147 lines
19 KiB
XML
147 lines
19 KiB
XML
<document id="CEDEADB4612A28A7F2B929AEE28E31AF" ID-CLB-Dataset="24598" ID-DOI="10.5281/zenodo.3579152" ID-GBIF-Dataset="920561e2-c548-4c19-8be6-aeccfc9579a8" ID-ISSN="0374-1036" ID-Zenodo-Dep="3579152" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1576509166237" checkinUser="carolina" docAuthor="Bianchi, Filipe M., Barcellos, Aline & Grazia, Jocelia" docDate="2016" docId="03D7430D3C41013C1EBDFA94E2C207AD" docLanguage="en" docName="ActaEntMusNatPra.56.2.557-566.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Anhanga Distant 1887" docType="treatment" docVersion="10" lastPageNumber="560" masterDocId="FFEE3B753C40013F1F65FFBFE135053E" masterDocTitle="Rediscovering Anhanga: redescription and considerations on its taxonomic placement (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="566" masterPageNumber="557" pageNumber="558" updateTime="1698774496928" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2C521AFCCF560CADC0934624199735C5">Rediscovering Anhanga: redescription and considerations on its taxonomic placement (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
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<mods:name id="FC9342821C023E59368A962E2650E2F1" type="personal">
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<mods:namePart id="2913E85CCC81B6E090869189C862B01D">Bianchi, Filipe M.</mods:namePart>
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<mods:affiliation id="D5C7CB456903BEC67D349C9449A80089">Universidade Federal do Rio Grande do Sul, Departamento de Zoologia, Av. Bento Gonçalves 9500, 91501 - 970, Prédio 43435, Porto Alegre RS, Brazil</mods:affiliation>
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<mods:nameIdentifier id="8152631A3CF2F8203537EC119E5C7882" type="email">bianchi.fm@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="F9648461CA3CD079D30ABE3EC125F4F3">Barcellos, Aline</mods:namePart>
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<mods:affiliation id="015801EE46A61F92E41874D976AA0A72">Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Rua Dr. Salvador França, 1427, Porto Alegre, RS 90690 - 000, Brazil</mods:affiliation>
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<mods:nameIdentifier id="DDD9CB71C80BD47578F22BE52944779C" type="email">alinebar.fzb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FB68BBB30196FC026DF2980098830B53">Grazia, Jocelia</mods:namePart>
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<mods:affiliation id="571E8DFAB402ABE3E48F82A9002E7DC7">Universidade Federal do Rio Grande do Sul, Departamento de Zoologia, Av. Bento Gonçalves 9500, 91501 - 970, Prédio 43435, Porto Alegre RS, Brazil</mods:affiliation>
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<mods:nameIdentifier id="BED9F3D6172E0DE8BCAC1DB628EB8392" type="email">jocelia@ufrgs.com</mods:nameIdentifier>
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<mods:title id="E93EDD723F77A2C092B8E8AD162901FC">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="34AD232258897A2FD5433CCC943CACF6">2016</mods:date>
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<mods:number id="D8A6CEAFDE959E02DE4591EEC3C937C3">2016-11-15</mods:number>
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<treatment id="03D7430D3C41013C1EBDFA94E2C207AD" ID-GBIF-Taxon="160747976" LSID="urn:lsid:plazi:treatment:03D7430D3C41013C1EBDFA94E2C207AD" httpUri="http://treatment.plazi.org/id/03D7430D3C41013C1EBDFA94E2C207AD" lastPageId="3" lastPageNumber="560" pageId="1" pageNumber="558">
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<subSubSection id="C364A1903C41013E1EBDFA94E3EF007C" box="[472,730,1323,1346]" pageId="1" pageNumber="558" type="nomenclature">
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<paragraph id="8BC1F21B3C41013E1EBDFA94E3EF007C" blockId="1.[472,730,1323,1346]" box="[472,730,1323,1346]" pageId="1" pageNumber="558">
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<heading id="D08945773C41013E1EBDFA94E3EF007C" bold="true" box="[472,730,1323,1346]" centered="true" fontSize="10" level="3" pageId="1" pageNumber="558" reason="0">
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<taxonomicName id="4C7E89983C41013E1EBDFA94E3EF007C" authority="Distant, 1887" authorityName=" Distant" authorityYear="1887" box="[472,730,1323,1346]" class="Insecta" family="Cerambycidae" genus="Anhanga" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="B90A2E093C41013E1EBDFA94E3EF007C" bold="true" box="[472,730,1323,1346]" pageId="1" pageNumber="558">
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<emphasis id="B90A2E093C41013E1EBDFA94E30B007C" bold="true" box="[472,574,1323,1346]" italics="true" pageId="1" pageNumber="558">Anhanga</emphasis>
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<bibRefCitation id="EFEF8FEA3C41013E1D20FA94E3EF007C" author="DISTANT W. L." box="[581,730,1323,1346]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="1" pageNumber="558" pagination="lvi-lxvi" part="31" refId="ref3544" refString="DISTANT W. L. 1887: Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I. Annales de la Societe Entomologique de Belgique 31 (3): lvi-lxvi." title="Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I" type="journal article" year="1887">Distant, 1887</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C364A1903C41013E1F14FAEEE0C700A7" pageId="1" pageNumber="558" type="reference_group">
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<paragraph id="8BC1F21B3C41013E1F14FAEEE0C700A7" blockId="1.[113,1053,1361,1433]" pageId="1" pageNumber="558">
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<treatmentCitationGroup id="AB6ED5353C41013E1F14FAEEE2730040" pageId="1" pageNumber="558">
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<treatmentCitation id="0ADFD40A3C41013E1F14FAEEE066005A" box="[113,339,1361,1380]" pageId="1" pageNumber="558">
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<taxonomicName id="4C7E89983C41013E1F14FAEEE00F005A" authority="Distant, 1887" authorityName="Distant" authorityYear="1887" box="[113,314,1361,1380]" class="Insecta" family="Cerambycidae" genus="Anhanga" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="B90A2E093C41013E1F14FAEEE18B005A" box="[113,190,1361,1380]" italics="true" pageId="1" pageNumber="558">Anhanga</emphasis>
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<bibRefCitation id="EFEF8FEA3C41013E1FA1FAEEE00F005A" author="DISTANT W. L." box="[196,314,1361,1380]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="1" pageNumber="558" pagination="lvi-lxvi" part="31" refId="ref3544" refString="DISTANT W. L. 1887: Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I. Annales de la Societe Entomologique de Belgique 31 (3): lvi-lxvi." title="Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I" type="journal article" year="1887">Distant, 1887</bibRefCitation>
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</taxonomicName>
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: lx
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</treatmentCitation>
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(original description).
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<typeStatus id="54C54CB93C41013E1D72FAEEE374005A" box="[535,577,1361,1380]" pageId="1" pageNumber="558">Type</typeStatus>
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species:
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<taxonomicName id="4C7E89983C41013E1DEAFAEEE296005A" authority="Distant, 1887" authorityName="Distant" authorityYear="1887" box="[655,931,1361,1380]" class="Insecta" family="Cerambycidae" genus="Anhanga" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="B90A2E093C41013E1DEAFAEEE21C005A" box="[655,809,1361,1380]" italics="true" pageId="1" pageNumber="558">Anhanga modesta</emphasis>
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<bibRefCitation id="EFEF8FEA3C41013E1C4BFAEEE296005A" author="DISTANT W. L." box="[814,931,1361,1380]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="1" pageNumber="558" pagination="lvi-lxvi" part="31" refId="ref3544" refString="DISTANT W. L. 1887: Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I. Annales de la Societe Entomologique de Belgique 31 (3): lvi-lxvi." title="Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I" type="journal article" year="1887">Distant, 1887</bibRefCitation>
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</taxonomicName>
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, by monotypy
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<taxonomicName id="4C7E89983C41013E1F14FAD4E1860040" authority="Distant, 1887" authorityName="Distant" authorityYear="1887" box="[113,179,1387,1406]" pageId="1" pageNumber="558">
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<emphasis id="B90A2E093C41013E1F14FAD4E1860040" box="[113,179,1387,1406]" italics="true" pageId="1" pageNumber="558">Ahanga</emphasis>
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</taxonomicName>
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(incorrect subsequent spelling):
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<treatmentCitation id="0ADFD40A3C41013E1EAEFAD4E3EB0040" box="[459,734,1387,1406]" pageId="1" pageNumber="558">
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<bibRefCitation id="EFEF8FEA3C41013E1EAEFAD4E3F40040" author="LETHIERRY L. & SEVERIN G." bookContentInfo="286 pp." box="[459,705,1387,1406]" pageId="1" pageNumber="558" refId="ref3762" refString="LETHIERRY L. & SEVERIN G. 1893. Catalogue general des Hemipteres. Vol. I. Pentatomidae. Bruxelles, x + 286 pp." title="Catalogue general des Hemipteres. Vol. I. Pentatomidae" type="book" year="1893">LETHIERRY & SEVERIN (1893)</bibRefCitation>
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: 89
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</treatmentCitation>
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||
(catalogue)
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||
</treatmentCitationGroup>
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||
.
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<treatmentCitationGroup id="AB6ED5353C41013E1F14FA39E0DB00A7" box="[113,494,1414,1433]" pageId="1" pageNumber="558">
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<taxonomicName id="4C7E89983C41013E1F14FA39E18B00A7" authority="Distant, 1887" authorityName="Distant" authorityYear="1887" box="[113,190,1414,1433]" class="Insecta" family="Cerambycidae" genus="Anhanga" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="B90A2E093C41013E1F14FA39E18B00A7" box="[113,190,1414,1433]" italics="true" pageId="1" pageNumber="558">Anhanga</emphasis>
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</taxonomicName>
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:
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<treatmentCitation id="0ADFD40A3C41013E1FAFFA39E0B300A7" box="[202,390,1414,1433]" pageId="1" pageNumber="558">
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<bibRefCitation id="EFEF8FEA3C41013E1FAFFA39E06B00A7" author="KIRKALDY G. W." bookContentInfo="392 pp." box="[202,350,1414,1433]" pageId="1" pageNumber="558" refId="ref3688" refString="KIRKALDY G. W. 1909: Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I. Cimicidae. Berlin, xl + 392 pp." title="Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I. Cimicidae" type="book" year="1909">KIRKALDY (1909)</bibRefCitation>
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: 221
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</treatmentCitation>
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(catalogue)
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</treatmentCitationGroup>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C364A1903C41013C1F14FA0EE32E046D" lastPageId="3" lastPageNumber="560" pageId="1" pageNumber="558" type="diagnosis">
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<paragraph id="8BC1F21B3C41013C1F14FA0EE32E046D" blockId="1.[113,1089,1457,1576]" lastBlockId="3.[113,1089,220,659]" lastPageId="3" lastPageNumber="560" pageId="1" pageNumber="558">
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<emphasis id="B90A2E093C41013E1F14FA0EE1D300F6" bold="true" box="[113,230,1457,1480]" pageId="1" pageNumber="558">Diagnosis.</emphasis>
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In addition to
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<bibRefCitation id="EFEF8FEA3C41013E1EEFFA0EE37400F6" author="DISTANT W. L." box="[394,577,1457,1480]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="1" pageNumber="558" pagination="lvi-lxvi" part="31" refId="ref3544" refString="DISTANT W. L. 1887: Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I. Annales de la Societe Entomologique de Belgique 31 (3): lvi-lxvi." title="Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I" type="journal article" year="1887">DISTANT’s (1887)</bibRefCitation>
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original description, the following features are highlighted. Antennae 5-segmented; first antennal segment reaching but never surpassing anterior margin of head. Head somewhat long, narrowed and rounded in front; mandibular plates reflexed, convergent at apex and much longer than clypeus. Basal segment of rostrum arising anteriorly to imaginary line crossing head at anterior margin of eyes (
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<figureCitation id="1345EE9E3C42013D1CB1FA2BE5120095" box="[980,1063,1428,1451]" captionStart="Fig" captionStartId="2.[151,182,1210,1229]" captionTargetBox="[177,1114,225,1193]" captionTargetId="figure@2.[160,1117,219,1194]" captionTargetPageId="2" captionText="Fig. 1. Anhanga modesta Distant, 1887. A – dorsal view; B – magnification of the head in ventral view, dashed line highlighting the basal segment of labium arising anteriorly to the imaginary line traversing head at anterior margin of eyes; C – ventral view; D – magnification of external scent efferent system of metathoracic scent glands; E – magnification of abdomen, highlighting the placement of the pair of trichobothria (arrows) along the imaginary line of spiracles on urosternites VI and VII. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3579156" httpUri="https://zenodo.org/record/3579156/files/figure.png" pageId="2" pageNumber="559">Fig. 1B</figureCitation>
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); first rostral segment surpassing posterior limits of bucculae; apex of rostrum surpassing mesocoxae. Anterolateral margins of pronotum slightly reflexed, crenulate along anterior half; lateral angles rounded and not projecting. Scutellum not exceeding posterior margin of connexival segment V. Prosternum slightly projected. Peritreme spout-shaped, very short, about three times diameter of ostiole; evaporatorium covering 2/3 of metapleuron. Hemelytranot covering connexivum. Tarsi 3-segmented. Abdominal venter not tuberculated, without median sulcus or spine. Spiracles placed on callus. Pair of trichobothria present onurosternites III–VII along imaginary line connecting the spiracles.
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</paragraph>
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||
</subSubSection>
|
||
<caption id="DF01A2933C42013D1FF2FB05E34E0006" ID-DOI="http://doi.org/10.5281/zenodo.3579156" ID-Zenodo-Dep="3579156" httpUri="https://zenodo.org/record/3579156/files/figure.png" pageId="2" pageNumber="559" startId="2.[151,182,1210,1229]" targetBox="[177,1114,225,1193]" targetPageId="2">
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<paragraph id="8BC1F21B3C42013D1FF2FB05E34E0006" blockId="2.[151,1127,1210,1336]" pageId="2" pageNumber="559">
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Fig. 1.
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<taxonomicName id="4C7E89983C42013D1FB6FB05E0D301F3" authority="modesta Distant, 1887" authorityName="modesta Distant" authorityYear="1887" box="[211,486,1210,1229]" class="Insecta" family="Cerambycidae" genus="Anhanga" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="B90A2E093C42013D1FB6FB05E05901F3" box="[211,364,1210,1229]" italics="true" pageId="2" pageNumber="559">Anhanga modesta</emphasis>
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||
<bibRefCitation id="EFEF8FEA3C42013D1E14FB05E0D301F3" author="DISTANT W. L." box="[369,486,1210,1229]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="2" pageNumber="559" pagination="lvi-lxvi" part="31" refId="ref3544" refString="DISTANT W. L. 1887: Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I. Annales de la Societe Entomologique de Belgique 31 (3): lvi-lxvi." title="Enumeration of the Van Volxem collection of Rhynchota contained in the Brussels Museum. Part I" type="journal article" year="1887">Distant, 1887</bibRefCitation>
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||
</taxonomicName>
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||
. A – dorsal view; B – magnification of the head in ventral view, dashed line highlighting the basal segment of labium arising anteriorly to the imaginary line traversing head at anterior margin of eyes; C – ventral view; D – magnification of external scent efferent system of metathoracic scent glands; E – magnification of abdomen, highlighting the placement of the pair of trichobothria (arrows) along the imaginary line of spiracles on urosternites VI and VII. Scale bar: 5 mm.
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</paragraph>
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</caption>
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||
<subSubSection id="C364A1903C43013C1FF5FEE3E2C207AD" pageId="3" pageNumber="560" type="description">
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||
<paragraph id="8BC1F21B3C43013C1FF5FEE3E2B304CD" blockId="3.[113,1089,220,659]" pageId="3" pageNumber="560">
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||
<emphasis id="B90A2E093C43013C1FF5FEE3E006044D" bold="true" box="[144,307,348,371]" italics="true" pageId="3" pageNumber="560">Male genitalia.</emphasis>
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Pygophore globose, posterolateral anglesprojected, spatulate, convergent. Dorsal rim concave and slightly projected over segment X. Ventral rim with open V-shaped excavation, bearing 1+1 acute projections close to posterolateral angles. Superior layer of ventral rim well developed, with 1+1 blunt and hairy projections directed dorsad. Parameres paddle-like. Segment X lozenge-shaped; posterior half grooved and hairy.
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</paragraph>
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<paragraph id="8BC1F21B3C43013C1FF5FE43E2C207AD" blockId="3.[113,1089,220,659]" pageId="3" pageNumber="560">
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<emphasis id="B90A2E093C43013C1FF5FE43E062072D" bold="true" box="[144,343,508,531]" italics="true" pageId="3" pageNumber="560">Female genitalia.</emphasis>
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Gonocoxites VIII quadrangular; anterolateral margin rounded and posterior margin emarginated. Gonocoxites IX trapezoidal. Laterotergites VIII presenting spiracles. Laterotergite VIII and IX similar in length. Laterotergites IX rounded at apex, not surpassing transverse band linking laterotergites VIII. Ring sclerites present. Vesicular area of receptaculum seminis well developed. Capsula seminalis with finger-like process.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |