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<document id="5BC8AF0BD1C975DA10B526A97F2296E0" ID-CLB-Dataset="25149" ID-DOI="10.11646/zootaxa.4688.4.5" ID-GBIF-Dataset="c704c5d3-8ef6-48b6-a346-efe5b5e1c032" ID-ISSN="1175-5326" ID-Zenodo-Dep="3517710" ID-ZooBank="F412A472-E501-494B-9BF9-CAF253E9BD92" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571896581567" checkinUser="plazi" docAuthor="Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo &amp; Szwedo, Jacek" docDate="2019" docId="03C5112FFFE9FFAFFF02FA82FAEC51D0" docLanguage="en" docName="zootaxa.4688.4.5.pdf" docOrigin="Zootaxa 4688 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Culicoides ellenbergeri Szadziewski, sp. nov." docType="treatment" docUuid="C343228E-064A-4344-A63B-C748F54A468D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="541" masterDocId="FFFC6957FFECFFA9FF95FFC5FFA05106" masterDocTitle="Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous" masterLastPageNumber="548" masterPageNumber="535" pageNumber="540" updateTime="1698760925119" updateUser="ExternalLinkService">
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<mods:title id="AD08AAA14BEF10375539DC41674200F2">Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous</mods:title>
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<mods:namePart id="1ED4A0850E1F108258B37BDE46CF50F6">Szadziewski, Ryszard</mods:namePart>
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<mods:namePart id="50E9F1CD290F42D3511237D8FC5B18F9">Dominiak, Patrycja</mods:namePart>
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<mods:namePart id="3E00099E799D7DEFBC1BD8201E88F929">Sontag, Elżbieta</mods:namePart>
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<mods:namePart id="D965BDAF0FC4BBE89DC422B1B110BA34">Krzemiński, Wiesław</mods:namePart>
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<mods:namePart id="5A0EA46700431B7E22EC40C62711C2C4">Wang, Bo</mods:namePart>
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<treatment id="03C5112FFFE9FFAFFF02FA82FAEC51D0" ID-DOI="http://doi.org/10.5281/zenodo.5698032" ID-GBIF-Taxon="159834508" ID-Zenodo-Dep="5698032" LSID="urn:lsid:zoobank.org:act:C343228E-064A-4344-A63B-C748F54A468D" httpUri="http://treatment.plazi.org/id/03C5112FFFE9FFAFFF02FA82FAEC51D0" lastPageId="6" lastPageNumber="541" pageId="5" pageNumber="540">
<subSubSection id="C376F3B2FFE9FFACFF02FA82FCFD5464" box="[151,861,1351,1378]" pageId="5" pageNumber="540" type="nomenclature">
<paragraph id="8BD3A039FFE9FFACFF02FA82FCFD5464" blockId="5.[151,960,1351,1443]" box="[151,861,1351,1378]" pageId="5" pageNumber="540">
<heading id="D09B1755FFE9FFACFF02FA82FCFD5464" bold="true" box="[151,861,1351,1378]" fontSize="11" level="1" pageId="5" pageNumber="540" reason="1">
<emphasis id="B9187C2BFFE9FFACFF02FA82FCFD5464" bold="true" box="[151,861,1351,1378]" pageId="5" pageNumber="540">
<taxonomicName id="4C6CDBBAFFE9FFACFF02FA82FDEE5467" authority="Szadziewski" authorityName="Szadziewski" box="[151,590,1351,1378]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="540" phylum="Arthropoda" rank="species" species="ellenbergeri">
<emphasis id="B9187C2BFFE9FFACFF02FA82FE0F5464" bold="true" box="[151,431,1351,1378]" italics="true" pageId="5" pageNumber="540">Culicoides ellenbergeri</emphasis>
Szadziewski
</taxonomicName>
&amp; Dominiak,
<taxonomicNameLabel id="A22BC150FFE9FFACFD6AFA8DFCFD5464" box="[767,861,1352,1378]" pageId="5" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C376F3B2FFE9FFACFF02FAAEFC6054A5" pageId="5" pageNumber="540" type="description">
<paragraph id="8BD3A039FFE9FFACFF02FAAEFEB15480" blockId="5.[151,960,1351,1443]" box="[151,273,1387,1414]" pageId="5" pageNumber="540">
<figureCitation id="1357BCBCFFE9FFACFF02FAAEFF7A5480" box="[151,218,1387,1414]" captionStart="FIGURE 5" captionStartId="6.[151,250,1226,1251]" captionTargetBox="[151,1436,281,1198]" captionTargetId="figure@6.[151,1436,275,1198]" captionTargetPageId="6" captionText="FIGURE 5. Fossil Culicoides (Groganomyia) ellenbergeri Szadziewski &amp; Dominiak, sp. nov., female holotype. A—total habitus, B—palpus, C—flagellum." figureDoi="http://doi.org/10.5281/zenodo.3517720" httpUri="https://zenodo.org/record/3517720/files/figure.png" pageId="5" pageNumber="540">Figs 5</figureCitation>
AC
</paragraph>
<paragraph id="8BD3A039FFE9FFACFF02FA4DFC6054A5" blockId="5.[151,960,1351,1443]" box="[151,960,1416,1443]" pageId="5" pageNumber="540">
<uri id="FFFDAC3BFFE9FFACFF02FA4DFC6054A5" box="[151,960,1416,1443]" pageId="5" pageNumber="540">
urn:lsid:zoobank.org:act:
<uuid id="FFCA9AECFFE9FFACFE30FA4DFC6054A5" box="[421,960,1416,1443]" pageId="5" pageNumber="540">C343228E-064A-4344-A63B-C748F54A468D</uuid>
</uri>
</paragraph>
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<subSubSection id="C376F3B2FFE9FFACFF02FA15FAF65751" pageId="5" pageNumber="540" type="diagnosis">
<paragraph id="8BD3A039FFE9FFACFF02FA15FAF65751" blockId="5.[151,1437,1488,2019]" pageId="5" pageNumber="540">
<emphasis id="B9187C2BFFE9FFACFF02FA15FEB254ED" bold="true" box="[151,274,1488,1515]" pageId="5" pageNumber="540">Diagnosis.</emphasis>
This new species is distinguished from other Cretaceous
<taxonomicName id="4C6CDBBAFFE9FFACFC3DFA14FB8054ED" box="[936,1056,1489,1515]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="B9187C2BFFE9FFACFC3DFA14FB8054ED" box="[936,1056,1489,1515]" italics="true" pageId="5" pageNumber="540">Culicoides</emphasis>
</taxonomicName>
by the following combination of female characters: antennal flagellum with flagellomeres 28 spherical, much shorter than the more elongate 913, only flagellomere 1 with 7 sensilla coeloconica; distal portion of wing membrane with macrotrichia; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla; proboscis short. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C376F3B2FFE9FFACFF52F9A5FE28569D" pageId="5" pageNumber="540" type="description">
<paragraph id="8BD3A039FFE9FFACFF52F9A5FE905671" blockId="5.[151,1437,1488,2019]" pageId="5" pageNumber="540">
<emphasis id="B9187C2BFFE9FFACFF52F9A5FE18577D" bold="true" box="[199,440,1632,1659]" pageId="5" pageNumber="540">Description. Female</emphasis>
(
<figureCitation id="1357BCBCFFE9FFACFE5DF9A4FD84577D" box="[456,548,1632,1659]" captionStart="FIGURE 5" captionStartId="6.[151,250,1226,1251]" captionTargetBox="[151,1436,281,1198]" captionTargetId="figure@6.[151,1436,275,1198]" captionTargetPageId="6" captionText="FIGURE 5. Fossil Culicoides (Groganomyia) ellenbergeri Szadziewski &amp; Dominiak, sp. nov., female holotype. A—total habitus, B—palpus, C—flagellum." figureDoi="http://doi.org/10.5281/zenodo.3517720" httpUri="https://zenodo.org/record/3517720/files/figure.png" pageId="5" pageNumber="540">Fig. 5A</figureCitation>
). Poorly preserved, distorted, abdomen missing. Eyes bare, separation not visible. Antennal flagellum (
<figureCitation id="1357BCBCFFE9FFACFE22F940FDAD5799" box="[439,525,1668,1695]" captionStart="FIGURE 5" captionStartId="6.[151,250,1226,1251]" captionTargetBox="[151,1436,281,1198]" captionTargetId="figure@6.[151,1436,275,1198]" captionTargetPageId="6" captionText="FIGURE 5. Fossil Culicoides (Groganomyia) ellenbergeri Szadziewski &amp; Dominiak, sp. nov., female holotype. A—total habitus, B—palpus, C—flagellum." figureDoi="http://doi.org/10.5281/zenodo.3517720" httpUri="https://zenodo.org/record/3517720/files/figure.png" pageId="5" pageNumber="540">Fig. 5C</figureCitation>
) length
<quantity id="4C940DDCFFE9FFACFDFFF940FD715799" box="[618,721,1669,1695]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="540" unit="mm" value="0.32">0.32 mm</quantity>
; flagellomere 1 with 7 sensilla coeloconica, flagellomeres 213 without sensilla coeloconica, but with distinct sensilla basiconica; flagellomeres 28 more or less spherical, much shorter than the more elongate 913; antennal ratio 1.13. Palpus (
<figureCitation id="1357BCBCFFE9FFACFCEAF908FC7657E1" box="[895,982,1740,1767]" captionStart="FIGURE 5" captionStartId="6.[151,250,1226,1251]" captionTargetBox="[151,1436,281,1198]" captionTargetId="figure@6.[151,1436,275,1198]" captionTargetPageId="6" captionText="FIGURE 5. Fossil Culicoides (Groganomyia) ellenbergeri Szadziewski &amp; Dominiak, sp. nov., female holotype. A—total habitus, B—palpus, C—flagellum." figureDoi="http://doi.org/10.5281/zenodo.3517720" httpUri="https://zenodo.org/record/3517720/files/figure.png" pageId="5" pageNumber="540">Fig. 5B</figureCitation>
) 5-segmented; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla on ventral surface. Wing (
<figureCitation id="1357BCBCFFE9FFACFB26F934FAAC560D" box="[1203,1292,1776,1803]" captionStart="FIGURE 5" captionStartId="6.[151,250,1226,1251]" captionTargetBox="[151,1436,281,1198]" captionTargetId="figure@6.[151,1436,275,1198]" captionTargetPageId="6" captionText="FIGURE 5. Fossil Culicoides (Groganomyia) ellenbergeri Szadziewski &amp; Dominiak, sp. nov., female holotype. A—total habitus, B—palpus, C—flagellum." figureDoi="http://doi.org/10.5281/zenodo.3517720" httpUri="https://zenodo.org/record/3517720/files/figure.png" pageId="5" pageNumber="540">Fig. 5A</figureCitation>
) length
<quantity id="4C940DDCFFE9FFACFAFEF934FF665629" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="540" unit="mm" value="0.61">0.61 mm</quantity>
; costal ratio 0.69; distal portion of wing with macrotrichia in cells r
<subScript id="17E8A27CFFE9FFACFC57F8E3FC6A5632" attach="left" box="[962,970,1830,1844]" fontSize="6" pageId="5" pageNumber="540">3</subScript>
, m
<subScript id="17E8A27CFFE9FFACFC7AF8E3FC575632" attach="left" box="[1007,1015,1830,1844]" fontSize="6" pageId="5" pageNumber="540">1</subScript>
and m
<subScript id="17E8A27CFFE9FFACFBD0F8E3FBED5632" attach="left" box="[1093,1101,1830,1844]" fontSize="6" pageId="5" pageNumber="540">2</subScript>
; basal radial and cubital cells without macrotrichia. Scutellum with about 8 long marginal setae. Tarsomere 1 of hind leg slender; TR
<subScript id="17E8A27CFFE9FFACFA8EF88FFA8D565E" attach="left" box="[1307,1325,1866,1880]" fontSize="6" pageId="5" pageNumber="540">(1)</subScript>
2.1, TR
<subScript id="17E8A27CFFE9FFACFA1CF88FFA3C565E" attach="left" box="[1417,1436,1866,1880]" fontSize="6" pageId="5" pageNumber="540">(2)</subScript>
2.0, TR
<subScript id="17E8A27CFFE9FFACFF7EF8ABFF5E567A" attach="left" box="[235,254,1902,1916]" fontSize="6" pageId="5" pageNumber="540">(3)</subScript>
1.8.
</paragraph>
<paragraph id="8BD3A039FFE9FFACFF52F844FE28569D" blockId="5.[151,1437,1488,2019]" box="[199,392,1921,1947]" pageId="5" pageNumber="540">
<emphasis id="B9187C2BFFE9FFACFF52F844FEAB569D" bold="true" box="[199,267,1921,1947]" pageId="5" pageNumber="540">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C376F3B2FFE9FFACFF52F861FADF56E5" pageId="5" pageNumber="540" type="materials_examined">
<paragraph id="8BD3A039FFE9FFACFF52F861FADF56E5" blockId="5.[151,1437,1488,2019]" pageId="5" pageNumber="540">
<emphasis id="B9187C2BFFE9FFACFF52F861FE1156B9" bold="true" box="[199,433,1956,1983]" pageId="5" pageNumber="540">Material examined.</emphasis>
<materialsCitation id="3B04AA64FFE9FFACFE2DF860FADF56E5" ID-GBIF-Occurrence="2429183020" country="Myanmar" county="University of Gdansk." location="Burmese" municipality="Museum of Amber Inclusions" pageId="5" pageNumber="540" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D71E9BFFE9FFACFE2DF860FD8056B9" box="[440,544,1957,1983]" pageId="5" pageNumber="540" type="holotype">Holotype</typeStatus>
female, MAI 5666.
<location id="8EB3F6E2FFE9FFACFC93F860FCCA56B9" LSID="urn:lsid:plazi:treatment:03C5112FFFE9FFAFFF02FA82FAEC51D0:8EB3F6E2FFE9FFACFC93F860FCCA56B9" box="[774,874,1957,1983]" country="Myanmar" county="University of Gdansk." municipality="Museum of Amber Inclusions" name="Burmese" pageId="5" pageNumber="540">Burmese</location>
amber,
<location id="8EB3F6E2FFE9FFACFC51F860FB2156B9" LSID="urn:lsid:plazi:treatment:03C5112FFFE9FFAFFF02FA82FAEC51D0:8EB3F6E2FFE9FFACFC51F860FB2156B9" box="[964,1153,1957,1983]" country="Myanmar" county="University of Gdansk." municipality="Museum of Amber Inclusions" name="Hukawng Valley" pageId="5" pageNumber="540">Hukawng Valley</location>
,
<collectingCountry id="F37BE0A9FFE9FFACFB19F860FB5D56B9" box="[1164,1277,1957,1983]" name="Myanmar" pageId="5" pageNumber="540">Myanmar</collectingCountry>
; purchased by
<location id="8EB3F6E2FFE9FFACFF02F80CFE2456E5" LSID="urn:lsid:plazi:treatment:03C5112FFFE9FFAFFF02FA82FAEC51D0:8EB3F6E2FFE9FFACFF02F80CFE2456E5" box="[151,388,1993,2019]" country="Myanmar" county="University of Gdansk." municipality="Museum of Amber Inclusions" name="Sieghard Ellenberger" pageId="5" pageNumber="540">Sieghard Ellenberger</location>
from
<location id="8EB3F6E2FFE9FFACFE5DF80CFDB556E5" LSID="urn:lsid:plazi:treatment:03C5112FFFE9FFAFFF02FA82FAEC51D0:8EB3F6E2FFE9FFACFE5DF80CFDB556E5" box="[456,533,1993,2019]" country="Myanmar" county="University of Gdansk." municipality="Museum of Amber Inclusions" name="Kassel" pageId="5" pageNumber="540">Kassel</location>
,
<collectingCountry id="F37BE0A9FFE9FFACFDB4F80CFD2B56E5" box="[545,651,1993,2019]" name="Germany" pageId="5" pageNumber="540">Germany</collectingCountry>
.
<location id="8EB3F6E2FFE9FFACFD00F80CFC2D56E5" LSID="urn:lsid:plazi:treatment:03C5112FFFE9FFAFFF02FA82FAEC51D0:8EB3F6E2FFE9FFACFD00F80CFC2D56E5" box="[661,909,1993,2019]" country="Myanmar" county="University of Gdansk." municipality="Museum of Amber Inclusions" name="Deposited in Museum" pageId="5" pageNumber="540">
Deposited in
<collectingMunicipality id="6BB73A43FFE9FFACFCBEF80CFC2D56E5" box="[811,909,1993,2019]" pageId="5" pageNumber="540">Museum</collectingMunicipality>
</location>
of Amber Inclusions,
<collectingCounty id="62B2D8B5FFE9FFACFB13F80CFADF56E5" box="[1158,1407,1993,2019]" pageId="5" pageNumber="540">University of Gdańsk.</collectingCounty>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376F3B2FFEAFFAFFF52FF5CFAEC51D0" pageId="6" pageNumber="541" type="etymology">
<paragraph id="8BD3A039FFEAFFAFFF52FF5CFAEC51D0" blockId="6.[151,1436,152,215]" pageId="6" pageNumber="541">
<emphasis id="B9187C2BFFEAFFAFFF52FF5CFEEE51B5" bold="true" box="[199,334,153,179]" pageId="6" pageNumber="541">Etymology.</emphasis>
This new species is dedicated to Sieghard Ellenberger of Kassel,
<collectingCountry id="F37BE0A9FFEAFFAFFBD4FF5CFB0A51B5" box="[1089,1194,153,179]" name="Germany" pageId="6" pageNumber="541">Germany</collectingCountry>
, a generous collector who purchased this biting midge and donated it to the Museum of Amber Inclusions, University of Gdańsk.
</paragraph>
</subSubSection>
</treatment>
</document>