F94D08445EC7471AA54">Nineteen new species of Megaselia Rondani (Diptera, Phoridae) from Denmark</mods:title>
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<mods:name id="AF88B9D51B81FF0FD0F4EBB4F983DECD" type="personal">
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<mods:namePart id="A67649CF29F66AF3CE1B6AF580569033">Henry, R.</mods:namePart>
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<mods:affiliation id="DC43E1ACFEB216D7E8D015E432D250B5">University Department of Zoology, Downing Street, Cambridge CB 2 3 EJ, UK ..</mods:affiliation>
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<mods:roleTerm id="210F3313D073796D0A02BAEDCBC8AB42">Author</mods:roleTerm>
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<mods:namePart id="3F5B4019EF586ECC98249246587ADB43">Bøggild, Esben</mods:namePart>
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<mods:affiliation id="69C5E381D36D2220E1DAEBF3D02ABE48">Fayesgade 10, DK- 9500, Hobro, Denmark.</mods:affiliation>
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<mods:title id="8B98D3241002274C327C458F6CEEC3BC">Zootaxa</mods:title>
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<mods:part id="5E60543E051714B63EA62BAA4D4A8B50">
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<mods:date id="F108A8FDAD269845C4D81F02EDE6455C">2021</mods:date>
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<mods:number id="79F79511A5000E75E9D3B7288457E75C">2021-05-25</mods:number>
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<mods:number id="2F07F20A762320F169C23511159E88F4">4975</mods:number>
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<treatment id="03E487AEFFCC7B0D78B7E28EFBFDADB1" ID-DOI="http://doi.org/10.5281/zenodo.4925300" ID-GBIF-Taxon="182645730" ID-Zenodo-Dep="4925300" LSID="urn:lsid:plazi:treatment:03E487AEFFCC7B0D78B7E28EFBFDADB1" httpUri="http://treatment.plazi.org/id/03E487AEFFCC7B0D78B7E28EFBFDADB1" lastPageNumber="308" pageId="2" pageNumber="308">
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<subSubSection id="C3576533FFCC7B0D78B7E28EFE03AE81" box="[151,506,223,250]" pageId="2" pageNumber="308" type="nomenclature">
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<paragraph id="8BF236B8FFCC7B0D78B7E28EFE03AE81" blockId="2.[151,506,223,285]" box="[151,506,223,250]" pageId="2" pageNumber="308">
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<heading id="D0BA81D4FFCC7B0D78B7E28EFE03AE81" bold="true" box="[151,506,223,250]" fontSize="11" level="1" pageId="2" pageNumber="308" reason="1">
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<emphasis id="B939EAAAFFCC7B0D78B7E28EFE03AE81" bold="true" box="[151,506,223,250]" pageId="2" pageNumber="308">
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<taxonomicName id="4C4D4D3BFFCC7B0D78B7E28EFE49AE82" ID-CoL="9HWT7" authority="Henry & Bøggild, 2021" authorityName="Henry & Bøggild" authorityYear="2021" box="[151,432,223,250]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="308" phylum="Arthropoda" rank="species" species="culmenensis" status="sp. nov.">
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<emphasis id="B939EAAAFFCC7B0D78B7E28EFE49AE82" bold="true" box="[151,432,223,250]" italics="true" pageId="2" pageNumber="308">Megaselia culmenensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57D1FFCC7B0D7998E28EFE03AE81" box="[440,506,223,249]" pageId="2" pageNumber="308" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576533FFCC7B0D78B7E352FEE1AF65" box="[151,280,259,285]" pageId="2" pageNumber="308" type="description">
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<paragraph id="8BF236B8FFCC7B0D78B7E352FEE1AF65" blockId="2.[151,506,223,285]" box="[151,280,259,285]" pageId="2" pageNumber="308">
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(
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<figureCitation id="13762A3DFFCC7B0D78BFE352FEF6AF65" box="[159,271,259,285]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Figs 1–11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576533FFCC7B0D78B7E31AFABFAD25" pageId="2" pageNumber="308" type="diagnosis">
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<paragraph id="8BF236B8FFCC7B0D78B7E31AFB07AFA9" blockId="2.[151,1436,331,970]" pageId="2" pageNumber="308">
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<emphasis id="B939EAAAFFCC7B0D78B7E31AFEEBAF1D" bold="true" box="[151,274,331,357]" pageId="2" pageNumber="308">Diagnosis.</emphasis>
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In the keys of
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<bibRefCitation id="EFDC4B49FFCC7B0D79E5E31AFD70AF1D" author="Lundbeck, W." box="[453,649,331,357]" pageId="2" pageNumber="308" refId="ref14900" refString="Lundbeck, W. (1922) Diptera Danica Part VI Pipunculidae, Phoridae. Wesley, London, 447 pp." type="book" year="1922">Lundbeck (1922)</bibRefCitation>
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it runs to Group IV couplet 4, but its hypopygium is different from both options. In the key to the males of species from the British Isles (
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<bibRefCitation id="EFDC4B49FFCC7B0D7BE9E33EFB98AFF1" author="Disney R. H. L." box="[969,1121,367,393]" pageId="2" pageNumber="308" pagination="1 - 155" refId="ref14009" refString="Disney R. H. L. (1989) Scuttle Flies-Diptera Phoridae Genus Megaselia. Handbooks for the Identification of British Insects 10 (8), Royal Entomologicai Society of London, 1 - 155. https: // doi. org / 10.1163 / 187631287 X 00115" type="book chapter" year="1989">Disney, 1989</bibRefCitation>
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) it runs to couplet 121 and in
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<bibRefCitation id="EFDC4B49FFCC7B0D7896E3C2FE80AFD6" author="Schmitz, H." box="[182,377,403,430]" pageId="2" pageNumber="308" pagination="465 - 512" refId="ref15017" refString="Schmitz, H. (1958) Phoridae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region 4 (33) (Lieferung 202). E. Schweizerbartsche Verlagsbuchhandlung, Stuttgart, 465 - 512." type="book chapter" year="1958">Schmitz’s (1958)</bibRefCitation>
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key for Abteilung IV Erste Reihe it runs to couplet 18, but in both cases is excluded by its distinctive hypopygium. Likewise in Borgmeier’s (1964b) key Group IV it runs to couplet 20 lead 1.
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</paragraph>
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<paragraph id="8BF236B8FFCC7B0D78E7E38AFABFAD25" blockId="2.[151,1436,331,970]" pageId="2" pageNumber="308">
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<emphasis id="B939EAAAFFCC7B0D78E7E38AFE9BAF8D" bold="true" box="[199,354,475,501]" pageId="2" pageNumber="308">`Description.</emphasis>
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Male. Frons as
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<figureCitation id="13762A3DFFCC7B0D7A30E38AFDA9AF8D" box="[528,592,475,501]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 1</figureCitation>
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, with dense but very fine microtrichia. Cheek with 6 robust bristles and jowl with 2 that are longer and even more robust. Postpedicels, with a few SPS vesicles (
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<figureCitation id="13762A3DFFCC7B0D7C66E3AEFB73AC61" box="[1094,1162,511,537]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 3</figureCitation>
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), palps and proboscis as
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<figureCitation id="13762A3DFFCC7B0D78B7E072FF25AC45" box="[151,220,547,573]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 2</figureCitation>
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, the labella lacking short spinules below. Thorax (
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<figureCitation id="13762A3DFFCC7B0D7B36E072FCA3AC45" box="[790,858,547,573]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 4</figureCitation>
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) with 3 notopleural bristles and no cleft in front of these, mesopleuron with hairs and scutellum with an anterior pair of hairs and a posterior pair of bristles.Abdominal tergites as
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<figureCitation id="13762A3DFFCC7B0D7931E03AFEACACFE" box="[273,341,619,646]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 5</figureCitation>
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. Venter brown, and with hairs on segments 3–6. Hypopygium as
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<figureCitation id="13762A3DFFCC7B0D7C1AE03AFB59ACFD" box="[1082,1184,619,645]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Figs 6–8</figureCitation>
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. Legs brown but front leg more yellowish brown. Fore tarsus (
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<figureCitation id="13762A3DFFCC7B0D7A44E0DEFD53ACD1" box="[612,682,655,681]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 9</figureCitation>
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) with posterodorsal hair palisades on segments 1–5 and 5 clearly longer than 4. Dorsal hair palisade of mid tibia extends about 0.78 times its length. Hairs below basal half of hind femur subequal to those of anteroventral row of outer half (
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<figureCitation id="13762A3DFFCC7B0D7B06E086FC81AC89" box="[806,888,727,753]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 10</figureCitation>
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). Hind tibia with 7–8 differentiated posterodorsal bristles, without anterodorsals, and spinules of apical comb simple. Wings (
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<figureCitation id="13762A3DFFCC7B0D7BD9E0AAFBB2AD6D" box="[1017,1099,763,789]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="2" pageNumber="308">Fig. 11</figureCitation>
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)
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<quantity id="4CB59B5DFFCC7B0D7C7DE0AAFB3AAD6E" box="[1117,1219,763,790]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="308" unit="mm" value="1.14">1.14 mm</quantity>
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long. Costal index 0.49. Costal ratios 4: 2.3: 1. Costal cilia (of section 3)
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<quantity id="4CB59B5DFFCC7B0D7ADBE14EFCA6AD42" box="[763,863,799,826]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="308" unit="mm" value="0.07">0.07 mm</quantity>
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||
long. Vein 3 with hair (but only its sockets present on both wings). 2 axillary bristles, the outermost being
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<quantity id="4CB59B5DFFCC7B0D7B23E112FC9EAD26" box="[771,871,835,862]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="308" unit="mm" value="0.07">0.07 mm</quantity>
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long. Sc not reaching R1. Halteree brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576533FFCC7B0D78E7E136FEAFADDD" pageId="2" pageNumber="308" type="materials_examined">
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<paragraph id="8BF236B8FFCC7B0D78E7E136FEAFADDD" blockId="2.[151,1436,331,970]" pageId="2" pageNumber="308">
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<emphasis id="B939EAAAFFCC7B0D78E7E136FE96ADF9" bold="true" box="[199,367,871,897]" pageId="2" pageNumber="308">Type material</emphasis>
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.
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<materialsCitation id="3B253CE5FFCC7B0D7952E136FEAFADDD" ID-GBIF-Occurrence="3128593311" collectingDate="2017-07-24" collectingDateMax="2017-08-08" collectingDateMin="2017-07-24" collectionCode="DK, EJ, UCMZ" country="Denmark" location="Norre Onsild Mose" pageId="2" pageNumber="308" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F6881AFFCC7B0D7952E136FE27ADF9" box="[370,478,871,897]" pageId="2" pageNumber="308" type="holotype">Holotype</typeStatus>
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male,
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<collectingCountry id="F35A7628FFCC7B0D7A0BE136FD40ADF9" box="[555,697,871,897]" name="Denmark" pageId="2" pageNumber="308">DENMARK</collectingCountry>
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,
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<collectionCode id="ED5CAE7DFFCC7B0D7AE7E139FD16ADFA" box="[711,751,872,898]" pageId="2" pageNumber="308">DK</collectionCode>
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||
<collectionCode id="ED5CAE7DFFCC7B0D7AD7E139FCEFADFA" box="[759,790,872,898]" country="Israel" name="Ein Yabrud collection catalogue entries at The Hebrew University" pageId="2" pageNumber="308">EJ</collectionCode>
|
||
,
|
||
<location id="8E926063FFCC7B0D7B02E139FC06ADFA" LSID="urn:lsid:plazi:treatment:03E487AEFFCC7B0D78B7E28EFBFDADB1:8E926063FFCC7B0D7B02E139FC06ADFA" box="[802,1023,871,898]" country="Denmark" name="Norre Onsild Mose" pageId="2" pageNumber="308">Nørre Onsild Mose</location>
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||
,
|
||
<date id="FFF31078FFCC7B0D7C2BE136FB10ADF9" box="[1035,1257,871,897]" pageId="2" pageNumber="308" value="2017-07-24" valueMax="2017-08-08" valueMin="2017-07-24">
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<collectingDate id="EFB7E990FFCC7B0D7C2BE136FB10ADF9" box="[1035,1257,871,897]" pageId="2" pageNumber="308" value="2017-07-24" valueMax="2017-08-08" valueMin="2017-07-24">24.VII–8.VIII.2017</collectingDate>
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||
</date>
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||
, Esben Bøggild (
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||
<collectionCode id="ED5CAE7DFFCC7B0D78BFE1DAFF0FADDD" box="[159,246,907,933]" pageId="2" pageNumber="308">UCMZ</collectionCode>
|
||
—9–14).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576533FFCC7B0D78E7E1FEFBFDADB1" box="[199,1028,943,970]" pageId="2" pageNumber="308" type="etymology">
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||
<paragraph id="8BF236B8FFCC7B0D78E7E1FEFBFDADB1" blockId="2.[151,1436,331,970]" box="[199,1028,943,970]" pageId="2" pageNumber="308">
|
||
<emphasis id="B939EAAAFFCC7B0D78E7E1FEFEB1ADB1" bold="true" box="[199,328,943,969]" pageId="2" pageNumber="308">Etymology</emphasis>
|
||
. Named after unusually short roof (culmen) of the epandrium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |