treatments-xml/data/03/DD/33/03DD3329EF665852FC71FBF4C7A65DA8.xml

<document ID-DOI="10.5252/cryptogamie-mycologie2019v40a5" ID-ISSN="1776-100X" ID-Zenodo-Dep="7814899" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1681174563244" checkinUser="felipe" docAuthor="Wang, Xiang-Hua, Das, Kanad, Bera, Ishika, Chen, Yu-Hui, Bhatt, Rajendra Prasad, Ghosh, Aniket, Hembrom, Manoj Emanuel, Hofstetter, Valérie, Parihar, Arvind, Vizzini, Alfredo, Xu, Tai-Min &amp; Zhao, Chang-Lin" docDate="2019" docId="03DD3329EF665852FC71FBF4C7A65DA8" docLanguage="en" docName="CryptogamieMycologie.40.5.57-95.pdf" docOrigin="Cryptogamie, Mycologie 20 (5)" docSource="https://bioone.org/journals/cryptogamie-mycologie/volume-40/issue-5/cryptogamie-mycologie2019v40a5/Fungal-Biodiversity-Profiles-81-90/10.5252/cryptogamie-mycologie2019v40a5.full" docStyle="DocumentStyle:81F49AB8619EB7E5D11A6328441E2653.1:CryptogamieMycologie.2019-.journal_article" docStyleId="81F49AB8619EB7E5D11A6328441E2653" docStyleName="CryptogamieMycologie.2019-.journal_article" docStyleVersion="1" docTitle="Russula capillaris Buyck 2019, sp. nov." docType="treatment" docVersion="4" lastPageNumber="83" masterDocId="FFE44B51EF7E584EFFD4FFE0C4635A44" masterDocTitle="Fungal Biodiversity Profiles 81 - 90" masterLastPageNumber="95" masterPageNumber="57" pageNumber="79" updateTime="1681391969980" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Fungal Biodiversity Profiles 81 - 90</mods:title>
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<mods:namePart>Wang, Xiang-Hua</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia,) xhwang @ mail. kib. ac. cn</mods:affiliation>
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<mods:namePart>Bera, Ishika</mods:namePart>
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<mods:namePart>Chen, Yu-Hui</mods:namePart>
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<mods:namePart>Ghosh, Aniket</mods:namePart>
<mods:affiliation>Department of Botany &amp; Microbiology, H. N. B. Garhwal University (A Central University), Srinagar, Garhwal, 246174, Uttarakhand (India) bhatt. rajendra 123 @ gmail. com ghosh. aniket 87 @ gmail. com</mods:affiliation>
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<mods:namePart>Hofstetter, Valérie</mods:namePart>
<mods:affiliation>Department of Plant Protection, Agroscope Changins-Wädenswil - ACW, Rte De Duiller, CH- 1260 Nyon 1 (Switzerland) valerie. hofstetter @ agroscope. admin. ch</mods:affiliation>
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88. 
<taxonomicName ID-CoL="4TRQD" ID-ENA="2579157" authority="Buyck" authorityName="Buyck" authorityYear="2019" box="[975,1242,1044,1071]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="24" pageNumber="79" phylum="Basidiomycota" rank="species" species="capillaris" status="sp. nov.">
<emphasis bold="true" box="[975,1164,1044,1071]" italics="true" pageId="24" pageNumber="79">Russula capillaris</emphasis>
Buyck
</taxonomicName>
, 
<taxonomicNameLabel box="[1253,1335,1045,1071]" pageId="24" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection pageId="24" pageNumber="79" type="description">
<paragraph blockId="24.[1067,1200,1076,1103]" box="[1067,1200,1076,1103]" pageId="24" pageNumber="79">
<heading box="[1067,1200,1076,1103]" centered="true" fontSize="11" level="3" pageId="24" pageNumber="79" reason="3">
(
<figureCitation box="[1075,1191,1076,1103]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="25.[132,143,1985,2003]" captionStartId-1="26.[132,143,1962,1980]" captionStartId-2="27.[132,143,1962,1980]" captionTargetBox-0="[132,1455,230,1878]" captionTargetBox-1="[184,1389,213,1920]" captionTargetBox-2="[132,1364,176,1945]" captionTargetId-0="figure-56@25.[132,1455,215,1878]" captionTargetId-1="figure-88@26.[184,1452,182,1927]" captionTargetId-2="figure-98@27.[132,1455,132,1953]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIG. 16. — Russula capillaris Buyck, sp. nov. (holotype), morphology of fresh basidioma: A, detail of pileus and stipe surface; B, section showing stipe interior; C, details of gills. Photos: B. Buyck." captionText-1="FIG. 17. — Russula capillaris Buyck, sp. nov. (holotype), microscopic features of pileipellis: A, pileocystidia near the trama-subpellis transition with schematic contents in one and indication of refringent conglomerates in both other cells; B, detail of zebroid incrustations observed on most subpellis and context hyphae; C, hyphal extremities of the pileus surface. Scale bar: 10 µm. Drawings: B. Buyck." captionText-2="FIG. 18. — Russula capillaris Buyck, sp. nov. (holotype), microscopic features of the hymenophore: A, spores; B, basidia and basidiola; C, marginal cells of the gill edge; D, gloeocystidia on gill sides; E, gloeocystidia on gill edge. Cystidial contents are mostly schematic, note also the presence of incrusting material that is easily observed near the base of many pleurogloeocystidia. Scale bar: 10 µm, but only 5 µm for spores. Drawings: B. Buyck." figureDoi-0="http://doi.org/10.5281/zenodo.7814929" figureDoi-1="http://doi.org/10.5281/zenodo.7814931" figureDoi-2="http://doi.org/10.5281/zenodo.7814933" httpUri-0="https://zenodo.org/record/7814929/files/figure.png" httpUri-1="https://zenodo.org/record/7814931/files/figure.png" httpUri-2="https://zenodo.org/record/7814933/files/figure.png" pageId="24" pageNumber="79">Figs 16-18</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="24.[811,1457,1133,1395]" pageId="24" pageNumber="79">
Well-characterized by the combination of its medium-sized, firm stature, white and hard stipe, smooth and cream to pale yellowish brown pileus, nearly globose spores, (6.9)7.13-7.48-7.8(8.1) - (6.4)6.6-6.95-7.2(7.5) Μm, Q = (1.03)1.05-1.08-1.11(1.13), with very low, dense spore ornamentation of minute warts and invisible suprahilar spot, a pileipellis composed of ascending, slender, irregularly wavy-undulate hyphal extremities that are often inflated at the tip, submerged pileocystidia, pale spore print and slightly unpleasant smell, as well as by its habitat in tapia (
<taxonomicName box="[1183,1314,1345,1369]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="24" pageNumber="79" phylum="Tracheophyta" rank="species" species="bojeri">
<emphasis box="[1183,1314,1345,1369]" italics="true" pageId="24" pageNumber="79">Uapaca bojeri</emphasis>
</taxonomicName>
) woodland on the Central Plateau in 
<collectingCountry box="[1032,1149,1371,1395]" name="Madagascar" pageId="24" pageNumber="79">Madagascar</collectingCountry>
.
</paragraph>
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<subSubSection pageId="24" pageNumber="79" type="materials_examined">
<paragraph blockId="24.[811,1456,1419,1496]" pageId="24" pageNumber="79">
<materialsCitation collectingDate="2008-01-26" collectionCode="PC" collectorName="Buyck" country="Madagascar" location="Antsirabe" municipality="Central Plateau" pageId="24" pageNumber="79" specimenCode="PC0124764" specimenCount="1" typeStatus="typus">
<typeStatus box="[813,871,1419,1442]" pageId="24" pageNumber="79">TYPUS</typeStatus>
. — 
<emphasis bold="true" box="[908,1020,1419,1443]" pageId="24" pageNumber="79">
<collectingCountry box="[908,1020,1419,1443]" name="Madagascar" pageId="24" pageNumber="79">Madagascar</collectingCountry>
</emphasis>
. 
<collectingMunicipality box="[1031,1175,1419,1443]" pageId="24" pageNumber="79">Central Plateau</collectingMunicipality>
, along RN7 between 
<location LSID="urn:lsid:plazi:treatment:03DD3329EF665852FC71FBF4C7A65DA8:8EABD4E4EF665856FA83FA6BC1CC5FE7" box="[1367,1455,1419,1443]" country="Madagascar" municipality="Central Plateau" name="Antsirabe" pageId="24" pageNumber="79">Antsirabe</location>
and 
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, at km 40, in secondary 
<taxonomicName box="[1179,1306,1445,1469]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="24" pageNumber="79" phylum="Tracheophyta" rank="species" species="bojeri">
<emphasis box="[1179,1306,1445,1469]" italics="true" pageId="24" pageNumber="79">Uapaca bojeri</emphasis>
</taxonomicName>
woodland with 
<taxonomicName authorityName="Perry" authorityYear="1811" box="[811,864,1473,1496]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="24" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis box="[811,864,1473,1496]" italics="true" pageId="24" pageNumber="79">Pinus</emphasis>
</taxonomicName>
ingression, 
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<collectingDate box="[978,1077,1472,1496]" pageId="24" pageNumber="79" value="2008-01-26">26.I.2008</collectingDate>
</date>
, 
<emphasis box="[1085,1258,1472,1496]" italics="true" pageId="24" pageNumber="79">
563/ 
<collectorName box="[1129,1185,1472,1496]" pageId="24" pageNumber="79">Buyck</collectorName>
08.172
</emphasis>
(
<collectionCode box="[1272,1305,1472,1496]" pageId="24" pageNumber="79">PC</collectionCode>
[
<specimenCode box="[1311,1439,1472,1496]" pageId="24" pageNumber="79">PC0124764</specimenCode>
]).
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</paragraph>
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<paragraph blockId="24.[813,1053,1520,1544]" box="[813,1053,1520,1544]" pageId="24" pageNumber="79">MYCOBANK. — 832120.</paragraph>
<paragraph blockId="24.[813,1456,1567,1644]" pageId="24" pageNumber="79">
GENBANK. — 
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(
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),
</materialsCitation>
<materialsCitation box="[1148,1335,1567,1591]" collectionCode="LSU" pageId="24" pageNumber="79" specimenCount="1" typeStatus="holotype">
<accessionNumber box="[1148,1260,1567,1591]" pageId="24" pageNumber="79">KU237553</accessionNumber>
(
<collectionCode box="[1274,1324,1567,1591]" pageId="24" pageNumber="79">LSU</collectionCode>
),
</materialsCitation>
<materialsCitation pageId="24" pageNumber="79" specimenCount="1" typeStatus="holotype">
<accessionNumber box="[1343,1455,1567,1591]" pageId="24" pageNumber="79">KU237399</accessionNumber>
(mitSSU),
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<materialsCitation box="[924,1126,1594,1618]" collectionCode="RPB" pageId="24" pageNumber="79" specimenCode="RPB2" specimenCount="1" typeStatus="holotype">
<accessionNumber box="[924,1036,1594,1618]" pageId="24" pageNumber="79">KU237839</accessionNumber>
(
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),
</materialsCitation>
<materialsCitation pageId="24" pageNumber="79" specimenCount="1" typeStatus="holotype">
<accessionNumber box="[1134,1247,1594,1618]" pageId="24" pageNumber="79">KU237982</accessionNumber>
(tef1alpha), all from 
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.
</materialsCitation>
</paragraph>
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<subSubSection pageId="24" pageNumber="79" type="etymology">
<paragraph blockId="24.[813,1455,1668,1718]" pageId="24" pageNumber="79">ETYMOLOGY. — named after the resemblance of the pileipellis structure with a nice hair cut.</paragraph>
</subSubSection>
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<paragraph blockId="24.[813,957,1778,1804]" box="[813,957,1778,1804]" pageId="24" pageNumber="79">
<heading box="[813,957,1778,1804]" fontSize="9" level="3" pageId="24" pageNumber="79" reason="5">DESCRIPTION</heading>
</paragraph>
<paragraph blockId="24.[813,1150,1810,1867]" box="[813,922,1810,1836]" pageId="24" pageNumber="79">
<heading box="[813,922,1810,1836]" fontSize="11" level="3" pageId="24" pageNumber="79" reason="9">
<emphasis box="[813,922,1810,1836]" italics="true" pageId="24" pageNumber="79">Basidioma</emphasis>
</heading>
</paragraph>
<paragraph blockId="24.[813,1150,1810,1867]" box="[813,1150,1841,1867]" pageId="24" pageNumber="79">Growing single, medium-sized.</paragraph>
<paragraph blockId="24.[813,874,1905,1931]" box="[813,874,1905,1931]" pageId="24" pageNumber="79">
<heading box="[813,874,1905,1931]" fontSize="11" level="3" pageId="24" pageNumber="79" reason="9">
<emphasis box="[813,874,1905,1931]" italics="true" pageId="24" pageNumber="79">Pileus</emphasis>
</heading>
</paragraph>
<paragraph blockId="24.[813,1455,1937,2028]" lastBlockId="28.[132,777,215,369]" lastPageId="28" lastPageNumber="83" pageId="24" pageNumber="79">
<quantity box="[813,893,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="24" pageNumber="79" unit="mm" value="83.0">83 mm</quantity>
diam., quite regular in shape to slightly wavy near the margin, hardly depressed in the center, not becoming deeply infundibuliform, smooth near margin; surface glabrous, hardly separable, not viscid, probably even not when wet, continuous and smooth in the pileus center, dull, pale-colored, from whitish to cream or pale yellowish to pale yellowish brown, not concentrically zonated, not particularly darker in the center.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814929" ID-Zenodo-Dep="7814929" httpUri="https://zenodo.org/record/7814929/files/figure.png" pageId="25" pageNumber="80" startId="25.[132,143,1985,2003]" targetBox="[132,1455,230,1878]" targetPageId="25" targetType="figure">
<paragraph blockId="25.[132,1457,1985,2025]" pageId="25" pageNumber="80">
FIG. 16. — 
<taxonomicName authority="Buyck" authorityName="Buyck" authorityYear="2019" box="[224,428,1985,2003]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="25" pageNumber="80" phylum="Basidiomycota" rank="species" species="capillaris" status="sp. nov.">
<emphasis box="[224,367,1985,2003]" italics="true" pageId="25" pageNumber="80">Russula capillaris</emphasis>
Buyck
</taxonomicName>
, 
<taxonomicNameLabel box="[436,503,1985,2003]" pageId="25" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
(holotype), morphology of fresh basidioma: 
<emphasis bold="true" box="[872,885,1985,2003]" pageId="25" pageNumber="80">A</emphasis>
, detail of pileus and stipe surface; 
<emphasis bold="true" box="[1180,1193,1985,2003]" pageId="25" pageNumber="80">B</emphasis>
, section showing stipe interior; 
<emphasis bold="true" box="[132,146,2007,2025]" pageId="25" pageNumber="80">C</emphasis>
, details of gills. Photos: B. Buyck.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814931" ID-Zenodo-Dep="7814931" httpUri="https://zenodo.org/record/7814931/files/figure.png" pageId="26" pageNumber="81" startId="26.[132,143,1962,1980]" targetBox="[184,1389,213,1920]" targetPageId="26" targetType="figure">
<paragraph blockId="26.[132,1457,1962,2025]" pageId="26" pageNumber="81">
FIG. 17. — 
<taxonomicName authority="Buyck" authorityName="Buyck" authorityYear="2019" box="[225,431,1962,1980]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="26" pageNumber="81" phylum="Basidiomycota" rank="species" species="capillaris" status="sp. nov.">
<emphasis box="[225,369,1962,1980]" italics="true" pageId="26" pageNumber="81">Russula capillaris</emphasis>
Buyck
</taxonomicName>
, 
<taxonomicNameLabel box="[440,508,1962,1980]" pageId="26" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
(holotype), microscopic features of pileipellis: 
<emphasis bold="true" box="[899,912,1962,1980]" pageId="26" pageNumber="81">A</emphasis>
, pileocystidia near the trama-subpellis transition with schematic contents in one and indication of refringent conglomerates in both other cells; 
<emphasis bold="true" box="[779,792,1985,2003]" pageId="26" pageNumber="81">B</emphasis>
, detail of zebroid incrustations observed on most subpellis and context hyphae; 
<emphasis bold="true" box="[132,146,2007,2025]" pageId="26" pageNumber="81">C</emphasis>
, hyphal extremities of the pileus surface. Scale bar: 10 µm. Drawings: B. Buyck.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814933" ID-Zenodo-Dep="7814933" httpUri="https://zenodo.org/record/7814933/files/figure.png" pageId="27" pageNumber="82" startId="27.[132,143,1962,1980]" targetBox="[132,1364,176,1945]" targetPageId="27" targetType="figure">
<paragraph blockId="27.[132,1455,1962,2025]" pageId="27" pageNumber="82">
FIG. 18. — 
<taxonomicName authority="Buyck" authorityName="Buyck" authorityYear="2019" box="[224,428,1962,1980]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="27" pageNumber="82" phylum="Basidiomycota" rank="species" species="capillaris" status="sp. nov.">
<emphasis box="[224,367,1962,1980]" italics="true" pageId="27" pageNumber="82">Russula capillaris</emphasis>
Buyck
</taxonomicName>
, 
<taxonomicNameLabel box="[436,503,1962,1980]" pageId="27" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
(holotype), microscopic features of the hymenophore: 
<emphasis bold="true" box="[959,972,1962,1980]" pageId="27" pageNumber="82">A</emphasis>
, spores; 
<emphasis bold="true" box="[1050,1063,1962,1980]" pageId="27" pageNumber="82">B</emphasis>
, basidia and basidiola; 
<emphasis bold="true" box="[1262,1276,1962,1980]" pageId="27" pageNumber="82">C</emphasis>
, marginal cells of the gill edge; 
<emphasis bold="true" box="[213,227,1985,2003]" pageId="27" pageNumber="82">D</emphasis>
, gloeocystidia on gill sides; 
<emphasis bold="true" box="[461,473,1985,2003]" pageId="27" pageNumber="82">E</emphasis>
, gloeocystidia on gill edge. Cystidial contents are mostly schematic, note also the presence of incrusting material that is easily observed near the base of many pleurogloeocystidia. Scale bar: 10 µm, but only 5 µm for spores. Drawings: B. Buyck.
</paragraph>
</caption>
<paragraph blockId="28.[132,180,406,432]" box="[132,180,406,432]" pageId="28" pageNumber="83">
<heading box="[132,180,406,432]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,180,406,432]" italics="true" pageId="28" pageNumber="83">Gills</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,775,438,560]" pageId="28" pageNumber="83">
Adnate, with frequent lamellulae but less than normal gills, not or rarely forking, 
<emphasis box="[366,383,471,496]" italics="true" pageId="28" pageNumber="83">c.</emphasis>
<quantity box="[390,478,470,497]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="28" pageNumber="83" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6-7 mm</quantity>
high, thin and fragile, normally spaced (
<emphasis box="[291,308,503,528]" italics="true" pageId="28" pageNumber="83">c.</emphasis>
1L+l/mm), pale yellowish; gill edge even, concolorous.
</paragraph>
<paragraph blockId="28.[132,182,598,624]" box="[132,182,598,624]" pageId="28" pageNumber="83">
<heading box="[132,182,598,624]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,182,598,624]" italics="true" pageId="28" pageNumber="83">Stipe</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,775,630,752]" pageId="28" pageNumber="83">
Central, 43 × 
<quantity box="[295,381,630,656]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="83" unit="mm" value="16.0">16 mm</quantity>
, not annulate, subcylindrical, longitudinally wrinkled, glabrous, off-white, surprisingly hard and firm considering the interior is developing up to four cavities.
</paragraph>
<paragraph blockId="28.[132,213,790,816]" box="[132,213,790,816]" pageId="28" pageNumber="83">
<heading box="[132,213,790,816]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,213,790,816]" italics="true" pageId="28" pageNumber="83">Context</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,775,821,911]" pageId="28" pageNumber="83">
Firm, fleshy, c. 
<quantity box="[303,370,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="83" unit="mm" value="9.0">9 mm</quantity>
thick above gill attachment to stipe, whitish, unchanged when cut, in the stipe with a faint creamish-pinkish tinge.
</paragraph>
<paragraph blockId="28.[132,184,949,974]" box="[132,184,949,974]" pageId="28" pageNumber="83">
<heading box="[132,184,949,974]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,184,949,974]" italics="true" pageId="28" pageNumber="83">Taste</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,285,980,1007]" box="[132,285,980,1007]" pageId="28" pageNumber="83">Not observed.</paragraph>
<paragraph blockId="28.[132,186,1044,1070]" box="[132,186,1044,1070]" pageId="28" pageNumber="83">
<heading box="[132,186,1044,1070]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,186,1044,1070]" italics="true" pageId="28" pageNumber="83">Odor</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,351,1076,1102]" box="[132,351,1076,1102]" pageId="28" pageNumber="83">Faintly disagreeable.</paragraph>
<paragraph blockId="28.[132,247,1140,1166]" box="[132,247,1140,1166]" pageId="28" pageNumber="83">
<heading box="[132,247,1140,1166]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,247,1140,1166]" italics="true" pageId="28" pageNumber="83">Spore print</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,440,1172,1198]" box="[132,440,1172,1198]" pageId="28" pageNumber="83">Unsufficient, obviously pale.</paragraph>
<paragraph blockId="28.[132,195,1236,1262]" box="[132,195,1236,1262]" pageId="28" pageNumber="83">
<heading box="[132,195,1236,1262]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,195,1236,1262]" italics="true" pageId="28" pageNumber="83">Spores</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,775,1267,1422]" pageId="28" pageNumber="83">Globose to subglobose, (6.9)7.13-7.48-7.8(8.1) × (6.4)6.6- 6.95-7.2(7.5) Μm, Q = (1.03)1.05-1.08-1.11(1.13); ornamentation consisting of a minute, dense, granular-reticulate pattern, &lt;0.1-0.2 Μm high and weakly amyloid; suprahilar spot invisible; apiculus very short and small.</paragraph>
<paragraph blockId="28.[132,209,1459,1485]" box="[132,209,1459,1485]" pageId="28" pageNumber="83">
<heading box="[132,209,1459,1485]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[132,209,1459,1485]" italics="true" pageId="28" pageNumber="83">Basidia</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[132,776,1491,1549]" pageId="28" pageNumber="83">Slender, widest in their upper part, 53-61 × 8-10 Μm, fourspored; sterigmata rather slender, approximately 7 × 1-2 Μm.</paragraph>
<paragraph blockId="28.[131,366,1586,1612]" box="[131,366,1586,1612]" pageId="28" pageNumber="83">
<heading box="[131,366,1586,1612]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[131,366,1586,1612]" italics="true" pageId="28" pageNumber="83">Hymenial gloeocystidia</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[130,777,1618,2028]" pageId="28" pageNumber="83">Very abundant (c. 4000/mm2), thin-walled or nearly so, not strongly emergent, on the gill sides, 60-90 × 8-9 Μm, fusiform to subclavate or almost cylindrical, frequently distinctly inflated at the apex and widely capitate (often wider than the rest of the cystidium), some also tapering, near the apex obtuse-rounded or more frequently minutely mucronate, containing poor contents in the form of dispersed, refringent masses or conglomerates; the basal part of the cystidia often sheathed with encrusting material; near the gill edge distinctly smaller and mostly simply clavate to almost subcylindrical, 34-46 × 6-7(-8) Μm, and then nearly subcylindrical to clavulate, obtuse rounded at the tip, filled with more abundant, granular-crystalline contents, weakly reacting to SV.</paragraph>
<paragraph blockId="28.[813,962,215,241]" box="[813,962,215,241]" pageId="28" pageNumber="83">
<heading box="[813,962,215,241]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[813,962,215,241]" italics="true" pageId="28" pageNumber="83">Subhymenium</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[811,1455,247,306]" pageId="28" pageNumber="83">Dense, composed of cells that are hardly wider than the basal parts of the basidia.</paragraph>
<paragraph blockId="28.[813,975,343,369]" box="[813,975,343,369]" pageId="28" pageNumber="83">
<heading box="[813,975,343,369]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[813,975,343,369]" italics="true" pageId="28" pageNumber="83">Lamellar trama</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[813,1164,375,401]" box="[813,1164,375,401]" pageId="28" pageNumber="83">With sphaerocytes, rather dense.</paragraph>
<paragraph blockId="28.[813,958,438,464]" box="[813,958,438,464]" pageId="28" pageNumber="83">
<heading box="[813,958,438,464]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[813,958,438,464]" italics="true" pageId="28" pageNumber="83">Marginal cells</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[811,1456,470,624]" pageId="28" pageNumber="83">Well-differentiated, thin-walled, smaller than basidia, consisting of a basal or stalk cell holding a terminal fusiform, clavate or lageniform cell that is often terminally inflated, 22-39 × 5-9 Μm, occupying the whole gill edge, optically empty, mixed with small gloeocystidia.</paragraph>
<paragraph blockId="28.[813,912,662,688]" box="[813,912,662,688]" pageId="28" pageNumber="83">
<heading box="[813,912,662,688]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[813,912,662,688]" italics="true" pageId="28" pageNumber="83">Pileipellis</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[811,1458,693,1390]" pageId="28" pageNumber="83">
Orthochromatic in cresyl blue, poorly delimited from the underlying trama, one-layered to obscurely two-layered, forming a very dense tissue up to 200 Μm thick of closely packed hyphae (but easily dissolving in KOH); the lower hyphae intertwined-intricate, gradually more to very irregularly inflated, strongly branching, emitting densely packed, vertical to ascending hyphal extremities; these 
<emphasis box="[1312,1329,886,911]" italics="true" pageId="28" pageNumber="83">c.</emphasis>
4 Μm wide, thin-walled, irregularly undulate-twisted, composed of 2-3 subcylindrical cells with the second or third cell often branching and giving rise to two or more terminal cells; these even more irregularly undulate or wavy in outline, not exceeding 60 Μm long, often distinctly inflated near the apex. Subpellis composed of cylindrical, distinctly zebroid encrusted hyphae. Pileocystidia quite common to abundant below the pileipellis, but apparently absent from the pileus surface; immerged, subcylindrical, thin-walled gloeocystidia are very apparent immediately below the subpellis because of their poor, yet very visible contents in the form of dispersed, refringent masses of variable size and because of their relatively important size compared to the surrounding hyphae and other elements, continuing but less abundant in trama underneath.
</paragraph>
<paragraph blockId="28.[813,1007,1427,1453]" box="[813,1007,1427,1453]" pageId="28" pageNumber="83">
<heading box="[813,1007,1427,1453]" fontSize="11" level="3" pageId="28" pageNumber="83" reason="9">
<emphasis box="[813,1007,1427,1453]" italics="true" pageId="28" pageNumber="83">Clamp connections</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[813,1031,1459,1485]" box="[813,1031,1459,1485]" pageId="28" pageNumber="83">Absent in all tissues.</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="83" type="discussion">
<paragraph blockId="28.[813,887,1523,1549]" box="[813,887,1523,1549]" pageId="28" pageNumber="83">
<heading box="[813,887,1523,1549]" fontSize="9" level="3" pageId="28" pageNumber="83" reason="5">NOTES</heading>
</paragraph>
<paragraph blockId="28.[811,1456,1554,2028]" pageId="28" pageNumber="83">
This apparently very rare species – we have only seen it once in more than twenty years of fungal exploration in Africa and 
<collectingCountry box="[811,938,1618,1644]" name="Madagascar" pageId="28" pageNumber="83">Madagascar</collectingCountry>
– has been placed by a recently published multigene phylogeny of 
<taxonomicName authority="(Buyck et al. 2018)" baseAuthorityName="Buyck" baseAuthorityYear="2018" box="[1010,1295,1650,1677]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="28" pageNumber="83" phylum="Basidiomycota" rank="genus">
<emphasis box="[1010,1088,1650,1676]" italics="true" pageId="28" pageNumber="83">Russula</emphasis>
(
<bibRefCitation author="BUYCK B. &amp; ZOLLER S. &amp; HOFSTETTER V." box="[1103,1285,1650,1676]" pageId="28" pageNumber="83" pagination="267 - 292" refId="ref20232" refString="BUYCK B., ZOLLER S. &amp; HOFSTETTER V. 2018. - Walking the thin line … ten years later: the dilemma of above- versus below-ground features to support phylogenies in the Russulaceae (Basidiomycota). Fungal Diversity 89 (1): 267 - 292. https: // doi. org / 10.1007 / s 13225 - 018 - 0397 - 5" type="journal article" year="2018">
Buyck 
<emphasis box="[1176,1226,1650,1676]" italics="true" pageId="28" pageNumber="83">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
in 
<emphasis box="[1326,1350,1651,1676]" italics="true" pageId="28" pageNumber="83">R.</emphasis>
subgenus 
<emphasis box="[813,914,1682,1708]" italics="true" pageId="28" pageNumber="83">Malodora</emphasis>
Buyck &amp; V. Hofst., where it occupies a more or less isolated position, belonging neither to section 
<emphasis box="[1283,1453,1714,1740]" italics="true" pageId="28" pageNumber="83">Pseudocompactae</emphasis>
Buyck &amp; V. Hofst. nor to section 
<emphasis box="[1179,1246,1746,1772]" italics="true" pageId="28" pageNumber="83">Edules</emphasis>
Buyck &amp; V. Hofst. (see 
<bibRefCitation author="DAS K. &amp; GHOSH A. &amp; BHATT R. P. &amp; CHAKRABORTY D. &amp; HOFSTETTER V. &amp; BUYCK B." box="[860,1023,1778,1804]" pageId="28" pageNumber="83" pagination="527 - 548" refId="ref20530" refString="DAS K., GHOSH A., BHATT R. P., CHAKRABORTY D., HOFSTETTER V. &amp; BUYCK B. 2017. - Fungal Biodiversity Profiles 41 - 50. Cryptogamie, Mycologie 38 (4): 527 - 548. https: // doi. org / 10.7872 / crym / v 38. iss 4.2017.527" type="journal article" year="2017">
Das 
<emphasis box="[909,960,1778,1804]" italics="true" pageId="28" pageNumber="83">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph blockId="28.[811,1456,1554,2028]" pageId="28" pageNumber="83">
Both in the field and under the microscope, it reminds rather of some species formerly placed in 
<emphasis box="[1272,1296,1842,1867]" italics="true" pageId="28" pageNumber="83">R.</emphasis>
subg. 
<taxonomicName authority="Buyck &amp; V. Hofst." authorityName="Buyck &amp; V. Hofst." class="Arachnida" family="Archaeidae" genus="Archaea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis box="[1369,1455,1841,1867]" italics="true" pageId="28" pageNumber="83">Archaea</emphasis>
Buyck &amp;V. Hofst.
</taxonomicName>
, in particular the North American 
<taxonomicName authority="Buyck" authorityName="Buyck" box="[813,1029,1905,1931]" class="Agaricomycetes" family="Russulaceae" genus="Russula" kingdom="Fungi" order="Russulales" pageId="28" pageNumber="83" phylum="Basidiomycota" rank="species" species="fattoensis">
<emphasis box="[813,946,1905,1931]" italics="true" pageId="28" pageNumber="83">R. fattoensis</emphasis>
Buyck
</taxonomicName>
, because of the very similar structure and composition of the pileipellis. It differs nevertheless from all species placed in subg. 
<taxonomicName box="[1228,1316,1969,1995]" class="Arachnida" family="Archaeidae" genus="Archaea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis box="[1228,1316,1969,1995]" italics="true" pageId="28" pageNumber="83">Archaea</emphasis>
</taxonomicName>
in its much larger spores.
</paragraph>
</subSubSection>
</treatment>
</document>