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<document id="DE5EBC87CB3C2DC3261ECEF5E9D9830F" ID-DOI="10.35463/j.apr.2021.02.01" ID-ISSN="1842-371x" ID-Zenodo-Dep="12583322" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719585668405" checkinUser="felipe" docAuthor="SCHLAGINTWEIT, FELIX &amp; RASHIDI, KOOROSH" docDate="2021" docId="03B887C22B4B4E61FF0BD7CEB0E9DB74" docLanguage="en" docName="ActaPalaeontolRom.17.2.3-13.pdf" docOrigin="Acta Palaeontologica Romaniae 17 (2)" docSource="http://dx.doi.org/10.35463/j.apr.2021.02.01" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Dictyoconella complanata Henson 1948" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FF81FFBA2B4F4E66FF9CD04CB245D838" masterDocTitle="Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)" masterLastPageNumber="13" masterPageNumber="3" pageNumber="7" updateTime="1719587089633" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D0CD3C3838BFC98415A165DFFD6878BF">Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)</mods:title>
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<subSubSection id="C30B655F2B4B4E62FF0BD7CEB009DFA2" box="[151,588,1922,1946]" pageId="4" pageNumber="7" type="nomenclature">
<paragraph id="8BAE36D42B4B4E62FF0BD7CEB009DFA2" blockId="4.[151,588,1922,1946]" box="[151,588,1922,1946]" pageId="4" pageNumber="7">
<heading id="D0E681B82B4B4E62FF0BD7CEB009DFA2" box="[151,588,1922,1946]" fontSize="10" level="2" pageId="4" pageNumber="7" reason="5">
<taxonomicName id="4C114D572B4B4E62FF0BD7CEB009DFA2" ID-CoL="82CTX" authority="Henson, 1948" authorityName="Henson" authorityYear="1948" box="[151,588,1922,1946]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B4B4E62FF0BD7CEB3F4DFA2" box="[151,433,1922,1946]" italics="true" pageId="4" pageNumber="7">Dictyoconella complanata</emphasis>
<bibRefCitation id="EF804B252B4B4E62FE24D7CEB009DFA2" author="Henson, F. R. S." box="[440,588,1922,1946]" pageId="4" pageNumber="7" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B655F2B4B4E61FF0BD7EFB370D8EF" lastPageId="7" lastPageNumber="10" pageId="4" pageNumber="7" type="description">
<paragraph id="8BAE36D42B4B4E62FF0BD7EFB323DF83" blockId="4.[151,771,1955,2045]" box="[151,358,1955,1979]" pageId="4" pageNumber="7">
<figureCitation id="132A2A512B4B4E62FF0BD7EFB29ADF83" box="[151,223,1955,1979]" captionStart="Fig" captionStartId="1.[240,275,1053,1074]" captionTargetBox="[251,1385,154,1015]" captionTargetId="figure-864@1.[251,1385,154,1015]" captionTargetPageId="1" captionText="Fig. 2 Microfacies with Dictyoconella complanata Henson (a-b) and Gusicella minima (Henson) gen. et comb. nov. (c- d) from the Maastrichtian Tarbur Formation of the Naghan (a-b) and Mandegan sections (c-d), SW Iran (Zagros zone). Abbreviations: D = Dictyoconella, Di = Dicyclina, G = Gyroconulina, Gu = Gusicella, L = Loftusia, O = Omphalocyclus, P = Pseudonummoloculina, Ps = Pseudocymopolia, T = Tarburina. Thin sections: 2Ng 81 (a), 2Ng 81-4 (b), Rt 109 (c), Rt 105 (d)." figureDoi="http://doi.org/10.5281/zenodo.12583326" httpUri="https://zenodo.org/record/12583326/files/figure.png" pageId="4" pageNumber="7">Figs. 2</figureCitation>
a-b pars, 3-4
</paragraph>
<paragraph id="8BAE36D42B4B4E62FF0BD788B011DFC5" blockId="4.[151,771,1955,2045]" pageId="4" pageNumber="7">
1948
<taxonomicName id="4C114D572B4B4E62FF49D788B3B4DFE4" authorityName="SCHLAGINTWEIT &amp; RASHIDI" authorityYear="2021" box="[213,497,1988,2012]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="complanata" status="n. gen., n. sp.">
<emphasis id="B965EAC62B4B4E62FF49D788B3B4DFE4" box="[213,497,1988,2012]" italics="true" pageId="4" pageNumber="7">Dictyoconella complanata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657BD2B4B4E62FE65D789B0C8DFE4" box="[505,653,1989,2013]" pageId="4" pageNumber="7" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Henson, p. 25, plate 6, figs. 2-3, 16, pl. 10, fig. 14.
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD598B18EDE34" blockId="4.[816,1437,1492,1747]" pageId="4" pageNumber="7">
1986
<taxonomicName id="4C114D572B4B4E62FCF2D598B6A6DDD3" authority="Henson" authorityName="Henson" authorityYear="1948" box="[878,1251,1492,1516]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B4B4E62FCF2D598B6CFDDD4" box="[878,1162,1492,1516]" italics="true" pageId="4" pageNumber="7">Dictyoconella complanata</emphasis>
Henson
</taxonomicName>
Köylüoğlu, pl. 86, fig. 1 pars.
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD65AB647DE77" blockId="4.[816,1437,1492,1747]" pageId="4" pageNumber="7">
1998
<taxonomicName id="4C114D572B4B4E62FCEED65AB6ABDE16" authority="Henson" authorityName="Henson" authorityYear="1948" box="[882,1262,1558,1582]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B4B4E62FCEED65AB6D4DE16" box="[882,1169,1558,1582]" italics="true" pageId="4" pageNumber="7">Dictyoconella complanata</emphasis>
Henson
</taxonomicName>
Whittaker et al., pl. 47, figs. 4-6.
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD614B63DDEA9" blockId="4.[816,1437,1492,1747]" pageId="4" pageNumber="7">
2016
<taxonomicName id="4C114D572B4B4E62FCF3D615B77CDE48" authority="Schlagintweit et al." authorityName="Schlagintweit" box="[879,1337,1624,1648]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconus" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="bakhtiari">
<emphasis id="B965EAC62B4B4E62FCF3D615B61ADE48" box="[879,1119,1624,1648]" italics="true" pageId="4" pageNumber="7">Dictyoconus bakhtiari</emphasis>
Schlagintweit et al.
</taxonomicName>
Schlagintweit et al., fig.
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, fig.
<specimenCount id="9D17FD5D2B4B4E62FBD2D635B63DDEA9" box="[1102,1144,1657,1681]" count="10" pageId="4" pageNumber="7" type="generic">10f.</specimenCount>
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD6D6B197DEEB" blockId="4.[816,1437,1492,1747]" pageId="4" pageNumber="7">
2020
<taxonomicName id="4C114D572B4B4E62FCEDD6D6B6AEDE8A" authority="Henson" authorityName="Henson" authorityYear="1948" box="[881,1259,1690,1714]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="7" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B4B4E62FCEDD6D6B6CADE8A" box="[881,1167,1690,1714]" italics="true" pageId="4" pageNumber="7">Dictyoconella complanata</emphasis>
Henson
</taxonomicName>
Schlagintweit, p. 75, fig. 4d-f.
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD6BBB69EDF37" blockId="4.[816,1243,1783,1808]" box="[816,1243,1783,1808]" pageId="4" pageNumber="7">
<emphasis id="B965EAC62B4B4E62FCACD6BBB1F8DF37" bold="true" box="[816,957,1783,1807]" pageId="4" pageNumber="7">Description:</emphasis>
See the genus description.
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD77FB1F8DF73" blockId="4.[816,1436,1843,2065]" box="[816,957,1843,1867]" pageId="4" pageNumber="7">
<emphasis id="B965EAC62B4B4E62FCACD77FB1F8DF73" bold="true" box="[816,957,1843,1867]" pageId="4" pageNumber="7">Dimensions:</emphasis>
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD718B1AEDFB5" blockId="4.[816,1436,1843,2065]" pageId="4" pageNumber="7">
Test diameter: up to
<quantity id="4CE99B312B4B4E62FB8CD718B626DF53" box="[1040,1123,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="7" unit="mm" value="4.8">4.8 mm</quantity>
(
<quantity id="4CE99B312B4B4E62FBEFD718B68DDF53" box="[1139,1224,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="7" unit="mm" value="3.3">3.3 mm</quantity>
,
<typeStatus id="54AA88762B4B4E62FB4FD718B774DF54" box="[1235,1329,1876,1900]" pageId="4" pageNumber="7" type="holotype">holotype</typeStatus>
specimen of
<bibRefCitation id="EF804B252B4B4E62FCD1D739B1A6DFB5" author="Henson, F. R. S." box="[845,995,1909,1933]" pageId="4" pageNumber="7" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
)
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD7DAB1E4DFF7" blockId="4.[816,1436,1843,2065]" pageId="4" pageNumber="7">
Test height:
<quantity id="4CE99B312B4B4E62FC2BD7DAB64CDF95" box="[951,1033,1942,1966]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="7" unit="mm" value="3.6">3.6 mm</quantity>
(
<quantity id="4CE99B312B4B4E62FB84D7DAB62BDF95" box="[1048,1134,1942,1966]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="7" unit="mm" value="2.9">2.9 mm</quantity>
,
<typeStatus id="54AA88762B4B4E62FBE5D7DAB693DF96" box="[1145,1238,1942,1966]" pageId="4" pageNumber="7" type="holotype">holotype</typeStatus>
specimen of
<bibRefCitation id="EF804B252B4B4E62FAFBD7DAB1DFDFF7" author="Henson, F. R. S." pageId="4" pageNumber="7" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
)
</paragraph>
<paragraph id="8BAE36D42B4B4E62FCACD794B18BD029" blockId="4.[816,1436,1843,2065]" pageId="4" pageNumber="7">
Thickness: mostly between 0.8 and
<quantity id="4CE99B312B4B4E62FB5AD794B758DFC8" box="[1222,1309,2008,2032]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="7" unit="mm" value="0.9">0.9 mm</quantity>
(~
<quantity id="4CE99B312B4B4E62FAA1D794B7DDDFC8" box="[1341,1432,2008,2032]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="7" unit="mm" value="0.8">0.8 mm</quantity>
,
<bibRefCitation id="EF804B252B4B4E62FCACD7B6B183D029" author="Henson, F. R. S." box="[816,966,2041,2065]" pageId="4" pageNumber="7" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
)
</paragraph>
<caption id="DF6E665C2B4A4E63FF09D6EDB10FDF51" ID-DOI="http://doi.org/10.5281/zenodo.12583336" ID-Zenodo-Dep="12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="5" pageNumber="8" startId="5.[149,184,1697,1718]" targetBox="[149,1438,157,1655]" targetPageId="5" targetType="figure">
<paragraph id="8BAE36D42B4A4E63FF09D6EDB10FDF51" blockId="5.[149,1439,1696,1897]" pageId="5" pageNumber="8">
<emphasis id="B965EAC62B4A4E63FF09D6EDB295DE8E" bold="true" box="[149,208,1697,1718]" pageId="5" pageNumber="8">Fig. 7</emphasis>
<taxonomicName id="4C114D572B4A4E63FF4BD6ECB3A7DE8E" authority="(Henson)" authorityName="SCHLAGINTWEIT &amp; RASHIDI" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT &amp; RASHIDI" baseAuthorityYear="1948" box="[215,482,1696,1718]" class="Globothalamea" family="Orbitolinidae" genus="Gusicella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="8" phylum="Foraminifera" rank="species" species="minima" status="gen. et comb. nov.">
<emphasis id="B965EAC62B4A4E63FF4BD6ECB3C7DE8E" box="[215,386,1696,1718]" italics="true" pageId="5" pageNumber="8">Gusicella minima</emphasis>
(Henson)
</taxonomicName>
<taxonomicNameLabel id="A25657BD2B4A4E63FE75D6EDB0DBDE8E" box="[489,670,1696,1718]" pageId="5" pageNumber="8" rank="species">gen. et comb. nov.</taxonomicNameLabel>
from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section:
<emphasis id="B965EAC62B4A4E63FF54D6F2B2ADDEEB" bold="true" box="[200,232,1726,1747]" pageId="5" pageNumber="8">b, l</emphasis>
; Naghan section:
<emphasis id="B965EAC62B4A4E63FE0BD6F2B396DEEB" bold="true" box="[407,467,1726,1747]" pageId="5" pageNumber="8">a, ck</emphasis>
).
<emphasis id="B965EAC62B4A4E63FE7BD6F2B3B6DEEB" bold="true" box="[487,499,1726,1747]" pageId="5" pageNumber="8">a</emphasis>
Oblique transverse section.
<emphasis id="B965EAC62B4A4E63FC99D6F2B157DEEB" bold="true" box="[773,786,1726,1747]" pageId="5" pageNumber="8">b</emphasis>
Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits.
<emphasis id="B965EAC62B4A4E63FC3CD690B183DEC9" bold="true" box="[928,966,1756,1777]" pageId="5" pageNumber="8">c, k</emphasis>
Subaxial sections.
<emphasis id="B965EAC62B4A4E63FB1BD690B6F7DEC9" bold="true" box="[1159,1202,1756,1777]" pageId="5" pageNumber="8">dg,</emphasis>
Axial sections showing initial spire.
<emphasis id="B965EAC62B4A4E63FE93D6B6B36ADF37" bold="true" box="[271,303,1786,1807]" pageId="5" pageNumber="8">hi</emphasis>
Details (from
<emphasis id="B965EAC62B4A4E63FE21D6B6B38DDF37" bold="true" box="[445,456,1786,1807]" pageId="5" pageNumber="8">e</emphasis>
and
<emphasis id="B965EAC62B4A4E63FE6BD6B6B046DF37" bold="true" box="[503,515,1786,1807]" pageId="5" pageNumber="8">g</emphasis>
) of the initial spire.
<emphasis id="B965EAC62B4A4E63FD4CD6B6B09DDF37" bold="true" box="[720,728,1786,1807]" pageId="5" pageNumber="8">j</emphasis>
Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (
<emphasis id="B965EAC62B4A4E63FC6BD77AB65BDF73" bold="true" box="[1015,1054,1846,1867]" pageId="5" pageNumber="8">a, k</emphasis>
), Rt 100 (
<emphasis id="B965EAC62B4A4E63FB1AD77AB6D6DF73" bold="true" box="[1158,1171,1846,1867]" pageId="5" pageNumber="8">b</emphasis>
), Ng 180-1 (
<emphasis id="B965EAC62B4A4E63FA8AD77AB764DF73" bold="true" box="[1302,1313,1846,1867]" pageId="5" pageNumber="8">c</emphasis>
), 2Ng 179-1 (
<emphasis id="B965EAC62B4A4E63FF01D718B2EFDF51" bold="true" box="[157,170,1876,1897]" pageId="5" pageNumber="8">d</emphasis>
), 2Ng 168 (
<emphasis id="B965EAC62B4A4E63FEBCD718B301DF51" bold="true" box="[288,324,1876,1897]" pageId="5" pageNumber="8">e, h</emphasis>
), 2Ng 175 (
<emphasis id="B965EAC62B4A4E63FE25D718B384DF51" bold="true" box="[441,449,1876,1897]" pageId="5" pageNumber="8">f</emphasis>
), 2Ng 174 (
<emphasis id="B965EAC62B4A4E63FDABD718B013DF51" bold="true" box="[567,598,1876,1897]" pageId="5" pageNumber="8">g, i</emphasis>
), 2Ng 177 (
<emphasis id="B965EAC62B4A4E63FD57D718B096DF51" bold="true" box="[715,723,1876,1897]" pageId="5" pageNumber="8">j</emphasis>
), Rt 113 (
<emphasis id="B965EAC62B4A4E63FCAAD718B178DF51" bold="true" box="[822,829,1876,1897]" pageId="5" pageNumber="8">l</emphasis>
).
</paragraph>
</caption>
<caption id="DF6E665C2B494E60FF52D6C6B6A3DF93" ID-DOI="http://doi.org/10.5281/zenodo.12583338" ID-Zenodo-Dep="12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="6" pageNumber="9" startId="6.[206,241,1674,1695]" targetBox="[219,1374,159,1632]" targetPageId="6" targetType="figure">
<paragraph id="8BAE36D42B494E60FF52D6C6B6A3DF93" blockId="6.[205,1382,1673,1963]" pageId="6" pageNumber="9">
<emphasis id="B965EAC62B494E60FF52D6C6B34DDEA7" bold="true" box="[206,264,1674,1695]" pageId="6" pageNumber="9">Fig. 8</emphasis>
<taxonomicName id="4C114D572B494E60FE93D6C5B05FDEA7" authority="(Henson)" authorityName="SCHLAGINTWEIT &amp; RASHIDI" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT &amp; RASHIDI" baseAuthorityYear="1948" box="[271,538,1673,1695]" class="Globothalamea" family="Orbitolinidae" genus="Gusicella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="9" phylum="Foraminifera" rank="species" species="minima" status="gen. et comb. nov.">
<emphasis id="B965EAC62B494E60FE93D6C5B3FFDEA7" box="[271,442,1673,1695]" italics="true" pageId="6" pageNumber="9">Gusicella minima</emphasis>
(Henson)
</taxonomicName>
<taxonomicNameLabel id="A25657BD2B494E60FDBDD6C6B093DEA7" box="[545,726,1673,1695]" pageId="6" pageNumber="9" rank="species">gen. et comb. nov.</taxonomicNameLabel>
from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section:
<emphasis id="B965EAC62B494E60FEC9D6EBB3E7DE84" bold="true" box="[341,418,1703,1724]" pageId="6" pageNumber="9">ab, j, l</emphasis>
; Mandegan section:
<emphasis id="B965EAC62B494E60FDF2D6EBB09ADE84" bold="true" box="[622,735,1703,1724]" pageId="6" pageNumber="9">ci, k, mp</emphasis>
).
<emphasis id="B965EAC62B494E60FD68D6EBB145DE84" bold="true" box="[756,768,1703,1724]" pageId="6" pageNumber="9">a</emphasis>
Subaxial section.
<emphasis id="B965EAC62B494E60FC2FD6EBB1A2DE84" bold="true" box="[947,999,1703,1724]" pageId="6" pageNumber="9">b, i, l</emphasis>
Oblique sections.
<emphasis id="B965EAC62B494E60FB00D6EBB69BDE84" bold="true" box="[1180,1246,1703,1724]" pageId="6" pageNumber="9">ce, m</emphasis>
Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next.
<emphasis id="B965EAC62B494E60FB38D689B6E9DEE2" bold="true" box="[1188,1196,1733,1754]" pageId="6" pageNumber="9">f</emphasis>
Shallow tangential section displaying subepidermal network of beams and rafters.
<emphasis id="B965EAC62B494E60FCD9D6AFB1F4DEC0" bold="true" box="[837,945,1763,1784]" pageId="6" pageNumber="9">g, j, l, op</emphasis>
Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in
<emphasis id="B965EAC62B494E60FB0ED74DB6DBDF2E" bold="true" box="[1170,1182,1793,1814]" pageId="6" pageNumber="9">o</emphasis>
). Note oval test outline in
<emphasis id="B965EAC62B494E60FE8FD753B364DF0C" bold="true" box="[275,289,1823,1844]" pageId="6" pageNumber="9">j.</emphasis>
Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone.
<emphasis id="B965EAC62B494E60FD5BD770B091DF69" bold="true" box="[711,724,1852,1873]" pageId="6" pageNumber="9">k</emphasis>
Detail from
<emphasis id="B965EAC62B494E60FCCDD770B11BDF69" bold="true" box="[849,862,1852,1873]" pageId="6" pageNumber="9">h</emphasis>
showing marginal foramina arranged in vertical lines between successive chambers.
<emphasis id="B965EAC62B494E60FE64D716B040DF57" bold="true" box="[504,517,1882,1903]" pageId="6" pageNumber="9">n</emphasis>
Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (
<emphasis id="B965EAC62B494E60FBE6D734B6C3DFB5" bold="true" box="[1146,1158,1912,1933]" pageId="6" pageNumber="9">a</emphasis>
), Ng 192 (
<emphasis id="B965EAC62B494E60FB6CD734B6B8DFB5" bold="true" box="[1264,1277,1912,1933]" pageId="6" pageNumber="9">b</emphasis>
), Rt 108-3 (
<emphasis id="B965EAC62B494E60FF4AD7DAB36DDF93" bold="true" box="[214,296,1942,1963]" pageId="6" pageNumber="9">ce, g, n</emphasis>
), Rt 104 (
<emphasis id="B965EAC62B494E60FE17D7DAB3D6DF93" bold="true" box="[395,403,1942,1963]" pageId="6" pageNumber="9">f</emphasis>
), Rt 96 (
<emphasis id="B965EAC62B494E60FE76D7DAB055DF93" bold="true" box="[490,528,1942,1963]" pageId="6" pageNumber="9">h, k</emphasis>
), Rt 108-6 (
<emphasis id="B965EAC62B494E60FD1BD7DAB0CBDF93" bold="true" box="[647,654,1942,1963]" pageId="6" pageNumber="9">i</emphasis>
), Ng 187 (
<emphasis id="B965EAC62B494E60FD6BD7DAB0BADF93" bold="true" box="[759,767,1942,1963]" pageId="6" pageNumber="9">j</emphasis>
), Ng 186 (
<emphasis id="B965EAC62B494E60FCF5D7DAB135DF93" bold="true" box="[873,880,1942,1963]" pageId="6" pageNumber="9">l</emphasis>
), Rt 107 (
<emphasis id="B965EAC62B494E60FC4ED7DAB1A3DF93" bold="true" box="[978,998,1942,1963]" pageId="6" pageNumber="9">m</emphasis>
), Rt 102-2 (
<emphasis id="B965EAC62B494E60FBC0D7DAB62DDF93" bold="true" box="[1116,1128,1942,1963]" pageId="6" pageNumber="9">o</emphasis>
), Rt 104 (
<emphasis id="B965EAC62B494E60FB57D7DAB69DDF93" bold="true" box="[1227,1240,1942,1963]" pageId="6" pageNumber="9">p</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BAE36D42B484E61FF0BD0D2B370D8EF" blockId="7.[151,771,158,215]" pageId="7" pageNumber="10">
Number of chambers last mm axial length: 7-9 (about 7,
<bibRefCitation id="EF804B252B484E61FF0BD08CB368D8EF" author="Henson, F. R. S." box="[151,301,191,215]" pageId="7" pageNumber="10" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B655F2B484E61FF0BD14EB0E9DB74" pageId="7" pageNumber="10" type="discussion">
<paragraph id="8BAE36D42B484E61FF0BD14EB0E9DB74" blockId="7.[151,772,257,844]" pageId="7" pageNumber="10">
<emphasis id="B965EAC62B484E61FF0BD14EB34DD922" bold="true" box="[151,264,258,282]" pageId="7" pageNumber="10">Remarks:</emphasis>
The dimorphism of
<taxonomicName id="4C114D572B484E61FD9DD14EB0E2D921" authorityName="Henson" authorityYear="1948" box="[513,679,257,281]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B484E61FD9DD14EB0E2D921" box="[513,679,257,281]" italics="true" pageId="7" pageNumber="10">D. complanata</emphasis>
</taxonomicName>
is well constrained by the biloculine embryo in the A-form and an indiscernible embryo within a close-coiled whorl including more chambers in the B-form (one section only, see
<figureCitation id="132A2A512B484E61FF5CD1C9B348D9A5" box="[192,269,389,413]" captionStart="Fig" captionStartId="3.[206,241,1801,1822]" captionTargetBox="[206,1408,152,1758]" captionTargetId="figure-21@3.[206,1408,152,1758]" captionTargetPageId="3" captionText="Fig. 4 Dictyoconella complanata Henson from the Maastrichtian of the Tarbur Formation of Iran (Naghan section, a-i, k) and the Simsima Formation of Qatar (j). a, d, oblique sections perpendicular to plane of compression. Note the inclination of the apical-juvenile part expressed by the parallel-sectioning of septum (s) and foramina (f) in d. b, oblique section passing through the initial part of a microspheric specimen. c, f, i-k, oblique sections in the plane of compression (j, paratype from Henson, 1948, pl. 10, fig. 14). Detail from i shows the fine subepidermal cellular network, marginal foramina and undivided marginal trough. e, g-h, oblique sections passing through the initial part of megalospheric specimens. Thin sections: 2Ng 81- 2 (a, d), Ng 76 (b), 2Ng 81-3 (c), Ng 85 (e), 2Ng 87-2 (f, i), 2Ng 80-1 (g), 2Ng 85-4 (h), 2Ng 81-1 (k). Abbreviations: b = beam, deu = deuteroconch, f = foramen m.f. = marginal foramen, m.t. = marginal trough, pi = pillar, pr = proloculus, T = Tarburina zagrosiana Schlagintweit et al. (in k)." figureDoi="http://doi.org/10.5281/zenodo.12583330" httpUri="https://zenodo.org/record/12583330/files/figure.png" pageId="7" pageNumber="10">Fig. 4b</figureCitation>
). The differences in the external test morphology between the two generations are unknown. In the family
<taxonomicName id="4C114D572B484E61FF77D184B339D9D8" authorityName="Martin" authorityYear="1890" box="[235,380,456,480]" class="Globothalamea" family="Orbitolinidae" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="family">Orbitolinidae</taxonomicName>
, B-specimens are generally larger and more flattened (e.g.
<bibRefCitation id="EF804B252B484E61FE2AD1A5B00BDA39" author="Hofker, J. Jr." box="[438,590,489,513]" pageId="7" pageNumber="10" pagination="1 - 34" refId="ref6016" refString="Hofker, J. Jr., 1966. Studies on the family Orbitolinidae. Palaeontographica, Abt. A, 126: 1 - 34." type="journal article" year="1966">Hofker, 1966</bibRefCitation>
). In the Iranian Tarbur Formation,
<taxonomicName id="4C114D572B484E61FEF9D247B040DA1A" authorityName="Henson" authorityYear="1948" box="[357,517,522,546]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B484E61FEF9D247B040DA1A" box="[357,517,522,546]" italics="true" pageId="7" pageNumber="10">D. complanata</emphasis>
</taxonomicName>
is comparably rare and only a couple of sections are available. Due to the revision presented herein,
<taxonomicName id="4C114D572B484E61FE05D200B06ADA5C" authorityName="HENSON" authorityYear="1948" box="[409,559,588,612]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="genus">
<emphasis id="B965EAC62B484E61FE05D200B06ADA5C" box="[409,559,588,612]" italics="true" pageId="7" pageNumber="10">Dictyoconella</emphasis>
</taxonomicName>
becomes a monospecific genus and an assumed Maastrichtian newcomer that became extinct shortly afterwards at the K-Pg boundary. Besides the Tarbur Formation of
<collectingCountry id="F30676442B484E61FDEFD2FCB0E5DAFF" box="[627,672,688,711]" name="Iran" pageId="7" pageNumber="10">Iran</collectingCountry>
,
<taxonomicName id="4C114D572B484E61FD30D2FCB2ACDAD1" authorityName="Henson" authorityYear="1948" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="complanata">
<emphasis id="B965EAC62B484E61FD30D2FCB2ACDAD1" italics="true" pageId="7" pageNumber="10">D. complanata</emphasis>
</taxonomicName>
has been reported so far from the Maastrichtian Garzan Formation of southeastern
<collectingCountry id="F30676442B484E61FDBAD2BEB036DB32" box="[550,627,754,778]" name="Turkey" pageId="7" pageNumber="10">Turkey</collectingCountry>
(
<bibRefCitation id="EF804B252B484E61FD16D2BEB28BDB13" author="Koyluoglu, M." pageId="7" pageNumber="10" pagination="1 - 53" refId="ref6278" refString="Koyluoglu, M., 1986. Guneydogu Anadolu otokton birimlerin kronostratigrafisi, mikrofasiyes ve mikrofosilleri, T. P. A. O. Arastirma Merkezi, Egitim Yayinlari, 9: 1 - 53." type="journal article" year="1986">Köylüoğlu, 1986</bibRefCitation>
) revealing some kind of bioprovincialism, which requires subsequent study (
<bibRefCitation id="EF804B252B484E61FE25D378B0D9DB74" author="Schlagintweit, F." box="[441,668,820,844]" pageId="7" pageNumber="10" pagination="69 - 83" refId="ref6563" refString="Schlagintweit, F., 2020 a. Maastrichtian larger benthic foraminifera from the Arabian Plate sensu lato: new data from Somalia, Turkey and Iran. Acta Palaeontologica Romaniae, 16 (2): 69 - 83." type="journal article" year="2020">Schlagintweit, 2020a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>