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<document id="56987F7B978205A8124C562CE5C9D38C" ID-CLB-Dataset="272979" ID-DOI="10.11646/zootaxa.5353.5.5" ID-GBIF-Dataset="c44c50a8-887f-449e-b454-bc4086743b34" ID-ISSN="1175-5326" ID-Zenodo-Dep="10010393" ID-ZooBank="D35586A6-9BB0-4FDF-AE51-BC1475B346A2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1697229707486" checkinUser="plazi" docAuthor="Watanabe, Kohei, Nakajima, Jun &amp; Hayashi, Masakazu" docDate="2023" docId="03D5120F332CFFB424FA57CEFDF5F956" docLanguage="en" docName="zootaxa.5353.5.5.pdf" docOrigin="Zootaxa 5353 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.5.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nagisavelia Watanabe, Nakajima et Hayashi 2023, gen. nov." docType="treatment" docVersion="3" lastPageNumber="469" masterDocId="FFEC6A77332DFFB5246D5614FFEFFFC4" masterDocTitle="Nagisavelia hikarui, a new genus and species of Mesoveliinae (Hemiptera: Heteroptera: Mesoveliidae) inhabiting shingle beaches in Japan" masterLastPageNumber="478" masterPageNumber="468" pageNumber="469" updateTime="1697480010607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AA768243C91BCE27EFA5232A950CFDB2">Nagisavelia hikarui, a new genus and species of Mesoveliinae (Hemiptera: Heteroptera: Mesoveliidae) inhabiting shingle beaches in Japan</mods:title>
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<mods:namePart id="0106EDE66D9127A916E9D737D49D0214">Watanabe, Kohei</mods:namePart>
<mods:affiliation id="34737F4AA31885E655B9FAB02CF5B643">Ishikawa Insect Museum, Hakusan, Ishikawa, 920 - 2113 Japan.</mods:affiliation>
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<mods:namePart id="3407254B21DD914F273DE91130265514">Nakajima, Jun</mods:namePart>
<mods:affiliation id="9D0C09C3C3BDC62E4D8839719C0F1C59">Fukuoka Institute of Health and Environmental Sciences, Dazaifu, Fukuoka, 811 - 0135 Japan.</mods:affiliation>
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<mods:namePart id="C4720A15B08AD1869EA7CAE5070C80BE">Hayashi, Masakazu</mods:namePart>
<mods:affiliation id="B7A3B654AFB780881C306CB84A2BA2DE">Hoshizaki Green Foundation, Izumo, Shimane, 691 - 0076 Japan.</mods:affiliation>
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<mods:title id="34885DB687BCC9968DB0A72C0C26E3A0">Zootaxa</mods:title>
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<mods:date id="EBCA70497E5659D9EB8A61B730699AC6">2023</mods:date>
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<mods:number id="63E1B9848A55A23ECA0EAD3C24A245BE">2023-10-12</mods:number>
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<treatment id="03D5120F332CFFB424FA57CEFDF5F956" ID-DOI="http://doi.org/10.5281/zenodo.10010395" ID-Zenodo-Dep="10010395" LSID="urn:lsid:plazi:treatment:03D5120F332CFFB424FA57CEFDF5F956" httpUri="http://treatment.plazi.org/id/03D5120F332CFFB424FA57CEFDF5F956" lastPageNumber="469" pageId="1" pageNumber="469">
<subSubSection id="C366F092332CFFB424FA57CEFD5CFDDD" pageId="1" pageNumber="469" type="nomenclature">
<paragraph id="8BC3A319332CFFB424FA57CEFCD0FE31" blockId="1.[151,831,474,573]" box="[151,831,474,501]" pageId="1" pageNumber="469">
<heading id="D08B1475332CFFB424FA57CEFCD0FE31" bold="true" box="[151,831,474,501]" fontSize="11" level="1" pageId="1" pageNumber="469" reason="1">
<emphasis id="B9087F0B332CFFB424FA57CEFCD0FE31" bold="true" box="[151,831,474,501]" pageId="1" pageNumber="469">
<taxonomicName id="4C7CD89A332CFFB424FA57CEFD2CFE30" authority="Watanabe, Nakajima et Hayashi" authorityName="Watanabe, Nakajima et Hayashi" authorityYear="2023" box="[151,707,474,501]" class="Insecta" family="Mesoveliidae" genus="Nagisavelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9087F0B332CFFB424FA57CEFECAFE31" bold="true" box="[151,293,474,501]" italics="true" pageId="1" pageNumber="469">Nagisavelia</emphasis>
Watanabe, Nakajima et Hayashi
</taxonomicName>
,
<taxonomicNameLabel id="A23BC270332CFFB426BC57CFFCD0FE31" box="[721,831,475,501]" pageId="1" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC3A319332CFFB424FA57EBFD5CFDDD" blockId="1.[151,831,474,573]" box="[151,691,511,537]" pageId="1" pageNumber="469">[Japanese name: Nagisa-Mizukamemushi-zoku]</paragraph>
</subSubSection>
<subSubSection id="C366F092332CFFB424FA5437FE8AFDF9" box="[151,357,547,573]" pageId="1" pageNumber="469" type="description">
<paragraph id="8BC3A319332CFFB424FA5437FE8AFDF9" blockId="1.[151,831,474,573]" box="[151,357,547,573]" pageId="1" pageNumber="469">
(
<figureCitation id="1347BF9C332CFFB424F25437FEE4FDF9" box="[159,267,547,573]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–10" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1544,1568]" captionTargetBox-0="[182,1406,189,1897]" captionTargetBox-1="[151,1436,453,1521]" captionTargetId-0="figure-17@2.[172,1415,181,1919]" captionTargetId-1="figure-164@3.[151,1436,453,1521]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Nagisavelia hikarui sp. nov. 1, 3, 5, male; 2, 4, 6, female. 1, Holotype; 2–6, paratypes. 1–3, Dorsal aspect; 4, ventral aspect. 5–6, Live specimen. Scale refers to 1–4." captionText-1="FIGURES 7–10. Nagisavelia hikarui sp. nov. and its type locality. 7, Adult male; 8, adult female; 9, type locality; 10, habitat." figureDoi-0="http://doi.org/10.5281/zenodo.10010397" figureDoi-1="http://doi.org/10.5281/zenodo.10010399" httpUri-0="https://zenodo.org/record/10010397/files/figure.png" httpUri-1="https://zenodo.org/record/10010399/files/figure.png" pageId="1" pageNumber="469">Figs. 1–8</figureCitation>
,
<figureCitation id="1347BF9C332CFFB4257A5437FEB2FDF9" box="[279,349,547,573]" captionStart-0="FIGURES 11–14" captionStart-1="FIGURES 15–18" captionStart-2="FIGURES 19–24" captionStartId-0="4.[151,264,1472,1496]" captionStartId-1="5.[151,264,1938,1962]" captionStartId-2="6.[151,264,1650,1674]" captionTargetBox-0="[182,1436,181,1427]" captionTargetBox-1="[191,1394,525,1915]" captionTargetBox-2="[151,1430,181,1626]" captionTargetId-0="figure-17@4.[151,1436,181,1448]" captionTargetId-1="figure-186@5.[191,1395,525,1915]" captionTargetId-2="figure-17@6.[151,1436,181,1627]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 11–14. Abdominal segment VIII, pygopore, and proctiger of Nagisavelia hikarui sp. nov. male, holotype. 11, Abdominal segment VIII, ventral view; 12, proctiger, dorsal view; 13, pygophore (segment IX) and proctiger, lateral view; 14, same, dorsal view. Scale bars = 0.05 mm." captionText-1="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." captionText-2="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi-0="http://doi.org/10.5281/zenodo.10010403" figureDoi-1="http://doi.org/10.5281/zenodo.10010407" figureDoi-2="http://doi.org/10.5281/zenodo.10010410" httpUri-0="https://zenodo.org/record/10010403/files/figure.png" httpUri-1="https://zenodo.org/record/10010407/files/figure.png" httpUri-2="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">11–24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C366F092332CFFB424FA5470FD98FDBA" box="[151,631,612,638]" pageId="1" pageNumber="469" type="type_taxon">
<paragraph id="8BC3A319332CFFB424FA5470FD98FDBA" blockId="1.[151,1437,612,1683]" box="[151,631,612,638]" pageId="1" pageNumber="469">
<emphasis id="B9087F0B332CFFB424FA5470FEDDFDBA" bold="true" box="[151,306,612,638]" pageId="1" pageNumber="469">
<typeStatus id="54C71DBB332CFFB424FA5470FF3DFDBA" box="[151,210,612,638]" pageId="1" pageNumber="469">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C7CD89A332CFFB425545470FDF8FDBA" authority="Watanabe &amp; Nakajima &amp; Hayashi, 2023" authorityName="Watanabe &amp; Nakajima &amp; Hayashi" authorityYear="2023" box="[313,535,612,638]" class="Insecta" family="Mesoveliidae" genus="Nagisavelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="species" species="hikarui" status="sp. nov.">
<emphasis id="B9087F0B332CFFB425545470FDF8FDBA" box="[313,535,612,638]" italics="true" pageId="1" pageNumber="469">Nagisavelia hikarui</emphasis>
</taxonomicName>
<emphasis id="B9087F0B332CFFB426735470FD98FDBA" bold="true" box="[542,631,612,638]" pageId="1" pageNumber="469">
<taxonomicNameLabel id="A23BC270332CFFB426735470FD98FDBA" box="[542,631,612,638]" pageId="1" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C366F092332CFFB424AA549CFB73FC06" pageId="1" pageNumber="469" type="description">
<paragraph id="8BC3A319332CFFB424AA549CFB73FC06" blockId="1.[151,1437,612,1683]" pageId="1" pageNumber="469">
<emphasis id="B9087F0B332CFFB424AA549CFEB6FD66" bold="true" box="[199,345,648,674]" pageId="1" pageNumber="469">Description.</emphasis>
Apterous form, small. Head very long; ocelli absent; eyes vestigial, with small ommatidia, located near base of head, with dense setae (
<figureCitation id="1347BF9C332CFFB4265654B8FD7FFD02" box="[571,656,684,710]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 21</figureCitation>
); antennae long and slender, filamentous, shorter than body, without spines; ventral head very long, weakly carinate (
<figureCitation id="1347BF9C332CFFB426A754C4FCCEFD2F" box="[714,801,720,747]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 15</figureCitation>
); labium long, reaching posterior margin of abdominal sternum II; pronotum broad, posterior margin slightly arched; mesonotum broad, posterior margin slightly arched; metanotum broad, posterior margin straight; without wings; middle thoracic venter with single metasternal scent gland opening; legs slender and long, hind legs longest, spines absent, but hind tibia with weak spines; tarsi with three segments, first segment shortest, third segment longest (
<figureCitation id="1347BF9C332CFFB427345574FC40FCBE" box="[857,943,864,890]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 22</figureCitation>
); lateral abdomen with longitudinal sulcus; male genital segment large, parameres symmetrical; female genital segment large, ovipositor ensiform, both sides moderated middle to apex (
<figureCitation id="1347BF9C332CFFB425A755BCFDF2FC06" box="[458,541,936,962]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 17</figureCitation>
); gonoplacs large (
<figureCitation id="1347BF9C332CFFB4269F55BCFCAAFC06" box="[754,837,936,962]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 16</figureCitation>
); gynatrial sac small (
<figureCitation id="1347BF9C332CFFB4205155BCFB60FC06" box="[1084,1167,936,962]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366F092332CFFB424AA55D8FE9BF98E" pageId="1" pageNumber="469" type="discussion">
<paragraph id="8BC3A319332CFFB424AA55D8FE9BF98E" blockId="1.[151,1437,612,1683]" pageId="1" pageNumber="469">
<emphasis id="B9087F0B332CFFB424AA55D8FED4FC22" bold="true" box="[199,315,972,998]" pageId="1" pageNumber="469">Remarks.</emphasis>
<taxonomicName id="4C7CD89A332CFFB4252B55D8FE23FC22" authority="Watanabe &amp; Nakajima &amp; Hayashi, 2023" authorityName="Watanabe &amp; Nakajima &amp; Hayashi" authorityYear="2023" box="[326,460,972,998]" class="Insecta" family="Mesoveliidae" genus="Nagisavelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9087F0B332CFFB4252B55D8FE23FC22" box="[326,460,972,998]" italics="true" pageId="1" pageNumber="469">Nagisavelia</emphasis>
</taxonomicName>
<emphasis id="B9087F0B332CFFB425B555D8FDAAFC22" bold="true" box="[472,581,972,998]" pageId="1" pageNumber="469">
<taxonomicNameLabel id="A23BC270332CFFB425B555D8FDAAFC22" box="[472,581,972,998]" pageId="1" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from nine of the current mesoveliine genera by its relatively vestigial eyes (
<tableCitation id="C6FE96A2332CFFB425D755E4FDF9FBCE" box="[442,534,1008,1034]" captionStart="TABLE 1" captionStartId="8.[151,239,478,504]" captionTargetPageId="8" captionText="TABLE 1. Character matrix of eight genera of Mesoveliinae (based on Andersen &amp; Polhemus 1980, 2003)." httpUri="http://table.plazi.org/id/DF03F3913325FFBD24FA57CAFAFDFE3C" pageId="1" pageNumber="469" tableUuid="DF03F3913325FFBD24FA57CAFAFDFE3C">Table 1</tableCitation>
;
<bibRefCitation id="EFEDDEE8332CFFB4264855E4FC96FBCE" author="Andersen, N. M. &amp; Polhemus, J. T." box="[549,889,1008,1034]" pageId="1" pageNumber="469" pagination="369 - 392" refId="ref5369" refString="Andersen, N. M. &amp; Polhemus, J. T. (1980) Four new genera of Mesoveliidae (Hemiptera, Gerromorpha) and the phylogeny and classification of the family. Entomologica Scandinavica, 11, 369 - 392. https: // doi. org / 10.1163 / 187631280794710060" type="journal article" year="1980">Andersen &amp; Polhemus 1980</bibRefCitation>
,
<bibRefCitation id="EFEDDEE8332CFFB427EB55E4FC2AFBCE" author="Andersen, N. M. &amp; Polhemus, D. A." box="[902,965,1008,1034]" pageId="1" pageNumber="469" pagination="12 - 21" refId="ref5423" refString="Andersen, N. M. &amp; Polhemus, D. A. (2003) A new genus of terrestrial Mesoveliidae from the Seychelles (Hemiptera: Gerromorpha). Journal of the New York Entomological Society, 111, 12 - 21. https: // doi. org / 10.1664 / 0028 - 7199 (2003) 111 [0012: ANGOTM] 2.0. CO; 2" type="journal article" year="2003">2003</bibRefCitation>
;
<bibRefCitation id="EFEDDEE8332CFFB427BF55E4FAF3FBCE" author="Malipatil, M. B. &amp; Monteith, B." box="[978,1308,1008,1034]" pageId="1" pageNumber="469" pagination="943 - 955" refId="ref5952" refString="Malipatil, M. B. &amp; Monteith, B. (1983) One new genus and four new species of terrestrial Mesoveliidae (Hemiptera: Gerromorpha) from Australia and New Caledonia. Australian Journal of Zoology, 31 (6), 943 - 955. https: // doi. org / 10.1071 / ZO 9830943" type="journal article" year="1983">Malipatil &amp; Monteith 1983</bibRefCitation>
;
<bibRefCitation id="EFEDDEE8332CFFB4214455E4FE99FBEA" author="Polhemus, J. T. &amp; Polhemus, D. A." pageId="1" pageNumber="469" pagination="73 - 82" refId="ref6129" refString="Polhemus, J. T. &amp; Polhemus, D. A. (1989) A new mesoveliid genus and two new species of Hebrus (Heteroptera: Mesoveliidae, Hebridae) from intertidal habitats in Southeast Asian mangrove swamps. Raffles Bulletin of Zoology, 37 (1 &amp; 2), 73 - 82." type="journal article" year="1989">Polhemus &amp; Polhemus 1989</bibRefCitation>
).
<taxonomicName id="4C7CD89A332CFFB425E05200FDC6FBEA" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[397,553,1044,1070]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB425E05200FDC6FBEA" box="[397,553,1044,1070]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7CD89A332CFFB426A95200FCA5FBEA" authority="Watanabe &amp; Nakajima &amp; Hayashi, 2023" authorityName="Watanabe &amp; Nakajima &amp; Hayashi" authorityYear="2023" box="[708,842,1044,1070]" class="Insecta" family="Mesoveliidae" genus="Nagisavelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9087F0B332CFFB426A95200FCA5FBEA" box="[708,842,1044,1070]" italics="true" pageId="1" pageNumber="469">Nagisavelia</emphasis>
</taxonomicName>
<emphasis id="B9087F0B332CFFB427395200FC50FBEA" bold="true" box="[852,959,1044,1070]" pageId="1" pageNumber="469">
<taxonomicNameLabel id="A23BC270332CFFB427395200FC50FBEA" box="[852,959,1044,1070]" pageId="1" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in regard to its very long head, vestigial eyes, filiform antennae, and the long labium which reaches to the middle of the abdomen, but can be distinguished by the following combination of characteristics (
<tableCitation id="C6FE96A2332CFFB426A85248FCF3FBB2" box="[709,796,1116,1142]" captionStart="TABLE 1" captionStartId="8.[151,239,478,504]" captionTargetPageId="8" captionText="TABLE 1. Character matrix of eight genera of Mesoveliinae (based on Andersen &amp; Polhemus 1980, 2003)." httpUri="http://table.plazi.org/id/DF03F3913325FFBD24FA57CAFAFDFE3C" pageId="1" pageNumber="469" tableUuid="DF03F3913325FFBD24FA57CAFAFDFE3C">Table 1</tableCitation>
): body royal-orange (body yellowish in
<taxonomicName id="4C7CD89A332CFFB4209D5248FA63FBB2" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[1264,1420,1116,1142]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB4209D5248FA63FBB2" box="[1264,1420,1116,1142]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); antennae shorter than body (antennae longer than body in
<taxonomicName id="4C7CD89A332CFFB427505294FC36FB5E" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[829,985,1152,1178]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427505294FC36FB5E" box="[829,985,1152,1178]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); antennae without spines (with spines in
<taxonomicName id="4C7CD89A332CFFB424DA52B0FEBCFB7A" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[183,339,1188,1214]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB424DA52B0FEBCFB7A" box="[183,339,1188,1214]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); eyes with dense hairs (
<figureCitation id="1347BF9C332CFFB4261552B0FD23FB7A" box="[632,716,1188,1214]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 21</figureCitation>
) (eyes with two short setae in
<taxonomicName id="4C7CD89A332CFFB4205552B0FB3BFB7A" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[1080,1236,1188,1214]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB4205552B0FB3BFB7A" box="[1080,1236,1188,1214]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
: see
<bibRefCitation id="EFEDDEE8332CFFB4217C52B0FEA7FB26" author="Andersen, N. M. &amp; Polhemus, J. T." pageId="1" pageNumber="469" pagination="369 - 392" refId="ref5369" refString="Andersen, N. M. &amp; Polhemus, J. T. (1980) Four new genera of Mesoveliidae (Hemiptera, Gerromorpha) and the phylogeny and classification of the family. Entomologica Scandinavica, 11, 369 - 392. https: // doi. org / 10.1163 / 187631280794710060" type="journal article" year="1980">Andersen &amp; Polhemus 1980</bibRefCitation>
: fig. 47); ventral head weakly carinate (
<figureCitation id="1347BF9C332CFFB4269052DCFCA1FB27" box="[765,846,1224,1251]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 15</figureCitation>
) (simple in
<taxonomicName id="4C7CD89A332CFFB427BF52DCFB81FB26" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[978,1134,1224,1250]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427BF52DCFB81FB26" box="[978,1134,1224,1250]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); legs short (legs very long in
<taxonomicName id="4C7CD89A332CFFB424D852F8FEBEFAC2" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[181,337,1260,1286]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB424D852F8FEBEFAC2" box="[181,337,1260,1286]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
: see
<bibRefCitation id="EFEDDEE8332CFFB425E652F8FD4FFAC3" author="Gagne, W. C. &amp; Howarth, F. G." box="[395,672,1260,1287]" pageId="1" pageNumber="469" pagination="399 - 413" refId="ref5700" refString="Gagne, W. C. &amp; Howarth, F. G. (1975) The cavernicolous fauna of Hawaiian lava tubes 6. Mesoveliidae or water treaders (Heteroptera). Pacific Insects, 16 (4), 399 - 413." type="journal article" year="1975">Gagné &amp; Howarth 1975</bibRefCitation>
: fig. 1); thoracic segment length I&gt; II&gt; III (
<figureCitation id="1347BF9C332CFFB420F152F8FB01FAC2" box="[1180,1262,1260,1286]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 19</figureCitation>
) (I&gt; III&gt; II in
<taxonomicName id="4C7CD89A332CFFB424FA5304FEDCFAEE" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[151,307,1296,1322]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB424FA5304FEDCFAEE" box="[151,307,1296,1322]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); spines on legs absent but hind tibia weak (weak in
<taxonomicName id="4C7CD89A332CFFB427E55304FBCBFAEE" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[904,1060,1296,1322]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427E55304FBCBFAEE" box="[904,1060,1296,1322]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
); metatarsal segment length III&gt; II&gt; I (
<figureCitation id="1347BF9C332CFFB4248C5320FEDDFA8A" box="[225,306,1332,1358]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 22</figureCitation>
) (II&gt; III&gt; I in
<taxonomicName id="4C7CD89A332CFFB4258E5320FD90FA8A" authorityName="Andersen &amp; Polhemus" authorityYear="1980" box="[483,639,1332,1358]" class="Insecta" family="Mesoveliidae" genus="Cavaticovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB4258E5320FD90FA8A" box="[483,639,1332,1358]" italics="true" pageId="1" pageNumber="469">Cavaticovelia</emphasis>
</taxonomicName>
). In
<collectingCountry id="F36BE389332CFFB426D85321FCCFFA8A" box="[693,800,1332,1359]" name="China" pageId="1" pageNumber="469">East Asia</collectingCountry>
,
<taxonomicName id="4C7CD89A332CFFB427415320FC76FA8A" authorityName="Esaki" authorityYear="1929" box="[812,921,1332,1358]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427415320FC76FA8A" box="[812,921,1332,1358]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7CD89A332CFFB420415320FB5DFA8A" authority="Watanabe &amp; Nakajima &amp; Hayashi, 2023" authorityName="Watanabe &amp; Nakajima &amp; Hayashi" authorityYear="2023" box="[1068,1202,1332,1358]" class="Insecta" family="Mesoveliidae" genus="Nagisavelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9087F0B332CFFB420415320FB5DFA8A" box="[1068,1202,1332,1358]" italics="true" pageId="1" pageNumber="469">Nagisavelia</emphasis>
</taxonomicName>
<emphasis id="B9087F0B332CFFB420D45320FACDFA8A" bold="true" box="[1209,1314,1332,1358]" pageId="1" pageNumber="469">
<taxonomicNameLabel id="A23BC270332CFFB420D45320FACDFA8A" box="[1209,1314,1332,1358]" pageId="1" pageNumber="469" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished by the following combination of characteristics (
<tableCitation id="C6FE96A2332CFFB4273C534CFC49FAB6" box="[849,934,1368,1394]" captionStart="TABLE 1" captionStartId="8.[151,239,478,504]" captionTargetPageId="8" captionText="TABLE 1. Character matrix of eight genera of Mesoveliinae (based on Andersen &amp; Polhemus 1980, 2003)." httpUri="http://table.plazi.org/id/DF03F3913325FFBD24FA57CAFAFDFE3C" pageId="1" pageNumber="469" tableUuid="DF03F3913325FFBD24FA57CAFAFDFE3C">Table 1</tableCitation>
): very long head, more than 1.8 times longer than width (less than 1.5 times as long as width in
<taxonomicName id="4C7CD89A332CFFB426815368FCB6FA52" authorityName="Esaki" authorityYear="1929" box="[748,857,1404,1430]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB426815368FCB6FA52" box="[748,857,1404,1430]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); eyes vestigial (
<figureCitation id="1347BF9C332CFFB4204A5368FB6FFA52" box="[1063,1152,1404,1430]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 21</figureCitation>
) (reduced in
<taxonomicName id="4C7CD89A332CFFB421455368FA7AFA52" authorityName="Esaki" authorityYear="1929" box="[1320,1429,1404,1430]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB421455368FA7AFA52" box="[1320,1429,1404,1430]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
:
<figureCitation id="1347BF9C332CFFB424FA53B4FF03FA7F" box="[151,236,1440,1467]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1958,1982]" captionTargetBox="[152,1436,489,1934]" captionTargetId="figure-177@7.[151,1436,489,1935]" captionTargetPageId="7" captionText="FIGURES 25–30. SEM photographs of Speovelia maritima Esaki. 25–28, Male; 29–30, female. 25, 29, Head to abdomen; 26, thorax and abdomen; 27, habitus; 28, hind tibia and tarsus; 30, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010412" httpUri="https://zenodo.org/record/10010412/files/figure.png" pageId="1" pageNumber="469">Fig. 25</figureCitation>
); eyes with dense setae (
<figureCitation id="1347BF9C332CFFB4266153B4FDB0FA7E" box="[524,607,1440,1466]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 21</figureCitation>
) (with two short setae in
<taxonomicName id="4C7CD89A332CFFB427EA53B4FC1BFA7E" authorityName="Esaki" authorityYear="1929" box="[903,1012,1440,1466]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427EA53B4FC1BFA7E" box="[903,1012,1440,1466]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); antennae shorter than body (longer than body in
<taxonomicName id="4C7CD89A332CFFB4255E53D0FE4FFA1A" authorityName="Esaki" authorityYear="1929" box="[307,416,1476,1502]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB4255E53D0FE4FFA1A" box="[307,416,1476,1502]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); antennae without spines (with spines in
<taxonomicName id="4C7CD89A332CFFB427E653D0FC17FA1A" authorityName="Esaki" authorityYear="1929" box="[907,1016,1476,1502]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB427E653D0FC17FA1A" box="[907,1016,1476,1502]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
:
<figureCitation id="1347BF9C332CFFB4206653D0FB8FFA1B" box="[1035,1120,1476,1503]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1958,1982]" captionTargetBox="[152,1436,489,1934]" captionTargetId="figure-177@7.[151,1436,489,1935]" captionTargetPageId="7" captionText="FIGURES 25–30. SEM photographs of Speovelia maritima Esaki. 25–28, Male; 29–30, female. 25, 29, Head to abdomen; 26, thorax and abdomen; 27, habitus; 28, hind tibia and tarsus; 30, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010412" httpUri="https://zenodo.org/record/10010412/files/figure.png" pageId="1" pageNumber="469">Fig. 25</figureCitation>
); labium extends to middle of abdomen (to mesocoxae in
<taxonomicName id="4C7CD89A332CFFB4259C53FCFDB1F9C6" authorityName="Esaki" authorityYear="1929" box="[497,606,1512,1538]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB4259C53FCFDB1F9C6" box="[497,606,1512,1538]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); ventral head weakly carinate (
<figureCitation id="1347BF9C332CFFB427A053FCFBCFF9C7" box="[973,1056,1512,1539]" captionStart="FIGURES 15–18" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[191,1394,525,1915]" captionTargetId="figure-186@5.[191,1395,525,1915]" captionTargetPageId="5" captionText="FIGURES 15–18. Head, second gonocoxa, ovipositor and gynatrial complex of Nagisavelia hikarui sp. nov. female. 15, Head, ventral view; 16, second gonocoxa and gonoplac; 17, ovipositor (first gonapophysis), ventrolateral view; 18, gynatrial complex. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10010407" httpUri="https://zenodo.org/record/10010407/files/figure.png" pageId="1" pageNumber="469">Fig. 15</figureCitation>
) (simple in
<taxonomicName id="4C7CD89A332CFFB420C653FCFAF7F9C6" authorityName="Esaki" authorityYear="1929" box="[1195,1304,1512,1538]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB420C653FCFAF7F9C6" box="[1195,1304,1512,1538]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); spines on legs absent but hind tibia weak (strong in
<taxonomicName id="4C7CD89A332CFFB426015018FD36F9E2" authorityName="Esaki" authorityYear="1929" box="[620,729,1548,1574]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB426015018FD36F9E2" box="[620,729,1548,1574]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
); metatarsal segment length III&gt; II&gt; I (
<figureCitation id="1347BF9C332CFFB420CC5018FB1DF9E2" box="[1185,1266,1548,1574]" captionStart="FIGURES 19–24" captionStartId="6.[151,264,1650,1674]" captionTargetBox="[151,1430,181,1626]" captionTargetId="figure-17@6.[151,1436,181,1627]" captionTargetPageId="6" captionText="FIGURES 19–24. Scanning electron microscope (SEM) photographs of Nagisavelia hikarui sp. nov. 19–22, Male; 23–24, female. 19, 23, Head to abdomen; 20, thorax and abdomen, 21, eye, lateral view; 22, hind tibia and tarsus; 24, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010410" httpUri="https://zenodo.org/record/10010410/files/figure.png" pageId="1" pageNumber="469">Fig. 22</figureCitation>
) (II&gt; III&gt; I in
<taxonomicName id="4C7CD89A332CFFB424FA5024FEEBF98E" authorityName="Esaki" authorityYear="1929" box="[151,260,1584,1610]" class="Insecta" family="Mesoveliidae" genus="Speovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B9087F0B332CFFB424FA5024FEEBF98E" box="[151,260,1584,1610]" italics="true" pageId="1" pageNumber="469">Speovelia</emphasis>
</taxonomicName>
:
<figureCitation id="1347BF9C332CFFB425795024FE87F98E" box="[276,360,1584,1610]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1958,1982]" captionTargetBox="[152,1436,489,1934]" captionTargetId="figure-177@7.[151,1436,489,1935]" captionTargetPageId="7" captionText="FIGURES 25–30. SEM photographs of Speovelia maritima Esaki. 25–28, Male; 29–30, female. 25, 29, Head to abdomen; 26, thorax and abdomen; 27, habitus; 28, hind tibia and tarsus; 30, abdomen." figureDoi="http://doi.org/10.5281/zenodo.10010412" httpUri="https://zenodo.org/record/10010412/files/figure.png" pageId="1" pageNumber="469">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366F092332CFFB424AA5040FDF5F956" pageId="1" pageNumber="469" type="etymology">
<paragraph id="8BC3A319332CFFB424AA5040FDF5F956" blockId="1.[151,1437,612,1683]" pageId="1" pageNumber="469">
<emphasis id="B9087F0B332CFFB424AA5040FEA1F9AA" bold="true" box="[199,334,1620,1646]" pageId="1" pageNumber="469">Etymology.</emphasis>
The genus name is derived from the habitat “Nagisa” of the
<typeStatus id="54C71DBB332CFFB420645041FBD6F9AB" box="[1033,1081,1621,1647]" pageId="1" pageNumber="469">type</typeStatus>
species. In Japanese, “Nagisa” means the water edges of beaches.
</paragraph>
</subSubSection>
</treatment>
</document>
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