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<document id="81A10D278FC91748D55E9DA73476B51F" ID-CLB-Dataset="20049" ID-DOI="10.11646/zootaxa.5120.4.6" ID-GBIF-Dataset="a0421c84-a047-4499-b92f-52cab23d6e79" ID-ISSN="1175-5326" ID-Zenodo-Dep="6392907" ID-ZooBank="58CAB25D-CA92-44EE-BF1D-67DB91EB9596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648538015601" checkinUser="plazi" docAuthor="Martins, Luciana &amp; Tavares, Marcos" docDate="2022" docId="03B887C14B01813BBEE16CC7FA289097" docLanguage="en" docName="zootaxa.5120.4.6.pdf" docOrigin="Zootaxa 5120 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clarkiella discoveryi Heding" docType="treatment" docVersion="7" lastPageNumber="570" masterDocId="FF81FFB94B0B8130BE766D38FFB09611" masterDocTitle="Additions to the morphology of the Phyllophoridae and the Sclerodactylidae. I. Type species of Phyllophorella, Selenkiella, Cladolella and Clarkiella, with the description a new species of Thorsonia (Holothuroidea: Dendrochirotida)" masterLastPageNumber="572" masterPageNumber="559" pageNumber="569" updateTime="1699318708264" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F3C4122522ED5790A9821427C199A5DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CAE7540223AACCE6DEAC83E0EB9A254F">
<mods:title id="55456D7CCCB1A854F825D125F3B263C8">Additions to the morphology of the Phyllophoridae and the Sclerodactylidae. I. Type species of Phyllophorella, Selenkiella, Cladolella and Clarkiella, with the description a new species of Thorsonia (Holothuroidea: Dendrochirotida)</mods:title>
</mods:titleInfo>
<mods:name id="65B4A1583EDC765F116DFF80E53AAE21" type="personal">
<mods:role id="49860D2734EB1C3AC730F674E4333C60">
<mods:roleTerm id="C1FB688511A844AFFCA71D244125E576">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C79006F32D7CF15E25D48A903699CD61">Martins, Luciana</mods:namePart>
<mods:nameIdentifier id="A3A4BB9A18C21050A927C56AC89AC24B" type="ORCID">0000-0002-8107-3265</mods:nameIdentifier>
<mods:affiliation id="3B400F9A207F48EFDD80B3B61AE74463">Museu de Zoologia, Universidade de São Paulo, São Paulo-SP, 04263 - 000, Brazil. &amp; martinsrluciana @ gmail. com, mdst @ usp. br; https: // orcid. org / 0000 - 0002 - 8107 - 3265</mods:affiliation>
<mods:nameIdentifier id="3BF7F4BFC335AEE423A367B5B6D62382" type="email">mdst@usp.br</mods:nameIdentifier>
</mods:name>
<mods:name id="3A29BFADEBFCAC729C8BECD1A61034E1" type="personal">
<mods:role id="B62ABDEDBA1F414B08361482FB9DE269">
<mods:roleTerm id="6C0F2D5B53E0A5F56ACD887360ED3C71">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4819B3F6C3277F39304B144B80D5C5E6">Tavares, Marcos</mods:namePart>
<mods:nameIdentifier id="D815B7FE233A103CA90E2EEBB6B16A33" type="ORCID">0000-0002-7186-5787</mods:nameIdentifier>
<mods:affiliation id="181C21E576C14533E3DF48073C19EB68">Museu de Zoologia, Universidade de São Paulo, São Paulo-SP, 04263 - 000, Brazil. &amp; https: // orcid. org / 0000 - 0002 - 7186 - 5787</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0783228A55BB03D91C6944682BE19E75">text</mods:typeOfResource>
<mods:relatedItem id="890191B14329EA0B2F34E03915537A7B" type="host">
<mods:titleInfo id="AF3459C6E27D8642C687643486D35784">
<mods:title id="BD2F2F43753120C83DDE3791EB62B820">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D0A9835F01075FD58BC35E0152B1A0BF">
<mods:date id="61EB3AD13FC86C07DC3DDE273FB7CECB">2022</mods:date>
<mods:detail id="83BD32CC019EBDCFE82F624D65FB18CC" type="pubDate">
<mods:number id="13BD078DFB4967CE804E22485202E01F">2022-03-29</mods:number>
</mods:detail>
<mods:detail id="DA03E10811D4853C34844AD455954410" type="volume">
<mods:number id="ECFC43A8158E6ED37AF2A3D1B4D83559">5120</mods:number>
</mods:detail>
<mods:detail id="8CC2041C865C7BD93BEB2C1FFD50A056" type="issue">
<mods:number id="0F8186A86BC3603A3286BF25A466A372">4</mods:number>
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<mods:start id="3A78631507098ABF87EFDCB9A82677EF">559</mods:start>
<mods:end id="99B486591AAAA7EFE5EE61D59908684E">572</mods:end>
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<mods:classification id="BD5C787674CA025DC6FEF22848938370">journal article</mods:classification>
<mods:identifier id="73DD58703DAE9392DFDE66329587D437" type="CLB-Dataset">20049</mods:identifier>
<mods:identifier id="590E42A133E5E566C341C1E9E1E616E4" type="DOI">10.11646/zootaxa.5120.4.6</mods:identifier>
<mods:identifier id="20E2631F14031624356035D412E1B148" type="GBIF-Dataset">a0421c84-a047-4499-b92f-52cab23d6e79</mods:identifier>
<mods:identifier id="E019A369EF8F560B0231A690DCD50882" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6CDB36FB97C59C5D0A2D5E03F6EA6B17" type="Zenodo-Dep">6392907</mods:identifier>
<mods:identifier id="A715639380FE2D5E13C8F6A7D65A1567" type="ZooBank">58CAB25D-CA92-44EE-BF1D-67DB91EB9596</mods:identifier>
</mods:mods>
<treatment id="03B887C14B01813BBEE16CC7FA289097" ID-DOI="http://doi.org/10.5281/zenodo.6402237" ID-GBIF-Taxon="194317681" ID-Zenodo-Dep="6402237" LSID="urn:lsid:plazi:treatment:03B887C14B01813BBEE16CC7FA289097" httpUri="http://treatment.plazi.org/id/03B887C14B01813BBEE16CC7FA289097" lastPageId="11" lastPageNumber="570" pageId="10" pageNumber="569">
<subSubSection id="C30B655C4B01813ABEE16CC7FCE59408" box="[151,853,511,538]" pageId="10" pageNumber="569" type="nomenclature">
<paragraph id="8BAE36D74B01813ABEE16CC7FCE59408" blockId="10.[151,853,511,573]" box="[151,853,511,538]" pageId="10" pageNumber="569">
<heading id="D0E681BB4B01813ABEE16CC7FCE59408" bold="true" box="[151,853,511,538]" fontSize="11" level="1" pageId="10" pageNumber="569" reason="1">
<emphasis id="B965EAC54B01813ABEE16CC7FCE59408" bold="true" box="[151,853,511,538]" pageId="10" pageNumber="569">
<taxonomicName id="4C114D544B01813ABEE16CC7FE449408" ID-CoL="VRNH" authority="Heding" authorityName="Heding" box="[151,500,511,538]" class="Holothuroidea" family="Sclerodactylidae" genus="Clarkiella" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="569" phylum="Echinodermata" rank="species" species="discoveryi">
<emphasis id="B965EAC54B01813ABEE16CC7FE20940B" bold="true" box="[151,400,511,538]" italics="true" pageId="10" pageNumber="569">Clarkiella discoveryi</emphasis>
Heding
</taxonomicName>
, in
<bibRefCitation id="EF804B264B01813ABC576CC7FCE59408" author="Heding, S. G. &amp; Panning, A." box="[545,853,511,537]" pageId="10" pageNumber="569" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B655C4B01813ABEE16F1BFEFA942C" box="[151,330,547,573]" pageId="10" pageNumber="569" type="description">
<paragraph id="8BAE36D74B01813ABEE16F1BFEFA942C" blockId="10.[151,853,511,573]" box="[151,330,547,573]" pageId="10" pageNumber="569">
(
<figureCitation id="132A2A524B01813ABEE96F1BFEF1942C" box="[159,321,547,573]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="10.[151,250,1800,1824]" captionStartId-1="11.[151,250,1500,1524]" captionTargetBox-0="[153,1430,664,1777]" captionTargetBox-1="[151,1436,777,1477]" captionTargetId-0="figure-100@10.[151,1436,664,1777]" captionTargetId-1="figure-283@11.[151,1436,777,1477]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." captionText-1="FIGURE 12. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). Ossicles from body wall (a–b) table in (a) top and (b) bottom view; (c–d) plates from tentacle; (e–f) rosettes from introvert." figureDoi-0="http://doi.org/10.5281/zenodo.6392932" figureDoi-1="http://doi.org/10.5281/zenodo.6392934" httpUri-0="https://zenodo.org/record/6392932/files/figure.png" httpUri-1="https://zenodo.org/record/6392934/files/figure.png" pageId="10" pageNumber="569">
Figures 11
<emphasis id="B965EAC54B01813ABF606F1CFE94942C" box="[278,292,548,573]" italics="true" pageId="10" pageNumber="569">–</emphasis>
12
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B655C4B01813BBEE16F50FC2D96E8" lastPageId="11" lastPageNumber="570" pageId="10" pageNumber="569" type="materials_examined">
<paragraph id="8BAE36D74B01813BBEE16F50FC2D96E8" blockId="10.[151,909,615,640]" lastBlockId="11.[151,1437,150,753]" lastPageId="11" lastPageNumber="570" pageId="10" pageNumber="569">
<taxonomicName id="4C114D544B01813ABEE16F50FE709491" authority="Heding" authorityName="Heding" box="[151,448,616,640]" class="Holothuroidea" family="Sclerodactylidae" genus="Clarkiella" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="569" phylum="Echinodermata" rank="species" species="discoveryi">
<emphasis id="B965EAC54B01813ABEE16F50FEDC9491" box="[151,364,616,640]" italics="true" pageId="10" pageNumber="569">Clarkiella discoveryi</emphasis>
Heding
</taxonomicName>
, in
<bibRefCitation id="EF804B264B01813ABF926F50FCF89491" author="Heding, S. G. &amp; Panning, A." box="[484,840,615,640]" pageId="10" pageNumber="569" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">
Heding &amp; Panning, 1954: 119
<emphasis id="B965EAC54B01813ABD626F50FC91946E" box="[788,801,616,639]" italics="true" pageId="10" pageNumber="569">–</emphasis>
120
</bibRefCitation>
, pl.49.
<emphasis id="B965EAC54B00813BBEE16DAEFE3596A1" bold="true" box="[151,389,150,176]" pageId="11" pageNumber="570">Material examined.</emphasis>
<materialsCitation id="3B793C8A4B00813BBFE76DAFFED996C7" ID-GBIF-Occurrence="3744207307" collectingDate="1930-11-19" collectionCode="ZMUC" collectorName="Mil W." country="South Georgia and the South Sandwich Islands" elevation="199" location="South Georgia" pageId="11" pageNumber="570" specimenCode="HOL 64" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AA88754B00813BBFE76DAFFE4E96A0" box="[401,510,151,177]" pageId="11" pageNumber="570" type="holotype">Holotype</typeStatus>
: 1
<collectorName id="26E453014B00813BBC516DAFFDCA96A0" box="[551,634,151,177]" pageId="11" pageNumber="570">Mil W.</collectorName>
off. Shag Rocks,
<collectingCountry id="F30676474B00813BBD236DAFFC4F96A0" box="[853,1023,151,177]" name="South Georgia and the South Sandwich Islands" pageId="11" pageNumber="570">South Georgia</collectingCountry>
,
<quantity id="4CE99B324B00813BBA786DAFFBEC96A0" box="[1038,1116,151,177]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" pageId="11" pageNumber="570" unit="m" value="199.0">
<elevation id="003CD1E44B00813BBA786DAFFBEC96A0" box="[1038,1116,151,177]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" pageId="11" pageNumber="570" unit="m" value="199.0">199 m</elevation>
</quantity>
,
<date id="FFAF10174B00813BBA1A6DAFFB5696A0" box="[1132,1254,151,177]" pageId="11" pageNumber="570" value="1930-11-19">
<collectingDate id="EFEBE9FF4B00813BBA1A6DAFFB5696A0" box="[1132,1254,151,177]" pageId="11" pageNumber="570" value="1930-11-19">
19.xi.
<quantity id="4CE99B324B00813BBADE6DAFFB5696A0" box="[1192,1254,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.9306" pageId="11" pageNumber="570" unit="mm" value="1930.6">1930</quantity>
</collectingDate>
</date>
, 60 mm length (
<collectionCode id="ED00AE124B00813BBEE86D83FF5F96C4" box="[158,239,187,213]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="570">ZMUC</collectionCode>
–
<specimenCode id="DBB79EAC4B00813BBE886D83FED096C7" box="[254,352,187,214]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="570">HOL 64</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B793C8A4B00813BBF026D83FBC196C4" ID-GBIF-Occurrence="3744207302" box="[372,1137,186,213]" collectingDate="1930-11-19" collectionCode="ZMUC" collectorName="Mil W." country="South Georgia and the South Sandwich Islands" elevation="199" location="South Georgia" pageId="11" pageNumber="570" specimenCode="HOL 247" specimenCount="1" typeStatus="paratype">
<typeStatus id="54AA88754B00813BBF026D83FE6A96C4" box="[372,474,187,213]" pageId="11" pageNumber="570" type="paratype">Paratype</typeStatus>
: same data as holotype,
<quantity id="4CE99B324B00813BBC9B6D82FC8D96C4" box="[749,829,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="570" unit="mm" value="45.0">45 mm</quantity>
length (
<collectionCode id="ED00AE124B00813BBDEF6D83FC5A96C4" box="[921,1002,187,213]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="570">ZMUC</collectionCode>
–
<specimenCode id="DBB79EAC4B00813BBD8F6D83FBD896C4" box="[1017,1128,187,213]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="570">HOL 247</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B793C8A4B00813BBA0A6D83FC2D96E8" ID-GBIF-Occurrence="3744207301" collectionCode="MNHN, IE" country="South Georgia and the South Sandwich Islands" pageId="11" pageNumber="570" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AA88754B00813BBA0A6D83FB5496C4" box="[1148,1252,187,213]" pageId="11" pageNumber="570" type="holotype">Holotype</typeStatus>
Slides
<collectionCode id="ED00AE124B00813BBB4F6D83FA3E96C4" box="[1337,1422,187,213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="570">MNHN</collectionCode>
–
<collectionCode id="ED00AE124B00813BBEE16DE6FF0296E8" box="[151,178,222,249]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="11" pageNumber="570">IE</collectionCode>
–2005–6324); (
<collectionCode id="ED00AE124B00813BBF2B6DE6FE0696E8" box="[349,438,222,249]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="570">MNHN</collectionCode>
–
<collectionCode id="ED00AE124B00813BBFB36DE6FE5096E8" box="[453,480,222,249]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="11" pageNumber="570">IE</collectionCode>
–2005–6325); (
<collectionCode id="ED00AE124B00813BBCFF6DE6FD5196E8" box="[649,737,222,249]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="570">MNHN</collectionCode>
–
<collectionCode id="ED00AE124B00813BBC996DE6FCBA96E8" box="[751,778,222,249]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="11" pageNumber="570">IE</collectionCode>
–2005–6326)
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF6E665F4B01813ABEE16A30FDBD91C4" ID-DOI="http://doi.org/10.5281/zenodo.6392932" ID-Zenodo-Dep="6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="10" pageNumber="569" startId="10.[151,250,1800,1824]" targetBox="[153,1430,664,1777]" targetPageId="10">
<paragraph id="8BAE36D74B01813ABEE16A30FDBD91C4" blockId="10.[151,1436,1800,2005]" pageId="10" pageNumber="569">
<emphasis id="B965EAC54B01813ABEE16A30FE909131" bold="true" box="[151,288,1800,1824]" pageId="10" pageNumber="569">FIGURE 11.</emphasis>
<taxonomicName id="4C114D544B01813ABF5E6A31FDE59130" authority="Heding" authorityName="Heding" box="[296,597,1801,1825]" class="Holothuroidea" family="Sclerodactylidae" genus="Clarkiella" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="569" phylum="Echinodermata" rank="species" species="discoveryi">
<emphasis id="B965EAC54B01813ABF5E6A31FE4F9130" box="[296,511,1801,1825]" italics="true" pageId="10" pageNumber="569">Clarkiella discoveryi</emphasis>
Heding
</taxonomicName>
, in
<bibRefCitation id="EF804B264B01813ABC0B6A31FC329131" author="Heding, S. G. &amp; Panning, A." box="[637,898,1800,1825]" pageId="10" pageNumber="569" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by
<bibRefCitation id="EF804B264B01813ABF586A85FE0E91C4" author="Ludwig, H." box="[302,446,1981,2005]" pageId="10" pageNumber="569" pagination="377 - 460" refId="ref5943" refString="Ludwig, H. (1889 - 1892) Die Seewalzen. In: Bronn HG (Ed.), Klassen und Ordnungen des Thier-Reichs, wissenschaftlich dargestellt in Wort und Bild, vol. 2, Part 3: Echinodermen (Stachelhauter), First book. Leipzig: C. F. Winter. [1889: 1 - 176, pls. 1 - 8; 1890: 177 - 240, pls. 9 - 12; 1891: 241 - 376, pls. 13 - 17; 1892: 377 - 460]" type="journal article" year="1889">Ludwig (1889</bibRefCitation>
–1892).
</paragraph>
</caption>
<subSubSection id="C30B655C4B00813BBEB16C3AFE8F94E0" pageId="11" pageNumber="570" type="description">
<paragraph id="8BAE36D74B00813BBEB16C3AFCB7942C" blockId="11.[151,1437,150,753]" pageId="11" pageNumber="570">
<emphasis id="B965EAC54B00813BBEB16C3AFEC5970D" bold="true" box="[199,373,258,284]" pageId="11" pageNumber="570">Redescription.</emphasis>
Body U–shaped; tube feet scattered throughout body (
<figureCitation id="132A2A524B00813BBD9D6C3BFBFD970C" box="[1003,1101,259,285]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11a</figureCitation>
). Color, in ethanol, beige. 15 dendritic tentacles, two ventral smaller (
<figureCitation id="132A2A524B00813BBC176C1FFD6E9750" box="[609,734,295,321]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11b, c</figureCitation>
). Anal papillae present (
<figureCitation id="132A2A524B00813BBD836C1FFBE69753" box="[1013,1110,295,322]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11d</figureCitation>
). Longitudinal, retractor and circular muscles thin (
<figureCitation id="132A2A524B00813BBFE16C73FDA39777" box="[407,531,331,358]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11e, f</figureCitation>
). Longitudinal muscles split posteriorly (
<figureCitation id="132A2A524B00813BBD9A6C73FBFB9777" box="[1004,1099,331,358]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11f</figureCitation>
). Retractor muscles short, attached to middle of radial plate (
<figureCitation id="132A2A524B00813BBC786C57FDC0979B" box="[526,624,367,394]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11h</figureCitation>
). Stone canal single, short, ending in elongated madreporite (
<figureCitation id="132A2A524B00813BBB5B6C57FA3F9798" box="[1325,1423,367,393]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">
Fig.
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</figureCitation>
). Radial and interradial plates of calcareous ring of same height, laterally connected to each other for most of their length (
<figureCitation id="132A2A524B00813BBE9B6C8FFED397C3" box="[237,355,439,466]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11h, i</figureCitation>
). MidIR (IR5) taller than other, lacking posterior projection (
<figureCitation id="132A2A524B00813BBA616C8FFBC297C3" box="[1047,1138,439,466]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="11" pageNumber="570">Fig. 10i</figureCitation>
); midR (RI) not modified. Radial plates undivided, rectangular, convex base with posterior projection and 2 AP, one wider than the other (
<figureCitation id="132A2A524B00813BBB196CE3FF75940B" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="11" pageNumber="570">Fig. 10h</figureCitation>
). Interradial plates undivided, elongated, with two posterior projections and pointed anterior end (
<figureCitation id="132A2A524B00813BBB666CC7FA3D940B" box="[1296,1421,511,538]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="11" pageNumber="570">Fig. 10 h, i</figureCitation>
). radial and interradial plates lacking posterior processes.
</paragraph>
<paragraph id="8BAE36D74B00813BBEB16F7FFE8F94E0" blockId="11.[151,1437,150,753]" pageId="11" pageNumber="570">
Body wall with two–pillared tables with circular disc (80–100 μm long;
<figureCitation id="132A2A524B00813BBD926F7FFBEC9473" box="[996,1116,583,610]" captionStart="FIGURE 12" captionStartId="11.[151,250,1500,1524]" captionTargetBox="[151,1436,777,1477]" captionTargetId="figure-283@11.[151,1436,777,1477]" captionTargetPageId="11" captionText="FIGURE 12. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). Ossicles from body wall (a–b) table in (a) top and (b) bottom view; (c–d) plates from tentacle; (e–f) rosettes from introvert." figureDoi="http://doi.org/10.5281/zenodo.6392934" httpUri="https://zenodo.org/record/6392934/files/figure.png" pageId="11" pageNumber="570">Fig.12 a, b</figureCitation>
); disc with four central holes and 8–10 marginal perforations, margin undulated; spire ending in about 12 spines. Tentacles with perforated plates (100–150 μm long;
<figureCitation id="132A2A524B00813BBF056FB7FE5694BB" box="[371,486,655,682]" captionStart="FIGURE 12" captionStartId="11.[151,250,1500,1524]" captionTargetBox="[151,1436,777,1477]" captionTargetId="figure-283@11.[151,1436,777,1477]" captionTargetPageId="11" captionText="FIGURE 12. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). Ossicles from body wall (a–b) table in (a) top and (b) bottom view; (c–d) plates from tentacle; (e–f) rosettes from introvert." figureDoi="http://doi.org/10.5281/zenodo.6392934" httpUri="https://zenodo.org/record/6392934/files/figure.png" pageId="11" pageNumber="570">Fig.12c, d</figureCitation>
). Introvert with closed rosettes (70–110 μm long;
<figureCitation id="132A2A524B00813BBA666FB7FB3594BB" box="[1040,1157,655,682]" captionStart="FIGURE 12" captionStartId="11.[151,250,1500,1524]" captionTargetBox="[151,1436,777,1477]" captionTargetId="figure-283@11.[151,1436,777,1477]" captionTargetPageId="11" captionText="FIGURE 12. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). Ossicles from body wall (a–b) table in (a) top and (b) bottom view; (c–d) plates from tentacle; (e–f) rosettes from introvert." figureDoi="http://doi.org/10.5281/zenodo.6392934" httpUri="https://zenodo.org/record/6392934/files/figure.png" pageId="11" pageNumber="570">Fig. 12e, f</figureCitation>
). Tube feet with endplate with small holes, larger ones around margin, smaller ones medially, endplates up to 150 μm in diameter and tables like body wall.
</paragraph>
</subSubSection>
<caption id="DF6E665F4B00813BBEE168E4FBCC9009" ID-DOI="http://doi.org/10.5281/zenodo.6392934" ID-Zenodo-Dep="6392934" httpUri="https://zenodo.org/record/6392934/files/figure.png" pageId="11" pageNumber="570" startId="11.[151,250,1500,1524]" targetBox="[151,1436,777,1477]" targetPageId="11">
<paragraph id="8BAE36D74B00813BBEE168E4FBCC9009" blockId="11.[151,1437,1500,1561]" pageId="11" pageNumber="570">
<emphasis id="B965EAC54B00813BBEE168E4FE9093E5" bold="true" box="[151,288,1500,1524]" pageId="11" pageNumber="570">FIGURE 12.</emphasis>
<taxonomicName id="4C114D544B00813BBF5068E4FDFF93E5" authority="Heding" authorityName="Heding" box="[294,591,1500,1524]" class="Holothuroidea" family="Sclerodactylidae" genus="Clarkiella" kingdom="Animalia" order="Dendrochirotida" pageId="11" pageNumber="570" phylum="Echinodermata" rank="species" species="discoveryi">
<emphasis id="B965EAC54B00813BBF5068E4FE4B93E5" box="[294,507,1500,1524]" italics="true" pageId="11" pageNumber="570">Clarkiella discoveryi</emphasis>
Heding
</taxonomicName>
, in
<bibRefCitation id="EF804B264B00813BBC0568E4FCC393E5" author="Heding, S. G. &amp; Panning, A." box="[627,883,1500,1525]" pageId="11" pageNumber="570" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
, holotype (ZMUC-HOL 64). Ossicles from body wall (a–b) table in (a) top and (b) bottom view; (c–d) plates from tentacle; (e–f) rosettes from introvert.
</paragraph>
</caption>
<subSubSection id="C30B655C4B00813BBEB16B7FFA289097" pageId="11" pageNumber="570" type="discussion">
<paragraph id="8BAE36D74B00813BBEB16B7FFA289097" blockId="11.[151,1437,1607,1670]" pageId="11" pageNumber="570">
<emphasis id="B965EAC54B00813BBEB16B7FFE8B9070" bold="true" box="[199,315,1607,1633]" pageId="11" pageNumber="570">Remarks.</emphasis>
Here we provide a photograph of the general aspect of the
<typeStatus id="54AA88754B00813BBDB16B7FFB999070" box="[967,1065,1607,1633]" pageId="11" pageNumber="570" type="holotype">holotype</typeStatus>
(
<figureCitation id="132A2A524B00813BBA416B7FFB239070" box="[1079,1171,1607,1633]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11a</figureCitation>
) and describe by means optical microscopy for the first time the morphology of its longitudinal retractor and circular muscles (
<figureCitation id="132A2A524B00813BBB626B53FA3B9097" box="[1300,1419,1643,1670]" captionStart="FIGURE 11" captionStartId="10.[151,250,1800,1824]" captionTargetBox="[153,1430,664,1777]" captionTargetId="figure-100@10.[151,1436,664,1777]" captionTargetPageId="10" captionText="FIGURE 11. Clarkiella discoveryi Heding, in Heding &amp; Panning, 1954, holotype (ZMUC-HOL 64). (a) specimen in lateral view; (b) top and (c) lateral view of tentacles; (d) anal papillae; (e–f) muscular system and gonads; (g) madreporite and stone canal; (h) photograph of the calcareous ring in outer view showing the insertion of the retractor muscle and (i) photograph of the calcareous ring in outer view showing the midIR (IR5), RIV and RV.Abbreviations: RM, retractor muscle, CM, circular muscle, LM, longitudinal muscle, GO, gonads, PV, Polian vesicle and M, madreporite, SC, stone canal. Plate identity follows the system established by Ludwig (1889–1892)." figureDoi="http://doi.org/10.5281/zenodo.6392932" httpUri="https://zenodo.org/record/6392932/files/figure.png" pageId="11" pageNumber="570">Fig. 11e, f</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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