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<document id="FF4C60ABD85820B14E9C34971B4277EB" ID-DOI="10.11646/zootaxa.5361.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10084538" ID-ZooBank="D9AE18C9-A265-44F0-A0AF-E663A59B2EAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1698932362298" checkinUser="plazi" docAuthor="Zamani, Alireza, Darvishnia, Hamid &amp; Marusik, Yuri M." docDate="2023" docId="03C5CF42FF92864522B5FD36FECC2D49" docLanguage="en" docName="zootaxa.5361.3.3.pdf" docOrigin="Zootaxa 5361 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.3.3/52195" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Troglohyphantes megagynus Zamani &amp; Darvishnia &amp; Marusik 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="357" masterDocId="FFFCB73AFF9886492222FFEDFFCC2911" masterDocTitle="New data on cave spiders (Arachnida: Araneae) of Iran, with new species and records" masterLastPageNumber="366" masterPageNumber="345" pageNumber="355" updateTime="1699464889253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D13A2DD5F0A4A425117D6B90E3B44FAE">
<mods:title id="C7E3E0AD00F5582BA5C137ED0DC85B83">New data on cave spiders (Arachnida: Araneae) of Iran, with new species and records</mods:title>
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<mods:namePart id="78ED65F3F2657E6989A1E601E4F5BF1C">Zamani, Alireza</mods:namePart>
<mods:affiliation id="3CC2B3C1E330AFE8856729B4ACF51B96">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku, Finland.</mods:affiliation>
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<mods:namePart id="7F17B34F9655AE5A533A8C52D2F134D4">Darvishnia, Hamid</mods:namePart>
<mods:affiliation id="9C21335CA62892D5825E42FCADF4B101">Department of Biology, Payame Noor University, Tehran, Iran.</mods:affiliation>
</mods:name>
<mods:name id="8EC02A5ADD07C15B9374EAA2BB8DAD45" type="personal">
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<mods:namePart id="3B38FD881CF6FD18B5E5C29163808921">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="410800F35C431270AB45E4A5E931EA1E">Institute for Biological Problems of the North, FEB RAS, Portovaya Str. 18. Magadan 685000, Russia. &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:title id="2743B907EE699C01BA432E496E99CA96">Zootaxa</mods:title>
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<mods:date id="A99993098383C398F24CF7833C9846A3">2023</mods:date>
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<mods:number id="C9DB9EF680019BDF87DEBF0652302165">2023-11-02</mods:number>
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<mods:number id="0756BE50A0B54748E5E523DC1A3EEEA6">5361</mods:number>
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<mods:identifier id="2461559EC0B2A1001BB32543800D8086" type="DOI">10.11646/zootaxa.5361.3.3</mods:identifier>
<mods:identifier id="A4CA512520A9E7202B89F3B58E0F9116" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="56F3600C6371DB3A3273297B85FDFECC" type="Zenodo-Dep">10084538</mods:identifier>
<mods:identifier id="86885D4264141B31C06FF5CD8569EA03" type="ZooBank">D9AE18C9-A265-44F0-A0AF-E663A59B2EAF</mods:identifier>
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<treatment id="03C5CF42FF92864522B5FD36FECC2D49" LSID="urn:lsid:plazi:treatment:03C5CF42FF92864522B5FD36FECC2D49" httpUri="http://treatment.plazi.org/id/03C5CF42FF92864522B5FD36FECC2D49" lastPageId="12" lastPageNumber="357" pageId="10" pageNumber="355">
<subSubSection id="C3762DDFFF92864322B5FD36FC8C2BE4" box="[151,832,731,758]" pageId="10" pageNumber="355" type="nomenclature">
<paragraph id="8BD37E54FF92864322B5FD36FC8C2BE4" blockId="10.[151,832,731,793]" box="[151,832,731,758]" pageId="10" pageNumber="355">
<heading id="D09BC938FF92864322B5FD36FC8C2BE4" bold="true" box="[151,832,731,758]" fontSize="11" level="1" pageId="10" pageNumber="355" reason="1">
<emphasis id="B918A246FF92864322B5FD36FC8C2BE4" bold="true" box="[151,832,731,758]" pageId="10" pageNumber="355">
<taxonomicName id="4C6C05D7FF92864322B5FD36FD3D2BE4" authority="Zamani &amp; Marusik" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[151,753,731,758]" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="megagynus" status="sp. nov.">
<emphasis id="B918A246FF92864322B5FD36FE382BE4" bold="true" box="[151,500,731,758]" italics="true" pageId="10" pageNumber="355">Troglohyphantes megagynus</emphasis>
Zamani &amp; Marusik
</taxonomicName>
,
<taxonomicNameLabel id="A22B1F3DFF92864320DCFD36FC8C2BE4" box="[766,832,731,757]" pageId="10" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF92864322B5FD12FEA32A08" box="[151,367,767,793]" pageId="10" pageNumber="355" type="description">
<paragraph id="8BD37E54FF92864322B5FD12FEA32A08" blockId="10.[151,832,731,793]" box="[151,367,767,793]" pageId="10" pageNumber="355">
<figureCitation id="135762D1FF92864322B5FD12FED82A08" box="[151,276,767,793]" captionStart="FIGURE 9" captionStartId="10.[151,250,2011,2035]" captionTargetBox="[169,1377,817,1981]" captionTargetId="figure-137@10.[151,1436,817,1987]" captionTargetPageId="10" captionText="FIGURE 9. Habitus of Troglohyphantes megagynus sp. n., female. A lateral view; B dorsal view; C prosoma, anterior view." figureDoi="http://doi.org/10.5281/zenodo.10084556" httpUri="https://zenodo.org/record/10084556/files/figure.png" pageId="10" pageNumber="355">Figs 9A–C</figureCitation>
,
<figureCitation id="135762D1FF928643233DFD12FEA32A08" box="[287,367,767,793]" captionStart="FIGURE 10" captionStartId="11.[151,250,1940,1964]" captionTargetBox="[160,1425,191,1908]" captionTargetId="figure-18@11.[151,1436,181,1916]" captionTargetPageId="11" captionText="FIGURE 10. Female of Troglohyphantes megagynus sp. n. A epigyne, ventral view; B same, dorsal view; C same, caudal view; D same, lateral view; E, abdomen, ventral view. Scale bars: 0.2 mm. Abbreviations: Be—base of epigyne, DPS—distal part of scape, Eg—epigynal groove, Ll—lateral lobe, PMP—posterior median plate, PPS—proximal part of scape, Sr—stretcher." figureDoi="http://doi.org/10.5281/zenodo.10084558" httpUri="https://zenodo.org/record/10084558/files/figure.png" pageId="10" pageNumber="355">10A–E</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF132EDCFF92864322B5F836FA492EE2" ID-DOI="http://doi.org/10.5281/zenodo.10084556" ID-Zenodo-Dep="10084556" box="[151,1413,2011,2035]" httpUri="https://zenodo.org/record/10084556/files/figure.png" pageId="10" pageNumber="355" startId="10.[151,250,2011,2035]" targetBox="[169,1377,817,1981]" targetPageId="10" targetType="figure">
<paragraph id="8BD37E54FF92864322B5F836FA492EE2" blockId="10.[151,1413,2011,2035]" box="[151,1413,2011,2035]" pageId="10" pageNumber="355">
<emphasis id="B918A246FF92864322B5F836FEDF2EE2" bold="true" box="[151,275,2011,2035]" pageId="10" pageNumber="355">FIGURE 9.</emphasis>
Habitus of
<taxonomicName id="4C6C05D7FF92864323A8F836FD602EE2" authority="Zamani &amp; Darvishnia &amp; Marusik, 2023" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[394,684,2011,2035]" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="megagynus" status="sp. nov.">
<emphasis id="B918A246FF92864323A8F836FD602EE2" box="[394,684,2011,2035]" italics="true" pageId="10" pageNumber="355">Troglohyphantes megagynus</emphasis>
</taxonomicName>
<emphasis id="B918A246FF9286432090F836FD272EE2" bold="true" box="[690,747,2011,2035]" pageId="10" pageNumber="355">
<taxonomicNameLabel id="A22B1F3DFF9286432090F836FD272EE2" box="[690,747,2011,2035]" pageId="10" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female. A lateral view; B dorsal view; C prosoma, anterior view.
</paragraph>
</caption>
<caption id="DF132EDCFF93864222B5F879FA932EE5" ID-DOI="http://doi.org/10.5281/zenodo.10084558" ID-Zenodo-Dep="10084558" httpUri="https://zenodo.org/record/10084558/files/figure.png" pageId="11" pageNumber="356" startId="11.[151,250,1940,1964]" targetBox="[160,1425,191,1908]" targetPageId="11" targetType="figure">
<paragraph id="8BD37E54FF93864222B5F879FA932EE5" blockId="11.[151,1436,1940,2036]" pageId="11" pageNumber="356">
<emphasis id="B918A246FF93864222B5F879FED32EBD" bold="true" box="[151,287,1940,1964]" pageId="11" pageNumber="356">FIGURE 10.</emphasis>
Female of
<taxonomicName id="4C6C05D7FF93864223ADF879FD7C2EBD" authority="Zamani &amp; Darvishnia &amp; Marusik, 2023" authorityName="Zamani &amp; Darvishnia &amp; Marusik" authorityYear="2023" box="[399,688,1940,1964]" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="megagynus" status="sp. nov.">
<emphasis id="B918A246FF93864223ADF879FD7C2EBD" box="[399,688,1940,1964]" italics="true" pageId="11" pageNumber="356">Troglohyphantes megagynus</emphasis>
</taxonomicName>
<emphasis id="B918A246FF9386422094F879FD222EBD" bold="true" box="[694,750,1940,1964]" pageId="11" pageNumber="356">
<taxonomicNameLabel id="A22B1F3DFF9386422094F879FD222EBD" box="[694,750,1940,1964]" pageId="11" pageNumber="356" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A epigyne, ventral view; B same, dorsal view; C same, caudal view; D same, lateral view; E, abdomen, ventral view. Scale bars: 0.2 mm. Abbreviations:
<emphasis id="B918A246FF93864221D5F854FBE42EC1" box="[1015,1064,1977,2000]" italics="true" pageId="11" pageNumber="356">Be—</emphasis>
base of epigyne,
<emphasis id="B918A246FF93864226F4F855FAD72EDE" box="[1238,1307,1976,1999]" italics="true" pageId="11" pageNumber="356">DPS—</emphasis>
distal part of scape,
<emphasis id="B918A246FF93864222FEF830FEC32EE5" box="[220,271,2013,2036]" italics="true" pageId="11" pageNumber="356">Eg—</emphasis>
epigynal groove,
<emphasis id="B918A246FF938642239DF831FE272EE5" box="[447,491,2012,2036]" italics="true" pageId="11" pageNumber="356">Ll—</emphasis>
lateral lobe,
<emphasis id="B918A246FF938642204BF830FD7F2EE5" box="[617,691,2013,2036]" italics="true" pageId="11" pageNumber="356">PMP—</emphasis>
posterior median plate,
<emphasis id="B918A246FF9386422181F831FC282EE2" box="[931,996,2012,2035]" italics="true" pageId="11" pageNumber="356">PPS—</emphasis>
proximal part of scape,
<emphasis id="B918A246FF93864226F7F831FACE2EE2" box="[1237,1282,2012,2035]" italics="true" pageId="11" pageNumber="356">Sr—</emphasis>
stretcher.
</paragraph>
</caption>
<subSubSection id="C3762DDFFF94864522B5FF7BFD4A29C5" pageId="12" pageNumber="357" type="materials_examined">
<paragraph id="8BD37E54FF94864522B5FF7BFD4A29C5" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522B5FF7BFE8629A1" bold="true" box="[151,330,150,176]" pageId="12" pageNumber="357">Type material.</emphasis>
<materialsCitation id="3B047409FF9486452375FF7BFD4A29C5" collectingDate="2014-09-17" collectionCode="ZMUT" collectorName="H. Darvishnia" country="Iran" latitude="33.211002" location="Kabootarlan Cave" longLatPrecision="1" longitude="47.050915" municipality="Zarrin Abad" pageId="12" pageNumber="357" specimenCount="1" specimenCount-female="1" stateProvince="Ilam" typeStatus="holotype">
<typeStatus id="54D7C0F6FF9486452375FF7BFE7329A1" box="[343,447,150,176]" pageId="12" pageNumber="357" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6AB5DDFF94864523E9FF7AFE2C29A0" box="[459,480,151,177]" count="1" pageId="12" pageNumber="357" type="female">♀</specimenCount>
(
<collectionCode id="ED7DE691FF94864523D6FF7BFD8529A1" box="[500,585,150,176]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="12" pageNumber="357">ZMUT</collectionCode>
),
<collectingCountry id="F37B3EC4FF9486452040FF7BFD6729A1" box="[610,683,150,176]" name="Iran" pageId="12" pageNumber="357">IRAN</collectingCountry>
:
<emphasis id="B918A246FF9486452098FF7BFC9229A1" box="[698,862,150,176]" italics="true" pageId="12" pageNumber="357">
<collectingRegion id="49A8B0B6FF9486452098FF7BFC9229A1" box="[698,862,150,176]" country="Iran" name="Ilam" pageId="12" pageNumber="357">Ilam Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6BB7E42EFF9486452150FF7BFBCE29A1" box="[882,1026,150,176]" pageId="12" pageNumber="357">Zarrin Abad</collectingMunicipality>
,
<location id="8EB3288FFF9486452631FF7BFB2A29A1" LSID="urn:lsid:plazi:treatment:03C5CF42FF92864522B5FD36FECC2D49:8EB3288FFF9486452631FF7BFB2A29A1" box="[1043,1254,150,176]" country="Iran" latitude="33.211002" longLatPrecision="1" longitude="47.050915" municipality="Zarrin Abad" name="Kabootarlan Cave" pageId="12" pageNumber="357" stateProvince="Ilam">Kabootarlan Cave</location>
,
<geoCoordinate id="EE581893FF94864526D5FF7BFA5029A1" box="[1271,1436,150,176]" degrees="33" direction="north" minutes="12" orientation="latitude" pageId="12" pageNumber="357" precision="1" seconds="39.60" value="33.211002">33°12'39.60''N</geoCoordinate>
<geoCoordinate id="EE581893FF94864522B5FF57FEF029C5" box="[151,316,186,212]" degrees="47" direction="east" minutes="03" orientation="longitude" pageId="12" pageNumber="357" precision="1" seconds="03.30" value="47.050915">47°03'03.30''E</geoCoordinate>
,
<date id="FFD25894FF948645236AFF57FE0429C5" box="[328,456,186,212]" pageId="12" pageNumber="357" value="2014-09-17">
<collectingDate id="EF96A17CFF948645236AFF57FE0429C5" box="[328,456,186,212]" pageId="12" pageNumber="357" value="2014-09-17">17.IX.2014</collectingDate>
</date>
(
<collectorName id="26991B82FF94864523F5FF57FDB629C5" box="[471,634,186,212]" pageId="12" pageNumber="357">H. Darvishnia</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF94864522E5FF33FCE2280D" pageId="12" pageNumber="357" type="etymology">
<paragraph id="8BD37E54FF94864522E5FF33FCE2280D" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522E5FF33FE8229E9" bold="true" box="[199,334,222,248]" pageId="12" pageNumber="357">Etymology.</emphasis>
The specific epithet is a combination of the Greek words ‘mega’ (large) and ‘gunḗ’ (female), referring to the unusually large epigyne of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF94864522E5FECBFCAB2875" pageId="12" pageNumber="357" type="diagnosis">
<paragraph id="8BD37E54FF94864522E5FECBFCAB2875" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522E5FECBFE8E2851" bold="true" box="[199,322,294,320]" pageId="12" pageNumber="357">Diagnosis.</emphasis>
The new species differs from all of the congeners by the wide epigyne (i.e., 0.68 of the abdomen’s width) and very thin scape (thinner than lateral sides,
<emphasis id="B918A246FF94864520D3FEA6FCC42875" box="[753,776,331,356]" italics="true" pageId="12" pageNumber="357">vs</emphasis>
. wider).
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF94864522E5FE83FE492A91" pageId="12" pageNumber="357" type="description">
<paragraph id="8BD37E54FF94864522E5FE83FC912B71" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522E5FE83FE952899" bold="true" box="[199,345,366,392]" pageId="12" pageNumber="357">Description.</emphasis>
Female. Habitus as in
<figureCitation id="135762D1FF948645207CFE83FD242899" box="[606,744,366,392]" captionStart="FIGURE 10" captionStartId="11.[151,250,1940,1964]" captionTargetBox="[160,1425,191,1908]" captionTargetId="figure-18@11.[151,1436,181,1916]" captionTargetPageId="11" captionText="FIGURE 10. Female of Troglohyphantes megagynus sp. n. A epigyne, ventral view; B same, dorsal view; C same, caudal view; D same, lateral view; E, abdomen, ventral view. Scale bars: 0.2 mm. Abbreviations: Be—base of epigyne, DPS—distal part of scape, Eg—epigynal groove, Ll—lateral lobe, PMP—posterior median plate, PPS—proximal part of scape, Sr—stretcher." figureDoi="http://doi.org/10.5281/zenodo.10084558" httpUri="https://zenodo.org/record/10084558/files/figure.png" pageId="12" pageNumber="357">Fig. 10A–D</figureCitation>
. Total length 1.85. Carapace 0.95 long, 0.75 wide. Eye sizes: ALE 0.03, PME 0.03, PLE 0.02; AME strongly reduced, all eyes vestigial. Carapace, sternum, labium, maxillae and legs light yellowish brown, chelicerae slightly darker. Chelicerae with 4–5 large pro- and 5 small retromarginal teeth. Legs slightly lighter than carapace. Abdomen beige, without any pattern. Spinnerets uniformly light beige. Measurements of legs: I: 6.36 (1.75, 0.26, 1.75, 1.68, 0.92), II: 5.89 (1.63, 0.30, 1.60, 1.53, 0.83), III: 4.59 (1.34, 0.26, 1.16, 1.20, 0.63), IV: 5.74 (1.67, 0.28, 1.49, 1.57, 0.73). Numbers of dorsal spines on tibiae: 2222. Position of trichobothrium on Mt I: 0.27. Trichobothrium on Mt IV absent.
</paragraph>
<paragraph id="8BD37E54FF94864522E5FD87FE532A4D" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
Epigyne as in
<figureCitation id="135762D1FF948645234EFD87FE372B95" box="[364,507,618,644]" captionStart="FIGURE 10" captionStartId="11.[151,250,1940,1964]" captionTargetBox="[160,1425,191,1908]" captionTargetId="figure-18@11.[151,1436,181,1916]" captionTargetPageId="11" captionText="FIGURE 10. Female of Troglohyphantes megagynus sp. n. A epigyne, ventral view; B same, dorsal view; C same, caudal view; D same, lateral view; E, abdomen, ventral view. Scale bars: 0.2 mm. Abbreviations: Be—base of epigyne, DPS—distal part of scape, Eg—epigynal groove, Ll—lateral lobe, PMP—posterior median plate, PPS—proximal part of scape, Sr—stretcher." figureDoi="http://doi.org/10.5281/zenodo.10084558" httpUri="https://zenodo.org/record/10084558/files/figure.png" pageId="12" pageNumber="357">Figs 10A–D</figureCitation>
; very broad, ca. 0.68 of abdomen’s width and very long (including base,
<emphasis id="B918A246FF9486452767FD86FAAF2B95" box="[1349,1379,619,644]" italics="true" pageId="12" pageNumber="357">Be</emphasis>
): ca. 0.58; base of epigyne subtrapezoidal, with distinct inclining grooves (
<emphasis id="B918A246FF948645219EFD62FC172BB9" box="[956,987,655,680]" italics="true" pageId="12" pageNumber="357">Eg</emphasis>
), narrowing posteriorly; epigynal plate sub-hexagonal, anteriorly as wide as 0.5 as maximal width; scape short and thin, ca. 0.28 of epigynal plate’s width, thinner than lateral side of epigynal plate, with lateral lobes (
<emphasis id="B918A246FF9486452174FD3BFCA12BE1" box="[854,877,726,752]" italics="true" pageId="12" pageNumber="357">Ll</emphasis>
); proximal part of scape (
<emphasis id="B918A246FF94864526B9FD3AFB072BE0" box="[1179,1227,727,753]" italics="true" pageId="12" pageNumber="357">PPS</emphasis>
) wider than distal part (
<emphasis id="B918A246FF94864522F0FD16FEC92A04" box="[210,261,763,789]" italics="true" pageId="12" pageNumber="357">DPS</emphasis>
); epigynal grooves (
<emphasis id="B918A246FF94864523CEFD16FDC72A05" box="[492,523,763,788]" italics="true" pageId="12" pageNumber="357">Eg</emphasis>
) diverging in anterior part, then converging, and subparallel in posterior part (
<figureCitation id="135762D1FF948645274DFD17FF062A29" captionStart="FIGURE 11" captionStartId="12.[151,250,2009,2033]" captionTargetBox="[161,1436,1377,1985]" captionTargetId="figure-449@12.[151,1436,1367,1986]" captionTargetPageId="12" captionText="FIGURE 11. Habitus of Scotargus pilosus, male. A dorsal view; B prosoma, anterior view." figureDoi="http://doi.org/10.5281/zenodo.10084560" httpUri="https://zenodo.org/record/10084560/files/figure.png" pageId="12" pageNumber="357">Fig. 11A</figureCitation>
); posterior median plate (
<emphasis id="B918A246FF94864523CBFCF2FDEF2A29" box="[489,547,799,824]" italics="true" pageId="12" pageNumber="357">PMP</emphasis>
) 2.5 times longer than wide; stretcher (
<emphasis id="B918A246FF94864521F9FCF2FC382A28" box="[987,1012,799,825]" italics="true" pageId="12" pageNumber="357">Sr</emphasis>
) long, ca. 4 times longer than wide in lateral view (
<figureCitation id="135762D1FF9486452308FCAFFE5F2A4D" box="[298,403,834,860]" captionStart="FIGURE 10" captionStartId="11.[151,250,1940,1964]" captionTargetBox="[160,1425,191,1908]" captionTargetId="figure-18@11.[151,1436,181,1916]" captionTargetPageId="11" captionText="FIGURE 10. Female of Troglohyphantes megagynus sp. n. A epigyne, ventral view; B same, dorsal view; C same, caudal view; D same, lateral view; E, abdomen, ventral view. Scale bars: 0.2 mm. Abbreviations: Be—base of epigyne, DPS—distal part of scape, Eg—epigynal groove, Ll—lateral lobe, PMP—posterior median plate, PPS—proximal part of scape, Sr—stretcher." figureDoi="http://doi.org/10.5281/zenodo.10084558" httpUri="https://zenodo.org/record/10084558/files/figure.png" pageId="12" pageNumber="357">Fig. 10D</figureCitation>
).
</paragraph>
<paragraph id="8BD37E54FF94864522E5FC8BFE492A91" blockId="12.[151,1437,150,1112]" box="[199,389,870,896]" pageId="12" pageNumber="357">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF94864522E5FC67FAB62D01" pageId="12" pageNumber="357" type="discussion">
<paragraph id="8BD37E54FF94864522E5FC67FAB62D01" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522E5FC67FE882AB5" bold="true" box="[199,324,906,932]" pageId="12" pageNumber="357">Comment.</emphasis>
The new species is placed in
<taxonomicName id="4C6C05D7FF94864520BEFC67FC3F2AB5" authority="Joseph, 1882" authorityName="Joseph" authorityYear="1882" box="[668,1011,906,932]" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="B918A246FF94864520BEFC67FC9B2AB5" box="[668,855,906,932]" italics="true" pageId="12" pageNumber="357">Troglohyphantes</emphasis>
Joseph, 1882
</taxonomicName>
tentatively, as many members of this genus are known to inhabit caves, with some possessing vestigial or absent eyes. Additionally, all cave-dwelling
<taxonomicName id="4C6C05D7FF94864522B5FC3FFE652AFD" box="[151,425,978,1004]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522B5FC3FFE9E2AFD" box="[151,338,978,1004]" italics="true" pageId="12" pageNumber="357">Troglohyphantes</emphasis>
species
</taxonomicName>
exhibit long legs. However, it is highly probable that this species belongs to an undescribed genus. Due to the absence of male specimens, we are currently unable to determine its exact generic placement.
</paragraph>
</subSubSection>
<subSubSection id="C3762DDFFF94864522E5FBF7FECC2D49" pageId="12" pageNumber="357" type="distribution">
<paragraph id="8BD37E54FF94864522E5FBF7FECC2D49" blockId="12.[151,1437,150,1112]" pageId="12" pageNumber="357">
<emphasis id="B918A246FF94864522E5FBF7FEAD2D25" bold="true" box="[199,353,1050,1076]" pageId="12" pageNumber="357">Distribution.</emphasis>
Known only from the
<typeStatus id="54D7C0F6FF9486452042FBF6FD5C2D24" box="[608,656,1051,1077]" pageId="12" pageNumber="357">type</typeStatus>
locality in Kabootarlan Cave (
<figureCitation id="135762D1FF94864521CBFBF7FB9D2D24" box="[1001,1105,1050,1077]" captionStart="FIGURE 15" captionStartId="16.[151,250,1977,2001]" captionTargetBox="[158,1430,525,1953]" captionTargetId="figure-141@16.[158,1430,525,1953]" captionTargetPageId="16" captionText="FIGURE 15. Some of the collection localities of the material reported in this study.A Barreh Zard; B Chehel Sotoon; C Gakal; D Kabootarlan; E Khoffash; F Malek. Photos by Mohammad Javad Malek-Hosseini." figureDoi="http://doi.org/10.5281/zenodo.10084568" httpUri="https://zenodo.org/record/10084568/files/figure.png" pageId="12" pageNumber="357">Fig. 15D</figureCitation>
),
<collectingRegion id="49A8B0B6FF9486452647FBF7FACA2D25" box="[1125,1286,1050,1076]" country="Iran" name="Ilam" pageId="12" pageNumber="357">Ilam Province</collectingRegion>
, western
<collectingCountry id="F37B3EC4FF948645274DFBF6FA502D24" box="[1391,1436,1051,1077]" name="Iran" pageId="12" pageNumber="357">Iran</collectingCountry>
(
<figureCitation id="135762D1FF94864522BDFBD3FF3F2D49" box="[159,243,1086,1112]" captionStart="FIGURE 17" captionStartId="18.[151,250,1232,1256]" captionTargetBox="[275,1311,345,1209]" captionTargetId="figure-69@18.[275,1311,345,1209]" captionTargetPageId="18" captionText="FIGURE 17. Partial map of Iran, showing the collection localities of the material reported in this study. 1 Ali Sheikh Cave; 2 Barreh Zard Cave; 3 Chehel Sotoon Cave; 4 Darreh-ye Ashkaft Cave; 5 Dengezlu Cave; 6 Gakal Cave; 7 Gelim Goosh Cave; 8 Hamzeh Zendan Cave; 9 Jabiglu Cave; 10 Kabootarlan Cave; 11 Kahman Cave; 12 Kerend Cave; 13 Khoffash Cave, Mar Kelga Cave; 14 Malek Cave, Markhareil Cave; 15 Mir Melas Cave; 16 Mooly Cave; 17 Pir Morad Cave; 18 Raad-e Gharbi Cave; 19 Tang-e Kaboutari; 20 Tang-e-Lor Cave; 21 Tang-e Zendan Cave; 22 Tashan Cave; 23 Torang Cave; 24 Var-e-Zamin Cave; 25 Yaran Cave." figureDoi="http://doi.org/10.5281/zenodo.10084572" httpUri="https://zenodo.org/record/10084572/files/figure.png" pageId="12" pageNumber="357">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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