treatments-xml/data/03/DC/56/03DC56194C5B3D2F81F9FF002D95C62E.xml
2024-06-21 12:22:17 +02:00

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<document id="7B0AE0F46696155F72A6431A9D454D97" ID-CLB-Dataset="45400" ID-DOI="10.11646/zootaxa.3821.4.3" ID-GBIF-Dataset="26306b5b-ed89-4007-888f-7cb564bfa282" ID-ISSN="1175-5326" ID-Zenodo-Dep="252865" ID-ZooBank="1249EDEE-817D-45E3-B564-26A55411A508" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460442192791" checkinUser="plazi" docAuthor="Norrbom, Allen L. &amp; Neder, Lilia Estela" docDate="2014" docId="03DC56194C5B3D2F81F9FF002D95C62E" docLanguage="en" docName="zt03821p456.pdf" docOrigin="Zootaxa 3821 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trupanea Schrank" docType="treatment" docVersion="9" lastPageNumber="444" masterDocId="FFE52E614C5A3D2E816EFF982800C007" masterDocTitle="New neotropical species of Trupanea (Diptera: Tephritidae) with unusual wing patterns" masterLastPageNumber="456" masterPageNumber="443" pageNumber="444" updateTime="1698333012524" updateUser="ExternalLinkService">
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<mods:title id="F80CC0B961770480172AF6BF472F1787">New neotropical species of Trupanea (Diptera: Tephritidae) with unusual wing patterns</mods:title>
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<mods:namePart id="FEDBBB9A9427B7F0D5FF17A46F7E0690">Norrbom, Allen L.</mods:namePart>
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<mods:namePart id="3B086231E722A311E06FCC97536794AA">Neder, Lilia Estela</mods:namePart>
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<treatment id="03DC56194C5B3D2F81F9FF002D95C62E" ID-DOI="http://doi.org/10.5281/zenodo.6135357" ID-GBIF-Taxon="119508209" ID-Zenodo-Dep="6135357" LSID="urn:lsid:plazi:treatment:03DC56194C5B3D2F81F9FF002D95C62E" httpUri="http://treatment.plazi.org/id/03DC56194C5B3D2F81F9FF002D95C62E" lastPageNumber="444" pageId="1" pageNumber="444">
<subSubSection id="C36FB4844C5B3D2F81F9FF00297EC0B5" box="[151,382,152,178]" pageId="1" pageNumber="444" type="nomenclature">
<paragraph id="8BCAE70F4C5B3D2F81F9FF00297EC0B5" blockId="1.[151,382,152,178]" box="[151,382,152,178]" pageId="1" pageNumber="444">
<heading id="D08250634C5B3D2F81F9FF00297EC0B5" bold="true" box="[151,382,152,178]" fontSize="11" level="1" pageId="1" pageNumber="444" reason="1">
<taxonomicName id="4C759C8C4C5B3D2F81F9FF00297EC0B5" ID-CoL="9G2TM" authority="Schrank" authorityName="Schrank" box="[151,382,152,178]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FF00297EC0B5" bold="true" box="[151,382,152,178]" pageId="1" pageNumber="444">
<emphasis id="B9013B1D4C5B3D2F81F9FF00290DC0B5" bold="true" box="[151,269,152,178]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
Schrank
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36FB4844C5B3D2F81F9FF782D95C62E" pageId="1" pageNumber="444" type="reference_group">
<paragraph id="8BCAE70F4C5B3D2F81F9FF782C73C0F3" blockId="1.[151,1367,222,306]" box="[151,1139,222,245]" pageId="1" pageNumber="444">
<taxonomicName id="4C759C8C4C5B3D2F81F9FF7829C0C0F3" ID-CoL="9G2TM" authority="Schrank 1795: 147" authorityName="Schrank" authorityPageNumber="147" authorityYear="1795" box="[151,448,222,245]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FF7828F8C0F2" box="[151,248,224,245]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
Schrank 1795: 147
</taxonomicName>
.
<typeStatus id="54CE59AD4C5B3D2F80A4FF4729FCC0F3" box="[458,508,223,244]" pageId="1" pageNumber="444">Type</typeStatus>
species—
<emphasis id="B9013B1D4C5B3D2F830BFF462AB0C0F3" box="[613,688,222,244]" italics="true" pageId="1" pageNumber="444">radiata</emphasis>
Schrank (=
<taxonomicName id="4C759C8C4C5B3D2F8242FF462BD5C0F3" ID-CoL="7CWKF" authority="Fuesslin" authorityName="Fuesslin" box="[812,981,222,244]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="stellata">
<emphasis id="B9013B1D4C5B3D2F8242FF462B77C0F3" box="[812,887,222,244]" italics="true" pageId="1" pageNumber="444">stellata</emphasis>
Fuesslin
</taxonomicName>
), by monotypy.
</paragraph>
<paragraph id="8BCAE70F4C5B3D2F81F9FF652D57C135" blockId="1.[151,1367,222,306]" pageId="1" pageNumber="444">
<treatmentCitationGroup id="AB65C0214C5B3D2F81F9FF652D57C135" pageId="1" pageNumber="444">
<taxonomicName id="4C759C8C4C5B3D2F81F9FF6529F8C114" ID-CoL="3K23" authority="Hendel 1914a: 86" authorityName="Hendel" authorityPageNumber="86" authorityYear="1914" box="[151,504,253,275]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FF65293CC114" box="[151,316,253,275]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
<treatmentCitation id="0AD4C11E4C5B3D2F802CFF6529F8C114" author="Hendel" box="[322,504,253,275]" page="86" pageId="1" pageNumber="444" year="1914">
<bibRefCitation id="EFE49AFE4C5B3D2F802CFF6529D3C114" author="Hendel" box="[322,467,253,275]" pageId="1" pageNumber="444" refString="Hendel, F. G. (1914 a) Die Gattungen der Bohrfliegen. (Analytische Ubersicht aller bisher bekannten Gattungen der Tephritinae.). Wiener Entomologische Zeitung, 33, 73 - 98." type="journal article" year="1914" yearSuffix="a">Hendel 1914a</bibRefCitation>
: 86
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54CE59AD4C5B3D2F836CFF662A34C114" box="[514,564,254,275]" pageId="1" pageNumber="444">Type</typeStatus>
species—
<taxonomicName id="4C759C8C4C5B3D2F83F3FF652B4EC114" ID-CoL="S3JN" authority="Hendel" authorityName="Hendel" box="[669,846,253,275]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F83F3FF652AFBC114" box="[669,763,253,275]" italics="true" pageId="1" pageNumber="444">maculata</emphasis>
Hendel
</taxonomicName>
, by original designation. New synonym.
<taxonomicName id="4C759C8C4C5B3D2F81F9FE8429E2C135" authority="Hering 1944: 14" authorityName="Hering" authorityPageNumber="14" authorityYear="1944" box="[151,482,284,306]" class="Insecta" family="Tephritidae" genus="Melanotrypana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FE842933C135" box="[151,307,284,306]" italics="true" pageId="1" pageNumber="444">Melanotrypana</emphasis>
Hering 1944: 14
</taxonomicName>
.
<typeStatus id="54CE59AD4C5B3D2F8082FE852A1EC135" box="[492,542,285,306]" pageId="1" pageNumber="444">Type</typeStatus>
species—
<taxonomicName id="4C759C8C4C5B3D2F83E9FE842BBDC135" ID-CoL="87LM7" authority="Seguy" authorityName="Seguy" box="[647,957,284,306]" class="Insecta" family="Tephritidae" genus="Acanthiophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="benoisti">
<emphasis id="B9013B1D4C5B3D2F83E9FE842B76C135" box="[647,886,284,306]" italics="true" pageId="1" pageNumber="444">Acanthiophilus benoisti</emphasis>
Séguy
</taxonomicName>
, by original designation. New synonym.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCAE70F4C5B3D2F81F9FECF2B78C28C" blockId="1.[151,1437,343,1577]" pageId="1" pageNumber="444">
<taxonomicName id="4C759C8C4C5B3D2F81F9FECF294DC177" ID-CoL="3K23" box="[151,333,343,368]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FECF294DC177" box="[151,333,343,368]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation id="EFE49AFE4C5B3D2F8377FECF2AC8C177" author="Hendel" box="[537,712,343,368]" pageId="1" pageNumber="444" refString="Hendel, F. G. (1914 a) Die Gattungen der Bohrfliegen. (Analytische Ubersicht aller bisher bekannten Gattungen der Tephritinae.). Wiener Entomologische Zeitung, 33, 73 - 98." type="journal article" year="1914" yearSuffix="a">Hendel (1914a)</bibRefCitation>
in a key to genera of
<taxonomicName id="4C759C8C4C5B3D2F82D1FECF2C41C177" ID-CoL="GZN" box="[959,1089,343,368]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
, and again by
<bibRefCitation id="EFE49AFE4C5B3D2F858AFECF2D97C177" author="Hendel" box="[1252,1431,343,368]" pageId="1" pageNumber="444" refString="Hendel, F. (1914 b) Die Bohrfliegen Sudamerikas. Ubersicht und Katalog der bisher aus der neotropischen Region beschriebenen Tephritinen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden, 14 (3), 1 - 84." type="journal article" year="1914" yearSuffix="b">Hendel (1914b)</bibRefCitation>
, with
<taxonomicName id="4C759C8C4C5B3D2F81BEFEE429B3C193" ID-CoL="S3JN" authority="Hendel" authorityName="Hendel" box="[208,435,379,404]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F81BEFEE4295AC193" box="[208,346,379,404]" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
Hendel
</taxonomicName>
from
<collectingCountry id="F362A79F4C5B3D2F8097FEE32A36C193" box="[505,566,379,404]" name="Chile" pageId="1" pageNumber="444">Chile</collectingCountry>
as
<typeStatus id="54CE59AD4C5B3D2F8333FEE42A8DC193" box="[605,653,380,404]" pageId="1" pageNumber="444">type</typeStatus>
species. It was not compared with
<taxonomicName id="4C759C8C4C5B3D2F8573FEE42C88C194" ID-CoL="9G2TM" box="[1053,1160,380,403]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F8573FEE42C88C194" box="[1053,1160,380,403]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
, but of the characters in the description and key, the main difference is that the wing pattern was “spotted, rather than stellate as in
<taxonomicName id="4C759C8C4C5B3D2F81F9FE5B2902C1DD" ID-CoL="9G2TM" box="[151,258,451,474]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FE5B2902C1DD" box="[151,258,451,474]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE49AFE4C5B3D2F8061FE5B29A0C1DC" author="Foote" box="[271,416,451,475]" pageId="1" pageNumber="444" refString="Foote, R. H. (1980) Fruit fly genera south of the United States (Diptera: Tephritidae). United States Department of Agriculture Technical Bulletin, No. 1600, iv + 79 pp." type="book" year="1980">Foote (1980)</bibRefCitation>
also considered the wing pattern significant and included species with a solid dark mark on the pterostigma and the area posterior to it, and a rectangular hyaline area distal to it. He included the
<taxonomicName id="4C759C8C4C5B3D2F845FFE7F2928C225" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="diespasmena">
<emphasis id="B9013B1D4C5B3D2F845FFE7F2928C225" italics="true" pageId="1" pageNumber="444">Trupanea diespasmena</emphasis>
</taxonomicName>
group sensu Aczél (1953) in the synonymy of
<taxonomicName id="4C759C8C4C5B3D2F825BFD912BEBC225" ID-CoL="3K23" box="[821,1003,521,546]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F825BFD912BEBC225" box="[821,1003,521,546]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
, although he did not list the individual species. He also considered
<taxonomicName id="4C759C8C4C5B3D2F808FFDB42B3EC242" authority="Hering (1944)" authorityName="Hering" authorityYear="1944" box="[481,830,556,581]" class="Insecta" family="Tephritidae" genus="Melanotrypana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F808FFDB42A8EC242" box="[481,654,556,581]" italics="true" pageId="1" pageNumber="444">Melanotrypana</emphasis>
Hering (1944)
</taxonomicName>
, with
<typeStatus id="54CE59AD4C5B3D2F82E5FDB52BBBC242" box="[907,955,557,581]" pageId="1" pageNumber="444">type</typeStatus>
species
<taxonomicName id="4C759C8C4C5B3D2F854FFDB42D80C242" ID-CoL="87LM7" authority="Seguy" authorityName="Seguy" box="[1057,1408,556,581]" class="Insecta" family="Tephritidae" genus="Acanthiophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="benoisti">
<emphasis id="B9013B1D4C5B3D2F854FFDB42D2FC242" box="[1057,1327,556,581]" italics="true" pageId="1" pageNumber="444">Acanthiophilus benoisti</emphasis>
Séguy
</taxonomicName>
, a synonym of
<taxonomicName id="4C759C8C4C5B3D2F804BFDC829DBC26E" ID-CoL="3K23" box="[293,475,592,617]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F804BFDC829DBC26E" box="[293,475,592,617]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
, although the latter species has a somewhat different, more interrupted wing pattern. Aczél (1953) considered
<taxonomicName id="4C759C8C4C5B3D2F80DFFDEB2A5EC28B" box="[433,606,627,652]" class="Insecta" family="Tephritidae" genus="Melanotrypana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F80DFFDEB2A5EC28B" box="[433,606,627,652]" italics="true" pageId="1" pageNumber="444">Melanotrypana</emphasis>
</taxonomicName>
a subgenus of
<taxonomicName id="4C759C8C4C5B3D2F826BFDED2B71C28B" ID-CoL="9G2TM" box="[773,881,629,652]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F826BFDED2B71C28B" box="[773,881,629,652]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCAE70F4C5B3D2F81A9FD0F2B2DC4ED" blockId="1.[151,1437,343,1577]" pageId="1" pageNumber="444">
The
<typeStatus id="54CE59AD4C5B3D2F8197FD002929C2B7" box="[249,297,664,688]" pageId="1" pageNumber="444">type</typeStatus>
species of
<emphasis id="B9013B1D4C5B3D2F80C9FD0F2A52C2B7" box="[423,594,663,688]" italics="true" pageId="1" pageNumber="444">Celidophenella</emphasis>
,
<taxonomicName id="4C759C8C4C5B3D2F830EFD002AE9C2B7" ID-CoL="S3JN" box="[608,745,663,688]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F830EFD002AE9C2B7" box="[608,745,663,688]" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
</taxonomicName>
, possesses the character states considered synapomorphies of the
<taxonomicName id="4C759C8C4C5B3D2F81AFFD24292CC2D4" ID-CoL="9G2TM" box="[193,300,700,723]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81AFFD24292CC2D4" box="[193,300,700,723]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
clade as recognized by
<bibRefCitation id="EFE49AFE4C5B3D2F8353FD232ACDC2D4" author="Merz" box="[573,717,699,723]" pageId="1" pageNumber="444" refString="Merz, B. (1999) Phylogeny of the Palearctic and Afrotropical genera of the Tephritis group (Tephritinae: Tephritini). In: Aluja, M. &amp; Norrbom, A. L. (Eds.), Fruit flies (Tephritidae): Phylogeny and evolution of behavior. CRC Press, Boca Raton, pp. 629 - 669." type="book chapter" year="1999">Merz (1999)</bibRefCitation>
. The acrophallus of the glans in the male terminalia is reduced; and the glans has a single, strongly sclerotized, movable, hooklike sclerite. Except perhaps in wing pattern,
<taxonomicName id="4C759C8C4C5B3D2F84ECFD4728FFC31D" ID-CoL="S3JN" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F84ECFD4728FFC31D" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
</taxonomicName>
otherwise falls within the traditional concept of
<taxonomicName id="4C759C8C4C5B3D2F825FFC9A2B9DC31E" ID-CoL="9G2TM" box="[817,925,770,793]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F825FFC9A2B9DC31E" box="[817,925,770,793]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE49AFE4C5B3D2F82C3FC9A2C76C31D" author="Foote" box="[941,1142,769,794]" pageId="1" pageNumber="444" refString="Foote, R. H., Blanc, F. L. &amp; Norrbom, A. L. (1993) Handbook of the fruit flies (Diptera: Tephritidae) of America north of Mexico. Comstock Publishing Associates, Ithaca, xii + 571 pp." type="book" year="1993">
Foote
<emphasis id="B9013B1D4C5B3D2F8296FC9A2C33C31D" box="[1016,1075,769,794]" italics="true" pageId="1" pageNumber="444">et al.</emphasis>
1993
</bibRefCitation>
,
<bibRefCitation id="EFE49AFE4C5B3D2F85EAFC992D31C31D" author="Norrbom" box="[1156,1329,769,794]" pageId="1" pageNumber="444" refString="Norrbom, A. L. (2010) Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. &amp; Zumbado, M. A. (Eds.), Manual of Central American Diptera, volume 2. NRC Research Press, Ottawa, pp. 909 - 954." type="journal article" year="2010">Norrbom 2010</bibRefCitation>
). It has 3 frontal setae, 2 orbital setae, and the apical scutellar setae are absent. The body is densely gray microtrichose. In
<taxonomicName id="4C759C8C4C5B3D2F84ECFCBE28FFC366" ID-CoL="S3JN" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F84ECFCBE28FFC366" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
</taxonomicName>
the apical rays of the wing pattern are reduced to spots at the apices of veins R4+5 and M, but its wing pattern certainly can be considered to be a modified stellate pattern. Its pattern is intermediate between typical
<taxonomicName id="4C759C8C4C5B3D2F81F9FC0B2903C3AD" ID-CoL="9G2TM" box="[151,259,915,938]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FC0B2903C3AD" box="[151,259,915,938]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
species and those of some of the other species that
<bibRefCitation id="EFE49AFE4C5B3D2F8223FC0B2BD8C3AC" author="Foote" box="[845,984,914,939]" pageId="1" pageNumber="444" refString="Foote, R. H. (1967) Family Tephritidae (Trypetidae, Trupaneidae). In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Fascicle 57. Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo., 91 pp." type="book" year="1967">Foote (1967</bibRefCitation>
,
<bibRefCitation id="EFE49AFE4C5B3D2F828BFC0A2C1EC3AD" author="Foote" box="[997,1054,914,938]" pageId="1" pageNumber="444" refString="Foote, R. H. (1980) Fruit fly genera south of the United States (Diptera: Tephritidae). United States Department of Agriculture Technical Bulletin, No. 1600, iv + 79 pp." type="book" year="1980">1980</bibRefCitation>
) included in
<taxonomicName id="4C759C8C4C5B3D2F85DAFC0A2D6AC3AC" ID-CoL="3K23" box="[1204,1386,914,939]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F85DAFC0A2D6AC3AC" box="[1204,1386,914,939]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
that have a more extensively dark pattern. The stellate pattern was considered by
<bibRefCitation id="EFE49AFE4C5B3D2F8283FC2E2C79C3C9" author="Merz" box="[1005,1145,950,974]" pageId="1" pageNumber="444" refString="Merz, B. (1999) Phylogeny of the Palearctic and Afrotropical genera of the Tephritis group (Tephritinae: Tephritini). In: Aluja, M. &amp; Norrbom, A. L. (Eds.), Fruit flies (Tephritidae): Phylogeny and evolution of behavior. CRC Press, Boca Raton, pp. 629 - 669." type="book chapter" year="1999">Merz (1999)</bibRefCitation>
a synapomorphy either of
<taxonomicName id="4C759C8C4C5B3D2F81F9FC422903C3F6" ID-CoL="9G2TM" box="[151,259,986,1009]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FC422903C3F6" box="[151,259,986,1009]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
or a higher taxon within the
<taxonomicName id="4C759C8C4C5B3D2F830EFC412AC3C3F5" ID-CoL="8YQP3" box="[608,707,985,1010]" class="Insecta" family="Tephritidae" genus="Tephritis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F830EFC412AC3C3F5" box="[608,707,985,1010]" italics="true" pageId="1" pageNumber="444">Tephritis</emphasis>
</taxonomicName>
group of genera in which
<taxonomicName id="4C759C8C4C5B3D2F856EFC422C6BC3F6" ID-CoL="9G2TM" box="[1024,1131,986,1009]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F856EFC422C6BC3F6" box="[1024,1131,986,1009]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
is included. Although the majority of species of
<taxonomicName id="4C759C8C4C5B3D2F80CFFC662A0CC412" ID-CoL="9G2TM" box="[417,524,1022,1045]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F80CFFC662A0CC412" box="[417,524,1022,1045]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
have a stellate pattern, there is considerable variation, particularly among the Neotropical species, and therefore we do not exclude those that otherwise fit the concept of the genus in chaetotaxy and genitalic characters. We have observed no characters that indicate that
<taxonomicName id="4C759C8C4C5B3D2F829AFBDD2C60C45B" ID-CoL="9G2TM" box="[1012,1120,1093,1116]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F829AFBDD2C60C45B" box="[1012,1120,1093,1116]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
without the inclusion of
<taxonomicName id="4C759C8C4C5B3D2F84ECFBDC28FFC478" ID-CoL="S3JN" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F84ECFBDC28FFC478" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
</taxonomicName>
and similar species is a monophyletic group, and we therefore consider
<taxonomicName id="4C759C8C4C5B3D2F853DFBFE2D09C478" ID-CoL="3K23" box="[1107,1289,1126,1151]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F853DFBFE2D09C478" box="[1107,1289,1126,1151]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
a subjective junior synonym of
<taxonomicName id="4C759C8C4C5B3D2F80E3FB1329F8C4A5" ID-CoL="9G2TM" box="[397,504,1163,1186]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F80E3FB1329F8C4A5" box="[397,504,1163,1186]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
. Although we have not examined
<taxonomicName id="4C759C8C4C5B3D2F82D4FB132C96C4A4" ID-CoL="87LM7" authority="Seguy" authorityName="Seguy" box="[954,1174,1162,1187]" class="Insecta" family="Tephritidae" genus="Acanthiophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="benoisti">
<emphasis id="B9013B1D4C5B3D2F82D4FB132C3DC4A4" box="[954,1085,1162,1187]" italics="true" pageId="1" pageNumber="444">A. benoisti</emphasis>
Séguy
</taxonomicName>
, the
<typeStatus id="54CE59AD4C5B3D2F858EFB132D10C4A4" box="[1248,1296,1163,1187]" pageId="1" pageNumber="444">type</typeStatus>
species of
<taxonomicName id="4C759C8C4C5B3D2F81F9FB352943C4C1" box="[151,323,1197,1222]" class="Insecta" family="Tephritidae" genus="Melanotrypana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FB352943C4C1" box="[151,323,1197,1222]" italics="true" pageId="1" pageNumber="444">Melanotrypana</emphasis>
</taxonomicName>
, it is consistent with
<taxonomicName id="4C759C8C4C5B3D2F8353FB372AA8C4C1" ID-CoL="9G2TM" box="[573,680,1199,1222]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F8353FB372AA8C4C1" box="[573,680,1199,1222]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
in chaetotaxy and probably falls within the concept of the genus. Study of its terminalia is needed to confirm this synonymy.
</paragraph>
<paragraph id="8BCAE70F4C5B3D2F81A9FB6E2C80C57F" blockId="1.[151,1437,343,1577]" pageId="1" pageNumber="444">
<bibRefCitation id="EFE49AFE4C5B3D2F81A9FB6E2959C50A" author="Merz" box="[199,345,1269,1293]" pageId="1" pageNumber="444" refString="Merz, B. (1999) Phylogeny of the Palearctic and Afrotropical genera of the Tephritis group (Tephritinae: Tephritini). In: Aluja, M. &amp; Norrbom, A. L. (Eds.), Fruit flies (Tephritidae): Phylogeny and evolution of behavior. CRC Press, Boca Raton, pp. 629 - 669." type="book chapter" year="1999">Merz (1999)</bibRefCitation>
considered the reduced size of the lower calypter to be a synapomorphy of
<taxonomicName id="4C759C8C4C5B3D2F859CFB6E2D5EC50A" ID-CoL="9G2TM" box="[1266,1374,1270,1293]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F859CFB6E2D5EC50A" box="[1266,1374,1270,1293]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
</taxonomicName>
. The calypter is no larger in
<taxonomicName id="4C759C8C4C5B3D2F80F5FA812A24C536" ID-CoL="S3JN" box="[411,548,1304,1329]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F80F5FA812A24C536" box="[411,548,1304,1329]" italics="true" pageId="1" pageNumber="444">C. maculata</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C759C8C4C5B3D2F83EFFA812B67C536" ID-CoL="7CWKF" authority="Fuesslin" authorityName="Fuesslin" box="[641,871,1304,1329]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="stellata">
<emphasis id="B9013B1D4C5B3D2F83EFFA812AEFC536" box="[641,751,1304,1329]" italics="true" pageId="1" pageNumber="444">T. stellata</emphasis>
(Fuesslin)
</taxonomicName>
or
<taxonomicName id="4C759C8C4C5B3D2F82E2FA812CAEC536" authority="Efflatoun" authorityName="Efflatoun" box="[908,1198,1304,1329]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="B9013B1D4C5B3D2F82E2FA812C28C536" box="[908,1064,1304,1329]" italics="true" pageId="1" pageNumber="444">T. desertorum</emphasis>
(Efflatoun)
</taxonomicName>
, the species included in his matrix, however, it is similar in size in the other
<taxonomicName id="4C759C8C4C5B3D2F8390FAA42B6CC552" ID-CoL="8YQ96" box="[766,876,1340,1365]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="tribe" tribe="Tephritini">Tephritini</taxonomicName>
genera that we compared (
<taxonomicName id="4C759C8C4C5B3D2F85F3FAA52CFCC553" ID-CoL="62R4P" box="[1181,1276,1341,1364]" class="Insecta" family="Tephritidae" genus="Dioxyna" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F85F3FAA52CFCC553" box="[1181,1276,1341,1364]" italics="true" pageId="1" pageNumber="444">Dioxyna</emphasis>
</taxonomicName>
,
<taxonomicName id="4C759C8C4C5B3D2F8464FAA52D95C553" ID-CoL="494X" box="[1290,1429,1341,1364]" class="Insecta" family="Tephritidae" genus="Dyseuaresta" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F8464FAA52D95C553" box="[1290,1429,1341,1364]" italics="true" pageId="1" pageNumber="444">Dyseuaresta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C759C8C4C5B3D2F81F9FAC72922C57F" ID-CoL="63FBZ" box="[151,290,1375,1400]" class="Insecta" family="Tephritidae" genus="Neotephritis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FAC72922C57F" box="[151,290,1375,1400]" italics="true" pageId="1" pageNumber="444">Neotephritis</emphasis>
</taxonomicName>
), so this is questionably a phylogenetically significant character at this level.
</paragraph>
<paragraph id="8BCAE70F4C5B3D2F81A9FA1A2D95C62E" blockId="1.[151,1437,343,1577]" pageId="1" pageNumber="444">
The following new or restored combinations are proposed for the species that have been classified in
<taxonomicName id="4C759C8C4C5B3D2F81F9FA3E294DC5B8" ID-CoL="3K23" box="[151,333,1446,1471]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B9013B1D4C5B3D2F81F9FA3E294DC5B8" box="[151,333,1446,1471]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
</taxonomicName>
(see Norrbom
<emphasis id="B9013B1D4C5B3D2F8364FA3F2A4AC5B8" box="[522,586,1446,1471]" italics="true" pageId="1" pageNumber="444">et al.</emphasis>
1999,
<bibRefCitation id="EFE49AFE4C5B3D2F83CAFA3E2B52C5B8" author="Norrbom" box="[676,850,1446,1471]" pageId="1" pageNumber="444" refString="Norrbom, A. L. (2004) Updates to Biosystematic Database of World Diptera for Tephritidae through 1999. Diptera Data Dissemination Disk (CD-ROM), 2." type="journal article" year="2004">Norrbom 2004</bibRefCitation>
) (original genus in parentheses):
<taxonomicName id="4C759C8C4C5B3D2F8585FA3F29C2C5E5" authority="(Blanchard 1852) (Acinia)" authorityName="Blanchard" authorityYear="1852" baseAuthorityName="Blanchard" baseAuthorityYear="1852" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B9013B1D4C5B3D2F8585FA3F2D9CC5B8" box="[1259,1436,1446,1471]" italics="true" pageId="1" pageNumber="444">Trupanea bella</emphasis>
(Blanchard 1852) (
<emphasis id="B9013B1D4C5B3D2F8000FA5229B7C5E6" box="[366,439,1482,1505]" italics="true" pageId="1" pageNumber="444">Acinia</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F80BEFA532BACC5E5" ID-CoL="87LM7" authority="(Seguy 1933) (Acanthiophilus)" authorityName="Seguy" authorityYear="1933" baseAuthorityName="Seguy" baseAuthorityYear="1933" box="[464,940,1481,1506]" class="Insecta" family="Tephritidae" genus="Acanthiophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="benoisti">
<emphasis id="B9013B1D4C5B3D2F80BEFA532A43C5E5" box="[464,579,1481,1506]" italics="true" pageId="1" pageNumber="444">T. benoisti</emphasis>
(Séguy 1933) (
<emphasis id="B9013B1D4C5B3D2F8399FA512BA1C5E5" box="[759,929,1481,1506]" italics="true" pageId="1" pageNumber="444">Acanthiophilus</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F82D4FA532D96C5E5" authority="(Schiner 1868) (Tephritis)" authorityName="Schiner" authorityYear="1868" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[954,1430,1481,1506]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="diespasmena">
<emphasis id="B9013B1D4C5B3D2F82D4FA532C67C5E5" box="[954,1127,1481,1506]" italics="true" pageId="1" pageNumber="444">T. diespasmena</emphasis>
(Schiner 1868) (
<emphasis id="B9013B1D4C5B3D2F8446FA512D8AC5E5" box="[1320,1418,1481,1506]" italics="true" pageId="1" pageNumber="444">Tephritis</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F81F9FA762AA1C601" ID-CoL="S3JN" authority="(Hendel 1914) (Celidosphenella)" authorityName="Hendel" authorityYear="1914" baseAuthorityName="Hendel" baseAuthorityYear="1914" box="[151,673,1517,1542]" class="Insecta" family="Tephritidae" genus="Celidosphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9013B1D4C5B3D2F81F9FA76291DC601" box="[151,285,1517,1542]" italics="true" pageId="1" pageNumber="444">T. maculata</emphasis>
(Hendel 1914) (
<emphasis id="B9013B1D4C5B3D2F808EFA752A96C601" box="[480,662,1517,1542]" italics="true" pageId="1" pageNumber="444">Celidosphenella</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F83DCFA762C63C601" ID-CoL="895Y4" authority="(Schiner 1868) (Sphenella)" authorityName="Schiner" authorityYear="1868" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[690,1123,1517,1542]" class="Insecta" family="Tephritidae" genus="Sphenella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="poecila">
<emphasis id="B9013B1D4C5B3D2F83DCFA762B23C601" box="[690,803,1517,1542]" italics="true" pageId="1" pageNumber="444">T. poecila</emphasis>
(Schiner 1868) (
<emphasis id="B9013B1D4C5B3D2F8284FA752C58C601" box="[1002,1112,1517,1542]" italics="true" pageId="1" pageNumber="444">Sphenella</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F851AFA762915C62E" authority="Malloch 1933" authorityName="Malloch" authorityYear="1933" class="Insecta" family="Tephritidae" genus="Trypanea" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="simulata">
<emphasis id="B9013B1D4C5B3D2F851AFA762CF3C601" box="[1140,1267,1517,1542]" italics="true" pageId="1" pageNumber="444">T. simulata</emphasis>
<bibRefCitation id="EFE49AFE4C5B3D2F8592FA752D9CC601" author="Malloch" box="[1276,1436,1517,1542]" pageId="1" pageNumber="444" refString="Malloch, J. R. (1933) Fascicle 4. - Acalyptrata [part]. In: Diptera of Patagonia and South Chile. Pt. 6. British Museum (Natural History), London, pp. 177 - 391, pls. 2 - 7." type="book chapter" year="1933">Malloch 1933</bibRefCitation>
(
<emphasis id="B9013B1D4C5B3D2F81CEF98A290AC62E" box="[160,266,1554,1577]" italics="true" pageId="1" pageNumber="444">Trypanea</emphasis>
)
</taxonomicName>
;
<taxonomicName id="4C759C8C4C5B3D2F804CF98A2AA0C62E" ID-CoL="89BDZ" authority="Stuardo 1946" authorityName="Stuardo" authorityYear="1946" box="[290,672,1552,1577]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="stonei">
<emphasis id="B9013B1D4C5B3D2F804CF98A297EC62E" box="[290,382,1554,1577]" italics="true" pageId="1" pageNumber="444">T. stonei</emphasis>
Stuardo 1946 (
<emphasis id="B9013B1D4C5B3D2F8342F98A2A97C62E" box="[556,663,1554,1577]" italics="true" pageId="1" pageNumber="444">Trupanea</emphasis>
)
</taxonomicName>
(=
<taxonomicName id="4C759C8C4C5B3D2F83A9F98A2BDBC62E" ID-CoL="7CWGY" authority="Malloch 1933" authorityName="Malloch" authorityYear="1933" box="[711,987,1552,1577]" class="Insecta" family="Tephritidae" genus="Trupanea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B9013B1D4C5B3D2F83A9F98A2B35C62E" box="[711,821,1552,1577]" italics="true" pageId="1" pageNumber="444">T. diversa</emphasis>
<bibRefCitation id="EFE49AFE4C5B3D2F8252F9882BDBC62E" author="Malloch" box="[828,987,1552,1577]" pageId="1" pageNumber="444" refString="Malloch, J. R. (1933) Fascicle 4. - Acalyptrata [part]. In: Diptera of Patagonia and South Chile. Pt. 6. British Museum (Natural History), London, pp. 177 - 391, pls. 2 - 7." type="book chapter" year="1933">Malloch 1933</bibRefCitation>
</taxonomicName>
); and
<taxonomicName id="4C759C8C4C5B3D2F8570F98A2D90C62E" authority="Hering 1942" authorityName="Hering" authorityYear="1942" box="[1054,1424,1552,1577]" class="Insecta" family="Tephritidae" genus="Trypanea" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="vidua">
<emphasis id="B9013B1D4C5B3D2F8570F98A2C78C62E" box="[1054,1144,1552,1577]" italics="true" pageId="1" pageNumber="444">T. vidua</emphasis>
Hering 1942 (
<emphasis id="B9013B1D4C5B3D2F8475F98A2D85C62E" box="[1307,1413,1554,1577]" italics="true" pageId="1" pageNumber="444">Trypanea</emphasis>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>