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<document id="F8F436BE949A39D600CF8CC58617A19B" ID-CLB-Dataset="29197" ID-DOI="10.11646/zootaxa.4479.1.1" ID-GBIF-Dataset="73cc6f7d-b3ec-40c5-adc7-52b9e0cbf236" ID-ISSN="1175-5326" ID-Zenodo-Dep="1452913" ID-ZooBank="93C84828-6EEF-4758-BEA1-97EEEF115245" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539087676732" checkinUser="plazi" docAuthor="Eiseman, Charles S. &amp; Lonsdale, Owen" docDate="2018" docId="03D287EFFFEDE404A8E5567345AEFA7C" docLanguage="en" docName="zootaxa.4479.1.1.pdf" docOrigin="Zootaxa 4479 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phytomyza palmeri Eiseman &amp; Lonsdale 2018, spec. nov." docType="treatment" docVersion="8" lastPageNumber="79" masterDocId="FFEBFF97FFA0E44AA872526F4103FFE1" masterDocTitle="New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species" masterLastPageNumber="156" masterPageNumber="1" pageNumber="78" updateTime="1698570476436" updateUser="ExternalLinkService">
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<mods:title id="160C8697C651666218E0C03412DB1397">New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species</mods:title>
</mods:titleInfo>
<mods:name id="BDAE1503BB4EAE3CC8D3B59F54DF5091" type="personal">
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<mods:namePart id="F6EE700B7298AC882293E134BBD52AFA">Eiseman, Charles S.</mods:namePart>
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<mods:namePart id="27D6B93EA3A6EFFB3E91FDD0A9BEF772">Lonsdale, Owen</mods:namePart>
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<mods:relatedItem id="897D51F73633BAF8EF5D236FB384C56B" type="host">
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<mods:title id="C5BC8664164D0BFF5C9ADC0FD6542C8F">Zootaxa</mods:title>
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<mods:date id="2D714BB3326C131412F02AD1AAFDEEF2">2018</mods:date>
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<mods:number id="96F2F83D0DCF752173C897B447F579A5">2018-09-14</mods:number>
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<mods:number id="6145ADAC64F31E27AEED40BEE7374053">4479</mods:number>
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<mods:classification id="02605D19ADFD9FA349842CED4215FAC6">journal article</mods:classification>
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<treatment id="03D287EFFFEDE404A8E5567345AEFA7C" ID-DOI="http://doi.org/10.5281/zenodo.5997917" ID-GBIF-Taxon="148663622" ID-Zenodo-Dep="5997917" LSID="urn:lsid:plazi:treatment:03D287EFFFEDE404A8E5567345AEFA7C" httpUri="http://treatment.plazi.org/id/03D287EFFFEDE404A8E5567345AEFA7C" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
<subSubSection id="C3616572FFEDE407A8E556734081FBB6" pageId="77" pageNumber="78" type="nomenclature">
<paragraph id="8BC436F9FFEDE407A8E5567340FDFBD7" blockId="77.[151,510,1052,1111]" box="[151,510,1052,1078]" pageId="77" pageNumber="78">
<heading id="D08C8195FFEDE407A8E5567340FDFBD7" bold="true" box="[151,510,1052,1078]" fontSize="11" level="1" pageId="77" pageNumber="78" reason="1">
<emphasis id="B90FEAEBFFEDE407A8E5567340FDFBD7" bold="true" box="[151,510,1052,1078]" pageId="77" pageNumber="78">
<taxonomicName id="4C7B4D7AFFEDE407A8E55673407CFBD7" ID-CoL="8DCN6" authority="Eiseman &amp; Lonsdale, 2018" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[151,383,1052,1078]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="palmeri" status="sp. nov.">
<emphasis id="B90FEAEBFFEDE407A8E55673407CFBD7" bold="true" box="[151,383,1052,1078]" italics="true" pageId="77" pageNumber="78">Phytomyza palmeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C5790FFEDE407A9F4567340FDFBD7" box="[390,510,1052,1078]" pageId="77" pageNumber="78" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC436F9FFEDE407A8E556514081FBB6" blockId="77.[151,510,1052,1111]" box="[151,386,1086,1111]" pageId="77" pageNumber="78">
(
<figureCitation id="13402A7CFFEDE407A8ED56514009FBB6" box="[159,266,1086,1111]" captionStart="FIGURES 189199" captionStartId="121.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@121.[151,1437,193,1639]" captionTargetPageId="121" captionText="FIGURES 189199. leaf mines; 189: Phytomyza hatfieldae spec. nov. in Osmorhiza claytonii; 190: P. hydrophyllivora spec. nov. in Hydrophyllum canadense; 191: P. hydrophyllivora in Hydrophyllum virginianum (yellow puparium at tip of leaf serration, upper right); 192: P. linnaeae (Griffiths) in Linnaea borealis (transmitted light); 193: P. loewii Hendel in Clematis virginiana; 194: P. minuscula Goureau in Thalictrum pubescens; 195: P. omlandi Scheffer &amp; Lonsdale in Gelsemium sempervirens; 196: P. palmeri spec. nov. in Symphoricarpos orbiculatus (transmitted light; puparium at upper left); 197: P. palustris spec. nov. in Micranthes pensylvanica; 198: same, five brown puparia in blisters on lower leaf surface (whitish object along left side of midrib is a coniopterygid cocoon); 199: P. penstemonis Spencer in Penstemon digitalis." httpUri="https://zenodo.org/record/1452959/files/figure.png" pageId="77" pageNumber="78">Figs. 196</figureCitation>
,
<figureCitation id="13402A7CFFEDE407A9645650407AFBB6" box="[278,377,1087,1111]" captionStart="FIGURES 343347" captionStartId="146.[151,264,1812,1835]" captionTargetBox="[182,1400,225,1761]" captionTargetId="figure@146.[151,1435,193,1792]" captionTargetPageId="146" captionText="FIGURES 343347. Phytomyza palmeri spec. nov., male genitalia; 343: external components, posterior; 344: postgonite, left lateral; 345: hypandrium, ventral; 346: distiphallus, ventral; 347: phallus, left lateral." httpUri="https://zenodo.org/record/1453009/files/figure.png" pageId="77" pageNumber="78">343347</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE407A8E556E9444AFACE" pageId="77" pageNumber="78" type="materials_examined">
<paragraph id="8BC436F9FFEDE407A8E556E942BAFB22" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<materialsCitation id="3B133CA4FFEDE407A8E556E942B7FB22" ID-GBIF-Occurrence="1915273650" collectingDate="2015-11-20" collectingDateMax="2015-11-30" collectingDateMin="2015-11-20" collectionCode="CNC" collectorName="M. W. Palmer" county="Payne Co." latitude="36.01434" location="Mehan" longLatPrecision="1" longitude="-96.99674" pageId="77" pageNumber="78" specimenCode="CNC634997" specimenCount="1" specimenCount-male="1" stateProvince="OKLAHOMA" typeStatus="holotype">
<emphasis id="B90FEAEBFFEDE407A8E556E940C9FB7E" bold="true" box="[151,458,1158,1183]" pageId="77" pageNumber="78">
<typeStatus id="54C0885BFFEDE407A8E556E94005FB7E" box="[151,262,1158,1183]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
.
<collectingRegion id="49BFF81BFFEDE407A96656E940C6FB7E" box="[276,453,1158,1183]" country="United States of America" name="Oklahoma" pageId="77" pageNumber="78">OKLAHOMA</collectingRegion>
:
</emphasis>
<collectingCounty id="62A54E75FFEDE407A9A156E84348FB7E" box="[467,587,1159,1183]" pageId="77" pageNumber="78">Payne Co.</collectingCounty>
,
<location id="8EA46022FFEDE407AA2B56E943A9FB7E" LSID="urn:lsid:plazi:treatment:03D287EFFFEDE404A8E5567345AEFA7C:8EA46022FFEDE407AA2B56E943A9FB7E" box="[601,682,1158,1183]" county="Payne Co." latitude="36.01434" longLatPrecision="1" longitude="-96.99674" name="Mehan" pageId="77" pageNumber="78" stateProvince="OKLAHOMA">Mehan</location>
,
<geoCoordinate id="EE4F503EFFEDE407AAC556E94259FB7F" box="[695,858,1158,1183]" direction="north" orientation="latitude" pageId="77" pageNumber="78" precision="1" value="36.01434">36.014339° N</geoCoordinate>
,
<geoCoordinate id="EE4F503EFFEDE407AB1556E9450EFB7F" box="[871,1037,1158,1183]" direction="west" orientation="longitude" pageId="77" pageNumber="78" precision="1" value="-96.99674">96.996744° W</geoCoordinate>
,
<collectingDate id="EF81E9D1FFEDE407AC6956E94596FB7E" box="[1051,1173,1158,1183]" pageId="77" pageNumber="78" value="2015-11-20">
<date id="FFC51039FFEDE407AC6956E94596FB7E" box="[1051,1173,1158,1183]" pageId="77" pageNumber="78" value="2015-11-20">20.xi.2015</date>
</collectingDate>
, em.
<collectingDate id="EF81E9D1FFEDE407ACA756E94453FB7E" box="[1237,1360,1158,1183]" pageId="77" pageNumber="78" value="2015-11-30">
<date id="FFC51039FFEDE407ACA756E94453FB7E" box="[1237,1360,1158,1183]" pageId="77" pageNumber="78" value="2015-11-30">30.xi.2015</date>
</collectingDate>
,
<collectorName id="268E532FFFEDE407AD2F56E841EBFB22" pageId="77" pageNumber="78">M.W. Palmer</collectorName>
, ex
<taxonomicName id="4C7B4D7AFFEDE407A96656C54350FB22" box="[276,595,1194,1219]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="77" pageNumber="78" phylum="Tracheophyta" rank="species" species="orbiculatus">
<emphasis id="B90FEAEBFFEDE407A96656C54350FB22" box="[276,595,1194,1219]" italics="true" pageId="77" pageNumber="78">Symphoricarpos orbiculatus</emphasis>
</taxonomicName>
, #CSE2655,
<specimenCode id="DBDD9E82FFEDE407AA9A56C54276FB22" box="[744,885,1194,1219]" collectionCode="CNC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="77" pageNumber="78">CNC634997</specimenCode>
(
<specimenCount id="9D7DFD70FFEDE407ABF556C442A8FB22" box="[903,939,1195,1219]" pageId="77" pageNumber="78" type="male">1♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC436F9FFEDE407A8E556A1444AFACE" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8E556A140DDFB06" bold="true" box="[151,478,1230,1255]" pageId="77" pageNumber="78">
<typeStatus id="54C0885BFFEDE407A8E556A14012FB06" box="[151,273,1230,1255]" pageId="77" pageNumber="78" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="49BFF81BFFEDE407A95B56A140D9FB06" box="[297,474,1230,1255]" country="United States of America" name="Oklahoma" pageId="77" pageNumber="78">OKLAHOMA</collectingRegion>
:
</emphasis>
same collection as
<typeStatus id="54C0885BFFEDE407AA9856A1424CFB06" box="[746,847,1230,1255]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED6AAE3CFFEDE407AB1456A142A3FB06" box="[870,928,1230,1255]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="77" pageNumber="78">CNC</collectionCode>
634987634998 (
<specimenCount id="9D7DFD70FFEDE407AC0056A04596FB06" box="[1138,1173,1231,1255]" pageId="77" pageNumber="78" type="male">3♂</specimenCount>
<specimenCount id="9D7DFD70FFEDE407ACDA56A045CFFB06" box="[1192,1228,1231,1255]" pageId="77" pageNumber="78" type="female">7♀</specimenCount>
);
<date id="FFC51039FFEDE407AC9D56A14458FB06" box="[1263,1371,1230,1255]" pageId="77" pageNumber="78" value="2015-11-07">7.xi.2015</date>
, em.
<date id="FFC51039FFEDE407A8E5569D4012FAEA" box="[151,273,1266,1291]" pageId="77" pageNumber="78" value="2015-11-17">17.xi.2015</date>
, M.W. Palmer, ex
<taxonomicName id="4C7B4D7AFFEDE407AA64569D4261FAEA" box="[534,866,1266,1291]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="77" pageNumber="78" phylum="Tracheophyta" rank="species" species="orbiculatus">
<emphasis id="B90FEAEBFFEDE407AA64569D4261FAEA" box="[534,866,1266,1291]" italics="true" pageId="77" pageNumber="78">Symphoricarpos orbiculatus</emphasis>
</taxonomicName>
, #CSE2163,
<collectionCode id="ED6AAE3CFFEDE407AC61569D454DFAEA" box="[1043,1102,1266,1291]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="77" pageNumber="78">CNC</collectionCode>
564616,
<collectionCode id="ED6AAE3CFFEDE407ACCE569D45F5FAEA" box="[1212,1270,1266,1291]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="77" pageNumber="78">CNC</collectionCode>
564617 (
<specimenCount id="9D7DFD70FFEDE407AD1B569C448FFAEA" box="[1385,1420,1267,1291]" pageId="77" pageNumber="78" type="female">2♀</specimenCount>
);
<date id="FFC51039FFEDE407A8E55779401BFACE" box="[151,280,1302,1327]" pageId="77" pageNumber="78" value="2015-12-11">11.xii.2015</date>
, em.
<date id="FFC51039FFEDE407A926577940D6FACE" box="[340,469,1302,1327]" pageId="77" pageNumber="78" value="2015-12-18">18.xii.2015</date>
, M.W. Palmer, ex
<taxonomicName id="4C7B4D7AFFEDE407AAD6577942E0FACE" box="[676,995,1302,1327]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="77" pageNumber="78" phylum="Tracheophyta" rank="species" species="orbiculatus">
<emphasis id="B90FEAEBFFEDE407AAD6577942E0FACE" box="[676,995,1302,1327]" italics="true" pageId="77" pageNumber="78">Symphoricarpos orbiculatus</emphasis>
</taxonomicName>
, #CSE2656,
<collectionCode id="ED6AAE3CFFEDE407AC0A577945B2FACE" box="[1144,1201,1302,1327]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="77" pageNumber="78">CNC</collectionCode>
634770 (
<specimenCount id="9D7DFD70FFEDE407AD6557784438FACE" box="[1303,1339,1303,1327]" pageId="77" pageNumber="78" type="male">1♂</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE407A8B557554394FA96" pageId="77" pageNumber="78" type="etymology">
<paragraph id="8BC436F9FFEDE407A8B557554394FA96" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B55755404DFAB2" bold="true" box="[199,334,1338,1363]" pageId="77" pageNumber="78">Etymology.</emphasis>
This species is named for Michael W. Palmer, botanist, who collected the
<typeStatus id="54C0885BFFEDE407ACE8575445C9FAB2" box="[1178,1226,1339,1363]" pageId="77" pageNumber="78">type</typeStatus>
series as well as a number of other specimens used in this study.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE407A8B557ED4509F9E6" pageId="77" pageNumber="78" type="biology_ecology">
<paragraph id="8BC436F9FFEDE407A8B557ED426BFA7A" blockId="77.[151,1437,1158,2011]" box="[199,872,1410,1435]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B557ED4005FA7A" bold="true" box="[199,262,1410,1435]" pageId="77" pageNumber="78">Host.</emphasis>
<taxonomicName id="4C7B4D7AFFEDE407A97F57ED40B6FA7A" box="[269,437,1410,1435]" class="Magnoliopsida" family="Caprifoliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="77" pageNumber="78" phylum="Tracheophyta" rank="family">Caprifoliaceae</taxonomicName>
:
<taxonomicName id="4C7B4D7AFFEDE407A9CD57ED426BFA7A" authority="Moench." authorityName="Moench." box="[447,872,1410,1435]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="77" pageNumber="78" phylum="Tracheophyta" rank="species" species="orbiculatus">
<emphasis id="B90FEAEBFFEDE407A9CD57ED43FEFA7A" box="[447,765,1410,1435]" italics="true" pageId="77" pageNumber="78">Symphoricarpos orbiculatus</emphasis>
Moench.
</taxonomicName>
</paragraph>
<paragraph id="8BC436F9FFEDE407A8B557C94509F9E6" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B557C94048FA5E" bold="true" box="[199,331,1446,1471]" pageId="77" pageNumber="78">Leaf mine.</emphasis>
(
<figureCitation id="13402A7CFFEDE407A92C57C940C1FA5E" box="[350,450,1446,1471]" captionStart="FIGURES 189199" captionStartId="121.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@121.[151,1437,193,1639]" captionTargetPageId="121" captionText="FIGURES 189199. leaf mines; 189: Phytomyza hatfieldae spec. nov. in Osmorhiza claytonii; 190: P. hydrophyllivora spec. nov. in Hydrophyllum canadense; 191: P. hydrophyllivora in Hydrophyllum virginianum (yellow puparium at tip of leaf serration, upper right); 192: P. linnaeae (Griffiths) in Linnaea borealis (transmitted light); 193: P. loewii Hendel in Clematis virginiana; 194: P. minuscula Goureau in Thalictrum pubescens; 195: P. omlandi Scheffer &amp; Lonsdale in Gelsemium sempervirens; 196: P. palmeri spec. nov. in Symphoricarpos orbiculatus (transmitted light; puparium at upper left); 197: P. palustris spec. nov. in Micranthes pensylvanica; 198: same, five brown puparia in blisters on lower leaf surface (whitish object along left side of midrib is a coniopterygid cocoon); 199: P. penstemonis Spencer in Penstemon digitalis." httpUri="https://zenodo.org/record/1452959/files/figure.png" pageId="77" pageNumber="78">Fig. 196</figureCitation>
) Upper surface, gradually widening, at least sometimes linear throughout, but often contorted to form a secondary blotch. The black frass is in closely spaced grains or beaded strips, allowing the larvas path to be traced even when the mine forms a nearly continuous blotch.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE407A8B5547D400EF9AE" pageId="77" pageNumber="78" type="description">
<paragraph id="8BC436F9FFEDE407A8B5547D400EF9AE" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B5547D404BF9CA" bold="true" box="[199,328,1554,1579]" pageId="77" pageNumber="78">Puparium.</emphasis>
Brown; formed within the mine, its anterior spiracles projecting ventrally through the upper epidermis.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE407A8B55435406CF93E" pageId="77" pageNumber="78" type="distribution">
<paragraph id="8BC436F9FFEDE407A8B5543540DDF992" blockId="77.[151,1437,1158,2011]" box="[199,478,1626,1651]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B554354063F992" bold="true" box="[199,352,1626,1651]" pageId="77" pageNumber="78">Distribution.</emphasis>
<collectingCountry id="F36C7669FFEDE407A915543440A0F992" box="[359,419,1627,1651]" name="United States of America" pageId="77" pageNumber="78">USA</collectingCountry>
: OK.
</paragraph>
<paragraph id="8BC436F9FFEDE407A8B55411406CF93E" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B5541140A3F976" bold="true" box="[199,416,1662,1687]" pageId="77" pageNumber="78">Adult description.</emphasis>
Wing length
<quantity id="4C839B1CFFEDE407AA4A541043BBF976" box="[568,696,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="77" pageNumber="78" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9mm</quantity>
(
<specimenCount id="9D7DFD70FFEDE407AABA541043DCF976" box="[712,735,1663,1687]" pageId="77" pageNumber="78" type="male"></specimenCount>
),
<quantity id="4C839B1CFFEDE407AA8754104277F976" box="[757,884,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="77" pageNumber="78" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.92.2mm</quantity>
(
<specimenCount id="9D7DFD70FFEDE407ABF554104299F976" box="[903,922,1663,1687]" pageId="77" pageNumber="78" type="female"></specimenCount>
). Eye height divided by gena height: 5.66.1. First flagellomere rounded; covered with longer hairs (except at base). Orbital plate ill-defined. Notum pruinose. Vein dm-cu absent.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEDE404A8B554854060FD64" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="description">
<paragraph id="8BC436F9FFEDE407A8B554854557F8AA" blockId="77.[151,1437,1158,2011]" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B554854044F8E2" box="[199,327,1770,1795]" italics="true" pageId="77" pageNumber="78">Chaetotaxy</emphasis>
: Two ors, two ori (anterior ori 1/31/2 length). Ocellar seta subequal to ors, postvertical slightly longer. Cephalic setae thinner. Four dorsocentral setae decreasing in length anteriorly, second seta approximately 3/ 5 first seta, fourth barely larger than setulae; only first seta strong with others thinner.
</paragraph>
<paragraph id="8BC436F9FFEDE404A8B55539408EFE69" blockId="77.[151,1437,1158,2011]" lastBlockId="78.[151,1437,151,1437]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
<emphasis id="B90FEAEBFFEDE407A8B555394041F88E" box="[199,322,1878,1903]" italics="true" pageId="77" pageNumber="78">Coloration</emphasis>
: Setae dark brown, genal setae slightly paler. Head mostly yellow; scape and pedicel brownish to light brown, first flagellomere slightly darker brown to dark brown; orbital plate light yellow; posterolateral corner of frons dark brown to base of inner vertical; head below antenna light yellow with center and ventrolateral region of face deeper yellow, venter of gena with light brown line and clypeus brownish; ocellar spot slightly larger than tubercle, concave posteriorly, region between spot and back of head brownish. Thorax brown with grayish to brownish pruinosity that is slightly bluish or coppery on anterior half of notum; postpronotum dirty white with brown spot; notopleuron paler brown with yellowish mottling; pleuron paler brown, anepisternum yellowish dorsally and sometimes yellow mottled. Calypter white with hairs light brown. Haltere white. Legs mostly light brown to brown, paler than thorax; fore coxa yellow with base brown; yellow streaking dominant on fore femur, less prominent on mid femur; fore tibia and tarsi yellowish, mid tibia and tarsi slightly less so. Abdomen brown with epandrium paler.
</paragraph>
<paragraph id="8BC436F9FFEEE404A8B553FB4060FD64" blockId="78.[151,1437,151,1437]" pageId="78" pageNumber="79">
<emphasis id="B90FEAEBFFEEE404A8B553FB4032FE4C" box="[199,305,404,429]" italics="true" pageId="78" pageNumber="79">Genitalia</emphasis>
: (
<figureCitation id="13402A7CFFEEE404A93B53FB40EFFE4D" box="[329,492,404,429]" captionStart="FIGURES 343347" captionStartId="146.[151,264,1812,1835]" captionTargetBox="[182,1400,225,1761]" captionTargetId="figure@146.[151,1435,193,1792]" captionTargetPageId="146" captionText="FIGURES 343347. Phytomyza palmeri spec. nov., male genitalia; 343: external components, posterior; 344: postgonite, left lateral; 345: hypandrium, ventral; 346: distiphallus, ventral; 347: phallus, left lateral." httpUri="https://zenodo.org/record/1453009/files/figure.png" pageId="78" pageNumber="79">Figs. 343347</figureCitation>
) Surstlyus small, rounded, fused to epandrium; epandrium with thin lobate process from posterior margin. Postgonite elongate, with narrow curved apex and single medial seta. Basiphallus divided into two dark band-like sclerites; basal arm on left sclerite wrapping around shaft; right sclerite dorsally fused to phallophorus and with rounded dorsal subbasal process. Hypophallus entirely membranous, with long anteromedial flagellum. Paraphallus absent. Mesophallus indistinct, incorporated into distiphallus as shallow basal stem. Distiphallus dark, flat, angled dorsally, medially divided past base into one pair of pointed lobes with paler distal annulations.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFEEE404A8B550E045AEFA7C" pageId="78" pageNumber="79" type="discussion">
<paragraph id="8BC436F9FFEEE404A8B550E04081FC45" blockId="78.[151,1437,151,1437]" pageId="78" pageNumber="79">
<emphasis id="B90FEAEBFFEEE404A8B550E0404CFD49" bold="true" box="[199,335,655,680]" pageId="78" pageNumber="79">Comments.</emphasis>
Three other Nearctic
<taxonomicName id="4C7B4D7AFFEEE404AA3B50E043C7FD49" box="[585,708,655,680]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEAEBFFEEE404AA3B50E043C7FD49" box="[585,708,655,680]" italics="true" pageId="78" pageNumber="79">Phytomyza</emphasis>
</taxonomicName>
species are known to mine leaves of
<taxonomicName id="4C7B4D7AFFEEE404AC0350E0442BFD49" authorityName="H.L. Duhamel du Monceau" authorityYear="1755" box="[1137,1320,655,680]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAEBFFEEE404AC0350E0442BFD49" box="[1137,1320,655,680]" italics="true" pageId="78" pageNumber="79">Symphoricarpos</emphasis>
</taxonomicName>
, all in the
<emphasis id="B90FEAEBFFEEE404A8E550DB4018FD2C" box="[151,283,692,717]" italics="true" pageId="78" pageNumber="79">periclymeni</emphasis>
superspecies. Their mines are discussed below under
<taxonomicName id="4C7B4D7AFFEEE404AB0650DB42ECFD2C" box="[884,1007,692,717]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEAEBFFEEE404AB0650DB42ECFD2C" box="[884,1007,692,717]" italics="true" pageId="78" pageNumber="79">Phytomyza</emphasis>
</taxonomicName>
sp. 4. These and other Caprifoliaceaemining species are best treated in
<bibRefCitation id="EFEA4B08FFEEE404AA6A50B843D4FD11" box="[536,727,727,752]" pageId="78" pageNumber="79" refId="ref101525" refString="Griffiths, G. C. D. (1974 a) Studies on boreal Agromyzidae (Diptera). V. On the genus Chromatomyia Hardy, with revision of Caprifoliaceae-mining species. Quaestiones Entomologicae, 10, 35 - 69." type="journal article">Griffiths (1974a)</bibRefCitation>
as part of the genus
<taxonomicName id="4C7B4D7AFFEEE404ABB750B8456FFD11" box="[965,1132,727,752]" class="Insecta" family="Agromyzidae" genus="Chromatomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEAEBFFEEE404ABB750B8456FFD11" box="[965,1132,727,752]" italics="true" pageId="78" pageNumber="79">Chromatomyia</emphasis>
</taxonomicName>
, where examination of the male genitalia is almost always required for species diagnosis. The hypophallus is usually present in the Caprifoliaceae-mining
<taxonomicName id="4C7B4D7AFFEEE404A9E851704316FCD9" box="[410,533,799,824]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEAEBFFEEE404A9E851704316FCD9" box="[410,533,799,824]" italics="true" pageId="78" pageNumber="79">Phytomyza</emphasis>
</taxonomicName>
, at least as a small vestigial sclerite (e.g.
<taxonomicName id="4C7B4D7AFFEEE404AB96514E45CAFCD9" authority="Hendel" authorityName="Hendel" authorityYear="1925" box="[996,1225,799,824]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="alpigenae">
<emphasis id="B90FEAEBFFEEE404AB96514E456DFCD9" box="[996,1134,799,824]" italics="true" pageId="78" pageNumber="79">P. alpigenae</emphasis>
Hendel
</taxonomicName>
), but it is absent in three known species—
<taxonomicName id="4C7B4D7AFFEEE404A9EF512A4310FCBC" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[413,531,836,861]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="palmeri">
<emphasis id="B90FEAEBFFEEE404A9EF512A4310FCBC" box="[413,531,836,861]" italics="true" pageId="78" pageNumber="79">P. palmeri</emphasis>
</taxonomicName>
on
<taxonomicName id="4C7B4D7AFFEEE404AA37512B43FFFCBC" authorityName="H.L. Duhamel du Monceau" authorityYear="1755" box="[581,764,836,861]" class="Magnoliopsida" family="Caprifoliaceae" genus="Symphoricarpos" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAEBFFEEE404AA37512B43FFFCBC" box="[581,764,836,861]" italics="true" pageId="78" pageNumber="79">Symphoricarpos</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D7AFFEEE404AB48512A42FEFCBD" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[826,1021,837,860]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sempervirentis">
<emphasis id="B90FEAEBFFEEE404AB48512A42FEFCBD" box="[826,1021,837,860]" italics="true" pageId="78" pageNumber="79">P. sempervirentis</emphasis>
</taxonomicName>
on
<taxonomicName id="4C7B4D7AFFEEE404AC5C512A4590FCBD" box="[1070,1171,837,860]" class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAEBFFEEE404AC5C512A4590FCBD" box="[1070,1171,837,860]" italics="true" pageId="78" pageNumber="79">Lonicera</emphasis>
</taxonomicName>
, both newly described here, and
<taxonomicName id="4C7B4D7AFFEEE404A97451064320FC61" authority="(Griffiths)" authorityName="Griffiths" box="[262,547,871,896]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrilineata">
<emphasis id="B90FEAEBFFEEE404A974510640A5FC61" box="[262,422,871,896]" italics="true" pageId="78" pageNumber="79">P. nigrilineata</emphasis>
(Griffiths)
</taxonomicName>
, also on
<taxonomicName id="4C7B4D7AFFEEE404AAF5510643EFFC61" box="[647,748,873,896]" class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Dipsacales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAEBFFEEE404AAF5510643EFFC61" box="[647,748,873,896]" italics="true" pageId="78" pageNumber="79">Lonicera</emphasis>
</taxonomicName>
(a minute hypophallus sclerite is erroneously depicted in the original description).
</paragraph>
<paragraph id="8BC436F9FFEEE404A8B551C040BEFAD1" blockId="78.[151,1437,151,1437]" pageId="78" pageNumber="79">
<taxonomicName id="4C7B4D7AFFEEE404A8B551C040A4FC29" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[199,423,943,968]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="palmeri">
<emphasis id="B90FEAEBFFEEE404A8B551C040A4FC29" box="[199,423,943,968]" italics="true" pageId="78" pageNumber="79">Phytomyza palmeri</emphasis>
</taxonomicName>
can be differentiated from the above by a wing length of
<quantity id="4C839B1CFFEEE404AC0D51DF4400FC29" box="[1151,1283,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="78" pageNumber="79" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9mm</quantity>
(
<specimenCount id="9D7DFD70FFEEE404AD7951DF441DFC29" box="[1291,1310,944,968]" pageId="78" pageNumber="79" type="male"></specimenCount>
) and
<quantity id="4C839B1CFFEEE404AD1951DF41F3FC0D" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="78" pageNumber="79" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9 2.2mm</quantity>
(
<specimenCount id="9D7DFD70FFEEE404A88B51BB4004FC0D" box="[249,263,980,1004]" pageId="78" pageNumber="79" type="female"></specimenCount>
), a narrower gena, a light yellow frons, a mostly white postpronotum, extensive yellow markings on the fore and mid femora, a short basiphallus and a distiphallus that is stouter and wider with distal annulations (
<figureCitation id="13402A7CFFEEE404AD16519841FDFBD5" captionStart="FIGURES 343347" captionStartId="146.[151,264,1812,1835]" captionTargetBox="[182,1400,225,1761]" captionTargetId="figure@146.[151,1435,193,1792]" captionTargetPageId="146" captionText="FIGURES 343347. Phytomyza palmeri spec. nov., male genitalia; 343: external components, posterior; 344: postgonite, left lateral; 345: hypandrium, ventral; 346: distiphallus, ventral; 347: phallus, left lateral." httpUri="https://zenodo.org/record/1453009/files/figure.png" pageId="78" pageNumber="79">Figs. 346, 347</figureCitation>
).
<taxonomicName id="4C7B4D7AFFEEE404A96656734338FBD5" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[276,571,1052,1077]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sempervirentis">
<emphasis id="B90FEAEBFFEEE404A96656734338FBD5" box="[276,571,1052,1077]" italics="true" pageId="78" pageNumber="79">Phytomyza sempervirentis</emphasis>
</taxonomicName>
is larger (wing length 2.12.2(
<specimenCount id="9D7DFD70FFEEE404ABD3567342B6FBD5" box="[929,949,1052,1076]" pageId="78" pageNumber="79" type="male"></specimenCount>
) and
<quantity id="4C839B1CFFEEE404AB8456734582FBD5" box="[1014,1153,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="78" pageNumber="79" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4mm</quantity>
(
<specimenCount id="9D7DFD70FFEEE404ACF85673459BFBD5" box="[1162,1176,1052,1076]" pageId="78" pageNumber="79" type="female"></specimenCount>
)), only the apex of the fore femur is yellow, and it has a higher gena, a brown frons, a brown postpronotum, a longer paler basiphallus and a more gracile distiphallus with one pair of lateromedial lobes (
<figureCitation id="13402A7CFFEEE404AB03560B4517FB9C" box="[881,1044,1124,1149]" captionStart="FIGURES 353359" captionStartId="148.[151,264,1820,1843]" captionTargetBox="[207,1423,222,1760]" captionTargetId="figure@148.[151,1436,193,1800]" captionTargetPageId="148" captionText="FIGURES 353359. Phytomyza sempervirentis spec. nov., male genitalia; 353: external components, posterior; 354: postgonite, left lateral; 355: hypandrium, ventral; 356: distiphallus, ventral (#CSE2553); 357: phallus, left lateral (#CSE2553); 358: distiphallus, ventral (#CSE279); 359: phallus, left lateral (#CSE279)." httpUri="https://zenodo.org/record/1453013/files/figure.png" pageId="78" pageNumber="79">Figs. 356-359</figureCitation>
).
<taxonomicName id="4C7B4D7AFFEEE404AC5B560B4432FB9C" box="[1065,1329,1124,1149]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrilineata">
<emphasis id="B90FEAEBFFEEE404AC5B560B4432FB9C" box="[1065,1329,1124,1149]" italics="true" pageId="78" pageNumber="79">Phytomyza nigrilineata</emphasis>
</taxonomicName>
is nearly identical to
<taxonomicName id="4C7B4D7AFFEEE404A96856E640DAFB41" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[282,473,1161,1184]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sempervirentis">
<emphasis id="B90FEAEBFFEEE404A96856E640DAFB41" box="[282,473,1161,1184]" italics="true" pageId="78" pageNumber="79">P. sempervirentis</emphasis>
</taxonomicName>
, but in the former there is yellowish mottling around the postpronotum, all femora are widely yellow apically (not just the fore femur), the distiphallus appears to have shorter, narrower distal arms with a darker accessory laterobasal process, and the single basal process of the distiphallus is stouter and as long as the distal section (based on the illustration in the original description, as the genitalia are now permanently embedded and only visible laterally).
</paragraph>
<paragraph id="8BC436F9FFEEE404A8B5575345AEFA7C" blockId="78.[151,1437,151,1437]" pageId="78" pageNumber="79">
The leaf mine most similar to that of
<taxonomicName id="4C7B4D7AFFEEE404AA1557534243FAB4" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[615,832,1340,1365]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="palmeri">
<emphasis id="B90FEAEBFFEEE404AA1557534243FAB4" box="[615,832,1340,1365]" italics="true" pageId="78" pageNumber="79">Phytomyza palmeri</emphasis>
</taxonomicName>
is apparently that of
<taxonomicName id="4C7B4D7AFFEEE404AC5F57524425FAB5" authority="Spencer" authorityName="Spencer" authorityYear="1969" box="[1069,1318,1340,1365]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caprifoliae">
<emphasis id="B90FEAEBFFEEE404AC5F575245C7FAB4" box="[1069,1220,1340,1365]" italics="true" pageId="78" pageNumber="79">P. caprifoliae</emphasis>
Spencer
</taxonomicName>
, but based on Griffiths (1974a) description and illustration for that species,
<taxonomicName id="4C7B4D7AFFEEE404AB05570E42E9FA99" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[887,1002,1375,1400]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="palmeri">
<emphasis id="B90FEAEBFFEEE404AB05570E42E9FA99" box="[887,1002,1375,1400]" italics="true" pageId="78" pageNumber="79">P. palmeri</emphasis>
</taxonomicName>
seems to have a more distinctly linear mine. It may not always be possible to distinguish the two should they prove to be sympatric.
</paragraph>
</subSubSection>
</treatment>
</document>