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<document ID-DOI="10.11646/zootaxa.5244.6.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7673035" ID-ZooBank="8182831B-25FC-4B92-B319-281659660565" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1677225701698" checkinUser="plazi" docAuthor="Kluge, Nikita" docDate="2023" docId="03AE0E6FFFD3160502DC923363D4FE19" docLanguage="en" docName="zootaxa.5244.6.2.pdf" docOrigin="Zootaxa 5244 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.6.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Teloganodes (Dudgeodes) rasus Kluge 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="551" masterDocId="FF977617FFDC161D024B957B6076FFC8" masterDocTitle="Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae)" masterLastPageNumber="552" masterPageNumber="527" pageNumber="542" updateTime="1677226200810" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kluge, Nikita</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5244</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>527</mods:start>
<mods:end>552</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.6.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5244.6.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7673035</mods:identifier>
<mods:identifier type="ZooBank">8182831B-25FC-4B92-B319-281659660565</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7673043" ID-Zenodo-Dep="7673043" LSID="urn:lsid:plazi:treatment:03AE0E6FFFD3160502DC923363D4FE19" httpUri="http://treatment.plazi.org/id/03AE0E6FFFD3160502DC923363D4FE19" lastPageId="24" lastPageNumber="551" pageId="15" pageNumber="542">
<subSubSection box="[151,608,1864,1891]" pageId="15" pageNumber="542" type="nomenclature">
<paragraph blockId="15.[151,608,1864,1927]" box="[151,608,1864,1891]" pageId="15" pageNumber="542">
<heading bold="true" box="[151,608,1864,1891]" fontSize="11" level="1" pageId="15" pageNumber="542" reason="1">
<emphasis bold="true" box="[151,608,1864,1891]" pageId="15" pageNumber="542">
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[151,534,1864,1891]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="542" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis bold="true" box="[151,301,1864,1891]" italics="true" pageId="15" pageNumber="542">Teloganodes</emphasis>
(
<emphasis bold="true" box="[319,453,1864,1891]" italics="true" pageId="15" pageNumber="542">Dudgeodes</emphasis>
)
<emphasis bold="true" box="[469,534,1865,1891]" italics="true" pageId="15" pageNumber="542">rasus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[542,608,1864,1890]" pageId="15" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,309,1900,1927]" pageId="15" pageNumber="542" type="description">
<paragraph blockId="15.[151,608,1864,1927]" box="[151,309,1900,1927]" pageId="15" pageNumber="542">
(
<figureCitation box="[159,301,1900,1927]" captionStart-0="FIGURES 50–60" captionStart-1="FIGURES 61–62" captionStart-2="FIGURES 63–68" captionStart-3="FIGURES 69–74" captionStart-4="FIGURES 75–83" captionStart-5="FIGURES 84–92" captionStart-6="FIGURES 93–94" captionStart-7="FIGURES 95–101" captionStartId-0="16.[151,264,1932,1956]" captionStartId-1="17.[151,264,1223,1247]" captionStartId-2="18.[151,264,1827,1851]" captionStartId-3="19.[151,264,1841,1865]" captionStartId-4="20.[151,264,1845,1869]" captionStartId-5="21.[151,264,1841,1865]" captionStartId-6="22.[151,264,1218,1242]" captionStartId-7="23.[151,264,1926,1950]" captionTargetBox-0="[151,1436,290,1909]" captionTargetBox-1="[159,1419,181,1200]" captionTargetBox-2="[151,1436,181,1797]" captionTargetBox-3="[151,1436,181,1817]" captionTargetBox-4="[151,1418,181,1821]" captionTargetBox-5="[151,1431,181,1808]" captionTargetBox-6="[151,1435,186,1189]" captionTargetBox-7="[151,1436,273,1903]" captionTargetId-0="figure-61@16.[151,1436,273,1909]" captionTargetId-1="figure-15@17.[159,1428,181,1200]" captionTargetId-2="figure-23@18.[151,1436,181,1803]" captionTargetId-3="figure-15@19.[151,1436,181,1817]" captionTargetId-4="figure-23@20.[151,1436,181,1821]" captionTargetId-5="figure-15@21.[151,1436,181,1817]" captionTargetId-6="figure-23@22.[151,1436,181,1193]" captionTargetId-7="figure-56@23.[151,1436,273,1903]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." captionText-1="FIGURES 61–62. Teloganodes (Dudgeodes) rasus sp. n. 61, male larva of last instar; 62, younger female larva." captionText-2="FIGURES 63–68. Teloganodes (Dudgeodes) rasus sp. n. 63–64, left and right mandibles; 65, maxilla, 66, apex of maxilla; 67, hypopharynx and superlingua; 68, labium. Abbreviations: in, incisor; kd, kinetodontium; m, mola; prs, prostheca." captionText-3="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." captionText-4="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." captionText-5="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." captionText-6="FIGURES 93–94. Teloganodes (Dudgeodes) rasus sp. n., wings of male imago (holotype). 93, in falling light; 94, in transmitted light (arrow shows constant crossvein cua-cup)." captionText-7="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi-0="http://doi.org/10.5281/zenodo.7673061" figureDoi-1="http://doi.org/10.5281/zenodo.7673063" figureDoi-2="http://doi.org/10.5281/zenodo.7673065" figureDoi-3="http://doi.org/10.5281/zenodo.7673067" figureDoi-4="http://doi.org/10.5281/zenodo.7673069" figureDoi-5="http://doi.org/10.5281/zenodo.7673071" figureDoi-6="http://doi.org/10.5281/zenodo.7673077" figureDoi-7="http://doi.org/10.5281/zenodo.7673083" httpUri-0="https://zenodo.org/record/7673061/files/figure.png" httpUri-1="https://zenodo.org/record/7673063/files/figure.png" httpUri-2="https://zenodo.org/record/7673065/files/figure.png" httpUri-3="https://zenodo.org/record/7673067/files/figure.png" httpUri-4="https://zenodo.org/record/7673069/files/figure.png" httpUri-5="https://zenodo.org/record/7673071/files/figure.png" httpUri-6="https://zenodo.org/record/7673077/files/figure.png" httpUri-7="https://zenodo.org/record/7673083/files/figure.png" pageId="15" pageNumber="542">Figs 54–101</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="542" type="etymology">
<paragraph blockId="15.[151,1436,1972,2034]" pageId="15" pageNumber="542">
<emphasis bold="true" box="[151,280,1972,1998]" pageId="15" pageNumber="542">Etymology</emphasis>
:
<taxonomicName authority="(Lat.)" baseAuthorityName="Lat." box="[296,422,1972,1999]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="542" phylum="Arthropoda" rank="species" species="rasus">rasus (Lat.)</taxonomicName>
—shaved; a hint that all stout setae on the larval head, body and legs are very short, as if the insect had been shaved.
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="551" pageId="16" pageNumber="543" type="materials_examined">
<paragraph blockId="16.[151,1436,150,249]" pageId="16" pageNumber="543">
<emphasis bold="true" box="[199,434,150,176]" pageId="16" pageNumber="543">Material examined.</emphasis>
<materialsCitation collectingDate="2009-08-23" collectorName="N. Kluge &amp; L. Sheyko." country="Indonesia" location="river Mamasa" pageId="16" pageNumber="543" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<emphasis bold="true" box="[441,560,151,177]" pageId="16" pageNumber="543">
<typeStatus box="[441,554,151,177]" pageId="16" pageNumber="543">Holotype</typeStatus>
:
</emphasis>
L-S-IJ {specimen [XI](8)A2009},
<collectingCountry box="[971,1131,151,177]" name="Indonesia" pageId="16" pageNumber="543">INDONESIA</collectingCountry>
,
<collectingRegion box="[1147,1293,151,177]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="543">SULAWESI</collectingRegion>
, tributary of
<location LSID="urn:lsid:plazi:treatment:03AE0E6FFFD3160502DC923363D4FE19:8ED8E9A2FFCC160D02DC95C06140FF1D" box="[151,310,187,213]" country="Indonesia" name="river Mamasa" pageId="16" pageNumber="543" stateProvince="Sulawesi Utara">river Mamasa</location>
<locationDeviation box="[320,525,186,213]" location="Mamasa" pageId="16" pageNumber="543">
<quantity box="[320,381,186,213]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="543" unit="km" value="5.0">5 km</quantity>
W Mamasa
</locationDeviation>
,
<date box="[540,689,187,213]" pageId="16" pageNumber="543" value="2009-08-23">
<collectingDate box="[540,689,187,213]" pageId="16" pageNumber="543" value="2009-08-23">23.VIII.2009</collectingDate>
</date>
, coll.
<collectorName box="[763,869,187,213]" pageId="16" pageNumber="543">N. Kluge</collectorName>
&amp;
<collectorName box="[912,1039,187,213]" pageId="16" pageNumber="543">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation collectingDate="2009-08-15" collectingDateMax="2009-08-27" collectingDateMin="2009-08-15" collectorName="N. Kluge &amp; L. Sheyko." country="Indonesia" location="river Mamasa" pageId="16" pageNumber="543" specimenCount="4" specimenCount-larva="4" stateProvince="L-SJ" typeStatus="paratype">
<emphasis bold="true" box="[1049,1178,187,213]" pageId="16" pageNumber="543">
<typeStatus box="[1049,1173,187,213]" pageId="16" pageNumber="543">Paratypes</typeStatus>
:
</emphasis>
the same locality and collectors,
<date box="[274,466,222,249]" pageId="16" pageNumber="543" value="2009-08-15" valueMax="2009-08-27" valueMin="2009-08-15">
<collectingDate box="[274,466,222,249]" pageId="16" pageNumber="543" value="2009-08-15" valueMax="2009-08-27" valueMin="2009-08-15">15–27.VIII.2009</collectingDate>
</date>
: 1
<collectingRegion box="[498,563,223,249]" country="Indonesia" name="L-SJ" pageId="16" pageNumber="543">L-SJ</collectingRegion>
,
<specimenCount box="[575,668,223,249]" pageId="16" pageNumber="543" type="larva">4 larvae</specimenCount>
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673061" ID-Zenodo-Dep="7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="16" pageNumber="543" startId="16.[151,264,1932,1956]" targetBox="[151,1436,290,1909]" targetPageId="16">
<paragraph blockId="16.[151,1436,1932,1992]" pageId="16" pageNumber="543">
<emphasis bold="true" box="[151,340,1932,1956]" pageId="16" pageNumber="543">FIGURES 50–60.</emphasis>
Tergalii. 50–53,
<taxonomicName box="[512,886,1932,1956]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="543" phylum="Arthropoda" rank="species" species="celebensis" subGenus="Dudgeodes">
<emphasis box="[512,639,1932,1956]" italics="true" pageId="16" pageNumber="543">Teloganodes</emphasis>
(
<emphasis box="[653,767,1932,1956]" italics="true" pageId="16" pageNumber="543">Dudgeodes</emphasis>
)
<emphasis box="[782,886,1932,1956]" italics="true" pageId="16" pageNumber="543">celebensis</emphasis>
</taxonomicName>
, tergalii II–V; 54–57,
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[1111,1436,1932,1956]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="543" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[1111,1238,1932,1956]" italics="true" pageId="16" pageNumber="543">Teloganodes</emphasis>
(
<emphasis box="[1253,1367,1932,1956]" italics="true" pageId="16" pageNumber="543">Dudgeodes</emphasis>
)
<emphasis box="[1381,1436,1933,1956]" italics="true" pageId="16" pageNumber="543">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[151,208,1968,1992]" pageId="16" pageNumber="543">
<taxonomicNameLabel box="[151,208,1968,1992]" pageId="16" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673063" ID-Zenodo-Dep="7673063" box="[151,1291,1223,1248]" httpUri="https://zenodo.org/record/7673063/files/figure.png" pageId="17" pageNumber="544" startId="17.[151,264,1223,1247]" targetBox="[159,1419,181,1200]" targetPageId="17">
<paragraph blockId="17.[151,1291,1223,1248]" box="[151,1291,1223,1248]" pageId="17" pageNumber="544">
<emphasis bold="true" box="[151,340,1223,1248]" pageId="17" pageNumber="544">FIGURES 61–62.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[346,672,1224,1248]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="544" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[346,473,1224,1248]" italics="true" pageId="17" pageNumber="544">Teloganodes</emphasis>
(
<emphasis box="[488,602,1224,1248]" italics="true" pageId="17" pageNumber="544">Dudgeodes</emphasis>
)
<emphasis box="[617,672,1225,1248]" italics="true" pageId="17" pageNumber="544">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[678,735,1224,1248]" pageId="17" pageNumber="544">
<taxonomicNameLabel box="[678,735,1224,1248]" pageId="17" pageNumber="544" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
61, male larva of last instar; 62, younger female larva.
</paragraph>
</caption>
<paragraph blockId="17.[151,1437,1288,2035]" box="[199,348,1288,1314]" pageId="17" pageNumber="544">
<emphasis bold="true" box="[199,348,1288,1314]" pageId="17" pageNumber="544">Descriptions</emphasis>
</paragraph>
<paragraph blockId="17.[151,1437,1288,2035]" pageId="17" pageNumber="544">
<emphasis bold="true" box="[199,275,1325,1351]" pageId="17" pageNumber="544">
<emphasis bold="true" box="[199,267,1325,1351]" italics="true" pageId="17" pageNumber="544">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head, pronotum and mesonotum ochre with diffusive brown maculae (
<figureCitation box="[159,292,1361,1387]" captionStart="FIGURES 69–74" captionStartId="19.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-15@19.[151,1436,181,1817]" captionTargetPageId="19" captionText="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." figureDoi="http://doi.org/10.5281/zenodo.7673067" httpUri="https://zenodo.org/record/7673067/files/figure.png" pageId="17" pageNumber="544">Figs 69–71</figureCitation>
). Femur of fore leg mostly ochre; femora of middle and hind legs mostly ochre, with proximal part diffusively darkened with brown; tibia of each leg diffusively darkened with brown in midlength-outer area (
<figureCitation captionStart="FIGURES 75–83" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1418,181,1821]" captionTargetId="figure-23@20.[151,1436,181,1821]" captionTargetPageId="20" captionText="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi="http://doi.org/10.5281/zenodo.7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="17" pageNumber="544">Figs 76–78</figureCitation>
). Abdominal terga with light brownish, with lateral areas of terga III–
<collectionCode box="[1005,1034,1433,1459]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="544">VI</collectionCode>
lighter (
<figureCitation box="[1130,1214,1433,1459]" captionStart="FIGURES 69–74" captionStartId="19.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-15@19.[151,1436,181,1817]" captionTargetPageId="19" captionText="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." figureDoi="http://doi.org/10.5281/zenodo.7673067" httpUri="https://zenodo.org/record/7673067/files/figure.png" pageId="17" pageNumber="544">Fig. 72</figureCitation>
).
</paragraph>
<paragraph blockId="17.[151,1437,1288,2035]" pageId="17" pageNumber="544">
HYPODERMAL COLORATION. Head and thorax with brown maculae or mostly brown. Fore protoptera with brown macula at base. On each leg, femur with black-brown apex, tibia with black-brown base; middle and hind femora with or without dark longitudinal maculae on proximal part; tibia of each leg with or without dark longitudinal stripe near middle (
<figureCitation box="[523,610,1577,1603]" captionStart="FIGURES 61–62" captionStartId="17.[151,264,1223,1247]" captionTargetBox="[159,1419,181,1200]" captionTargetId="figure-15@17.[159,1428,181,1200]" captionTargetPageId="17" captionText="FIGURES 61–62. Teloganodes (Dudgeodes) rasus sp. n. 61, male larva of last instar; 62, younger female larva." figureDoi="http://doi.org/10.5281/zenodo.7673063" httpUri="https://zenodo.org/record/7673063/files/figure.png" pageId="17" pageNumber="544">Fig. 62</figureCitation>
). Abdominal terga I–II mostly brown, terga III–
<collectionCode box="[1166,1195,1577,1603]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="544">VI</collectionCode>
with variable brown marking; terga VII–IX mostly brown (
<figureCitation box="[580,711,1613,1639]" captionStart="FIGURES 61–62" captionStartId="17.[151,264,1223,1247]" captionTargetBox="[159,1419,181,1200]" captionTargetId="figure-15@17.[159,1428,181,1200]" captionTargetPageId="17" captionText="FIGURES 61–62. Teloganodes (Dudgeodes) rasus sp. n. 61, male larva of last instar; 62, younger female larva." figureDoi="http://doi.org/10.5281/zenodo.7673063" httpUri="https://zenodo.org/record/7673063/files/figure.png" pageId="17" pageNumber="544">Figs 61–62</figureCitation>
).
</paragraph>
<paragraph blockId="17.[151,1437,1288,2035]" pageId="17" pageNumber="544">
<collectionCode box="[199,503,1649,1675]" pageId="17" pageNumber="544">SHAPE AND SETATION</collectionCode>
. Labrum with simple setae forming transverse field all over labrum width, with feathered setae located distad of this field (as in
<figureCitation box="[685,785,1685,1711]" captionStart="FIGURES 2–7" captionStartId="5.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-15@5.[151,1436,181,1817]" captionTargetPageId="5" captionText="FIGURES 2–7. Details of Teloganodes s.l. 2–4, Teloganodes (Derlethina) eloisae, mature larvae (Malaysia, Sabah, Kindssang, Ranau, Negeri, 9.IX.1994, coll. S.C. Kang): 2, apical part of fore protopteron; 3, hind protopteron; 4, hind leg (si, outer margin of subimaginal femur developing inside). 5, egg of T. (Teloganodes) insignis (Sri Lanka, south-eastern slope of Sri Pada, river Battulu oya, Fishing Hut, 10.II.2020, coll. N. Kluge &amp; L. Sheyko). 6–7, T. (Dudgeodes) celebensis, dorsal cuticle from labrum: 6, focus on proximal part; 7, focus on distal part." figureDoi="http://doi.org/10.5281/zenodo.7673041" httpUri="https://zenodo.org/record/7673041/files/figure.png" pageId="17" pageNumber="544">Figs 6–7</figureCitation>
). Nearly all setae forming rows on lateral margins of head, pronotum, mesonotum, on femora and tibiae of legs, on posterior and lateral margins of abdominal terga, equally short and stout (
<figureCitation box="[338,601,1757,1783]" captionStart-0="FIGURES 69–74" captionStart-1="FIGURES 75–83" captionStartId-0="19.[151,264,1841,1865]" captionStartId-1="20.[151,264,1845,1869]" captionTargetBox-0="[151,1436,181,1817]" captionTargetBox-1="[151,1418,181,1821]" captionTargetId-0="figure-15@19.[151,1436,181,1817]" captionTargetId-1="figure-23@20.[151,1436,181,1821]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." captionText-1="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi-0="http://doi.org/10.5281/zenodo.7673067" figureDoi-1="http://doi.org/10.5281/zenodo.7673069" httpUri-0="https://zenodo.org/record/7673067/files/figure.png" httpUri-1="https://zenodo.org/record/7673069/files/figure.png" pageId="17" pageNumber="544">Figs 70–72, 74–80, 82</figureCitation>
); only few setae longer: few setae on lateral parts of posterior margin of abdominal tergum I more or less elongate and bifid (
<figureCitation box="[738,821,1793,1819]" captionStart="FIGURES 69–74" captionStartId="19.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-15@19.[151,1436,181,1817]" captionTargetPageId="19" captionText="FIGURES 69–74. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII." figureDoi="http://doi.org/10.5281/zenodo.7673067" httpUri="https://zenodo.org/record/7673067/files/figure.png" pageId="17" pageNumber="544">Fig. 73</figureCitation>
); setae forming regular row on dorsal side of hind tibia along its inner margin, either all short and stout (as setae of other rows), or some of them long and thin (
<figureCitation box="[1339,1422,1829,1855]" captionStart="FIGURES 75–83" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1418,181,1821]" captionTargetId="figure-23@20.[151,1436,181,1821]" captionTargetPageId="20" captionText="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi="http://doi.org/10.5281/zenodo.7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="17" pageNumber="544">Fig. 81</figureCitation>
), or most of them long and thin. Outer side of each femur with two parallel rows of short, stout setae (
<figureCitation box="[1276,1361,1865,1891]" captionStart="FIGURES 75–83" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1418,181,1821]" captionTargetId="figure-23@20.[151,1436,181,1821]" captionTargetPageId="20" captionText="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi="http://doi.org/10.5281/zenodo.7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="17" pageNumber="544">Fig. 79</figureCitation>
). Each claw with row of 2–4 (usually 4) denticles on inner margin, without distal denticles, with 2 subapical longitudinal rows of arched setae (by one on anterior and posterior sides) (
<figureCitation box="[864,949,1937,1963]" captionStart="FIGURES 75–83" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1418,181,1821]" captionTargetId="figure-23@20.[151,1436,181,1821]" captionTargetPageId="20" captionText="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi="http://doi.org/10.5281/zenodo.7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="17" pageNumber="544">Fig. 83</figureCitation>
). Tergalii II–IV with two-branched ventral lobe, tergalius
<collectionCode box="[317,337,1973,1999]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="544" type="Museum">V</collectionCode>
without ventral lobe; dorsal lamella of each tergalius III–IV deeply cleft (
<figureCitation box="[1171,1302,1972,1999]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="17" pageNumber="544">Figs 54–57</figureCitation>
; tergalius
<collectionCode box="[1416,1436,1973,1999]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="544" type="Museum">V</collectionCode>
either without cleft (
<figureCitation box="[379,509,2008,2035]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="17" pageNumber="544">Figs 59–60</figureCitation>
), or with cleft much smaller than on previous tergalii (
<figureCitation box="[1119,1250,2008,2035]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="17" pageNumber="544">Figs 57–58</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673065" ID-Zenodo-Dep="7673065" httpUri="https://zenodo.org/record/7673065/files/figure.png" pageId="18" pageNumber="545" startId="18.[151,264,1827,1851]" targetBox="[151,1436,181,1797]" targetPageId="18">
<paragraph blockId="18.[151,1436,1827,1887]" pageId="18" pageNumber="545">
<emphasis bold="true" box="[151,340,1827,1851]" pageId="18" pageNumber="545">FIGURES 63–68.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[346,671,1827,1851]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="545" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[346,473,1827,1851]" italics="true" pageId="18" pageNumber="545">Teloganodes</emphasis>
(
<emphasis box="[488,602,1827,1851]" italics="true" pageId="18" pageNumber="545">Dudgeodes</emphasis>
)
<emphasis box="[616,671,1828,1851]" italics="true" pageId="18" pageNumber="545">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[677,734,1827,1851]" pageId="18" pageNumber="545">
<taxonomicNameLabel box="[677,734,1827,1851]" pageId="18" pageNumber="545" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
63–64, left and right mandibles; 65, maxilla, 66, apex of maxilla; 67, hypopharynx and superlingua; 68, labium. Abbreviations: in, incisor; kd, kinetodontium; m, mola; prs, prostheca.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673067" ID-Zenodo-Dep="7673067" httpUri="https://zenodo.org/record/7673067/files/figure.png" pageId="19" pageNumber="546" startId="19.[151,264,1841,1865]" targetBox="[151,1436,181,1817]" targetPageId="19">
<paragraph blockId="19.[151,1437,1841,1901]" pageId="19" pageNumber="546">
<emphasis bold="true" box="[151,340,1841,1865]" pageId="19" pageNumber="546">FIGURES 69–74.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[345,670,1841,1865]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="546" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[345,472,1841,1865]" italics="true" pageId="19" pageNumber="546">Teloganodes</emphasis>
(
<emphasis box="[487,601,1841,1865]" italics="true" pageId="19" pageNumber="546">Dudgeodes</emphasis>
)
<emphasis box="[615,670,1842,1865]" italics="true" pageId="19" pageNumber="546">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[675,732,1841,1865]" pageId="19" pageNumber="546">
<taxonomicNameLabel box="[675,732,1841,1865]" pageId="19" pageNumber="546" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, larval exuviae (holotype). 69–70, parts of head; 71, half of pronotum and mesonotum; 72, abdomen; 73, fragment of abdominal segment I; 74, fragment of abdominal segment VIII.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673069" ID-Zenodo-Dep="7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="20" pageNumber="547" startId="20.[151,264,1845,1869]" targetBox="[151,1418,181,1821]" targetPageId="20">
<paragraph blockId="20.[151,1437,1845,1905]" pageId="20" pageNumber="547">
<emphasis bold="true" box="[151,341,1845,1869]" pageId="20" pageNumber="547">FIGURES 75–83.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[348,675,1845,1870]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="547" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[348,475,1845,1869]" italics="true" pageId="20" pageNumber="547">Teloganodes</emphasis>
(
<emphasis box="[491,605,1845,1869]" italics="true" pageId="20" pageNumber="547">Dudgeodes</emphasis>
)
<emphasis box="[620,675,1846,1869]" italics="true" pageId="20" pageNumber="547">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[682,740,1845,1869]" pageId="20" pageNumber="547">
<taxonomicNameLabel box="[682,740,1845,1869]" pageId="20" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673071" ID-Zenodo-Dep="7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="21" pageNumber="548" startId="21.[151,264,1841,1865]" targetBox="[151,1431,181,1808]" targetPageId="21">
<paragraph blockId="21.[151,1437,1841,1938]" pageId="21" pageNumber="548">
<emphasis bold="true" box="[151,343,1841,1865]" pageId="21" pageNumber="548">FIGURES 84–92.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[351,682,1841,1866]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="548" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[351,478,1841,1865]" italics="true" pageId="21" pageNumber="548">Teloganodes</emphasis>
(
<emphasis box="[496,610,1841,1865]" italics="true" pageId="21" pageNumber="548">Dudgeodes</emphasis>
)
<emphasis box="[627,682,1842,1865]" italics="true" pageId="21" pageNumber="548">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[691,751,1842,1866]" pageId="21" pageNumber="548">
<taxonomicNameLabel box="[691,751,1842,1866]" pageId="21" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673077" ID-Zenodo-Dep="7673077" httpUri="https://zenodo.org/record/7673077/files/figure.png" pageId="22" pageNumber="549" startId="22.[151,264,1218,1242]" targetBox="[151,1435,186,1189]" targetPageId="22">
<paragraph blockId="22.[151,1436,1218,1278]" pageId="22" pageNumber="549">
<emphasis bold="true" box="[151,339,1218,1242]" pageId="22" pageNumber="549">FIGURES 93–94.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[344,667,1218,1242]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="549" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[344,471,1218,1242]" italics="true" pageId="22" pageNumber="549">Teloganodes</emphasis>
(
<emphasis box="[484,598,1218,1242]" italics="true" pageId="22" pageNumber="549">Dudgeodes</emphasis>
)
<emphasis box="[612,667,1219,1242]" italics="true" pageId="22" pageNumber="549">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[671,727,1218,1242]" pageId="22" pageNumber="549">
<taxonomicNameLabel box="[671,727,1218,1242]" pageId="22" pageNumber="549" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, wings of male imago (holotype). 93, in falling light; 94, in transmitted light (arrow shows constant crossvein
<emphasis box="[540,622,1255,1278]" italics="true" pageId="22" pageNumber="549">cua-cup</emphasis>
).
</paragraph>
</caption>
<paragraph blockId="22.[151,1437,1323,2035]" pageId="22" pageNumber="549">
<emphasis bold="true" box="[199,323,1323,1350]" italics="true" pageId="22" pageNumber="549">Subimago.</emphasis>
CUTICULAR COLORATION. Pronotum light brown. Mesonotum with anterior scutal chromozone and latero-posterior scutal chromozones light brown, medioparapsidal suture lighter; achromozone contrastingly colorless (
<figureCitation box="[266,398,1396,1422]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="22" pageNumber="549">Figs 98–99</figureCitation>
). Thoracic pleura and sterna with light brown and colorless areas. Wings brown. Fore femur with longitudinal dark brown and colorless bands; middle and hind femora colorless with light brown band along outer margin; fore tibia darker than tarsus, darkened with brown basally; middle and hind tibiae lighter, darkened with light brownish near base (
<figureCitation box="[511,645,1504,1530]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Figs 89–90</figureCitation>
). Abdominal terga and sterna light brown, with contrasting colorless blanks (
<figureCitation box="[240,323,1540,1567]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 85</figureCitation>
).
</paragraph>
<paragraph blockId="22.[151,1437,1323,2035]" box="[199,754,1576,1603]" pageId="22" pageNumber="549">HYPODERMAL COLORATION. As in imago.</paragraph>
<paragraph blockId="22.[151,1437,1323,2035]" pageId="22" pageNumber="549">TEXTURE: On fore leg of male, 1st tarsomere (fused with tibia) covered with microtrichia (as tibia), 2nd–5th tarsomeres covered with blunt microlepides; on middle and hind legs 1st–4th tarsomeres covered with microtrichia (as tibia), 5th tarsomere covered with blunt microlepides.</paragraph>
<paragraph blockId="22.[151,1437,1323,2035]" lastBlockId="23.[151,1436,151,249]" lastPageId="23" lastPageNumber="550" pageId="22" pageNumber="549">
<emphasis bold="true" box="[199,350,1719,1746]" italics="true" pageId="22" pageNumber="549">Imago, male.</emphasis>
Head brown. Dorsal eyes gray (
<figureCitation box="[706,790,1720,1746]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 84</figureCitation>
). Thorax brown (
<figureCitation box="[980,1064,1720,1746]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 84</figureCitation>
). Fore wing with most membrane colorless, area proximad of costal brace brown; longitudinal veins colorless or ochre; cross veins colorless; crossveins of pterostigma simple (
<figureCitation box="[412,544,1792,1818]" captionStart="FIGURES 93–94" captionStartId="22.[151,264,1218,1242]" captionTargetBox="[151,1435,186,1189]" captionTargetId="figure-23@22.[151,1436,181,1193]" captionTargetPageId="22" captionText="FIGURES 93–94. Teloganodes (Dudgeodes) rasus sp. n., wings of male imago (holotype). 93, in falling light; 94, in transmitted light (arrow shows constant crossvein cua-cup)." figureDoi="http://doi.org/10.5281/zenodo.7673077" httpUri="https://zenodo.org/record/7673077/files/figure.png" pageId="22" pageNumber="549">Figs 93–94</figureCitation>
). Hind wing widened distally, with costal projection in distal part; basal part of membrane and veins dark brown, other membrane and veins colorless (
<figureCitation box="[969,1130,1828,1854]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="22" pageNumber="549">Figs 100–101</figureCitation>
). On foreleg, femur brown with apex black-brown; tibia mostly ochre, with black-brown base and apex; tarsus mostly ochre, with black-dark brown apex of each tarsomere II–
<collectionCode box="[528,548,1901,1927]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="549" type="Museum">V</collectionCode>
(
<figureCitation box="[564,647,1900,1926]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 91</figureCitation>
). On middle and hind legs, femur mostly ochre with apex brown; tibia mostly ochre, with black-brown base and apex; tarsus mostly ochre, with black-brown apex (
<figureCitation box="[1206,1291,1936,1962]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 92</figureCitation>
). Abdominal terga I–II mostly brown, terga III–IV mostly ochre with reddish, medially-anteriorly brown; terga
<collectionCode box="[1286,1306,1973,1999]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="549" type="Museum">V</collectionCode>
–
<collectionCode box="[1321,1350,1973,1999]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="22" pageNumber="549">VI</collectionCode>
mostly ochre with reddish; terga VII–IX mostly brown; sterna lighter (
<figureCitation box="[845,928,2008,2034]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="22" pageNumber="549">Fig. 86</figureCitation>
). Styliger light ochre-brownish, gonostyli and penis ochre (
<figureCitation box="[293,376,151,177]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="23" pageNumber="550">Fig. 97</figureCitation>
). Penis longer than 1st segment of gonostylus, widest near middle, with apices pressed together, with common unpaired gonopore exposed dorsally (
<figureCitation box="[737,869,186,213]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="23" pageNumber="550">Figs 95–96</figureCitation>
). Cerci ochre beginning from very base, with dark brown apex of each segment (
<figureCitation box="[486,618,223,249]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="23" pageNumber="550">Figs 87–88</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673083" ID-Zenodo-Dep="7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="23" pageNumber="550" startId="23.[151,264,1926,1950]" targetBox="[151,1436,273,1903]" targetPageId="23">
<paragraph blockId="23.[151,1436,1926,2022]" pageId="23" pageNumber="550">
<emphasis bold="true" box="[151,354,1926,1950]" pageId="23" pageNumber="550">FIGURES 95–101.</emphasis>
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[362,690,1926,1950]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="550" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[362,489,1926,1950]" italics="true" pageId="23" pageNumber="550">Teloganodes</emphasis>
(
<emphasis box="[505,619,1926,1950]" italics="true" pageId="23" pageNumber="550">Dudgeodes</emphasis>
)
<emphasis box="[635,690,1927,1950]" italics="true" pageId="23" pageNumber="550">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[698,756,1926,1950]" pageId="23" pageNumber="550">
<taxonomicNameLabel box="[698,756,1926,1950]" pageId="23" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light.
</paragraph>
</caption>
<paragraph blockId="24.[151,1437,150,465]" box="[199,498,150,177]" pageId="24" pageNumber="551">
<emphasis bold="true" box="[199,373,150,177]" italics="true" pageId="24" pageNumber="551">Imago, female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="24.[151,1437,150,465]" box="[199,378,187,213]" pageId="24" pageNumber="551">
<emphasis bold="true" box="[199,252,187,213]" italics="true" pageId="24" pageNumber="551">Egg.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="24.[151,1437,150,465]" box="[199,625,222,249]" pageId="24" pageNumber="551">
<emphasis bold="true" box="[199,334,222,248]" pageId="24" pageNumber="551">Dimension.</emphasis>
Forewing length
<quantity box="[532,621,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="24" pageNumber="551" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="551" type="distribution">
<paragraph blockId="24.[151,1437,150,465]" box="[199,468,258,285]" pageId="24" pageNumber="551">
<emphasis bold="true" box="[199,353,258,284]" pageId="24" pageNumber="551">Distribution.</emphasis>
<collectingRegion box="[359,464,259,285]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="551">Sulawesi</collectingRegion>
.
</paragraph>
<paragraph blockId="24.[151,1437,150,465]" pageId="24" pageNumber="551">
<emphasis bold="true" box="[199,354,294,320]" pageId="24" pageNumber="551">Comparison.</emphasis>
Larva of the new species
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[646,787,295,321]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[646,667,295,320]" italics="true" pageId="24" pageNumber="551">T.</emphasis>
(
<emphasis box="[682,710,295,320]" italics="true" pageId="24" pageNumber="551">D.</emphasis>
)
<emphasis box="[726,787,295,320]" italics="true" pageId="24" pageNumber="551">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[793,857,295,321]" pageId="24" pageNumber="551">
<taxonomicNameLabel box="[793,857,295,321]" pageId="24" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from all other species of
<taxonomicName authorityName=": Sartori, Peters &amp; Hubbard" authorityYear="2008" box="[1221,1362,295,321]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis box="[1221,1362,295,321]" italics="true" pageId="24" pageNumber="551">Teloganodes</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1369,1402,295,321]" pageId="24" pageNumber="551" sensu="lato">s.l.</taxonomicNameLabel>
by very short stout setae in all setal rows of femora and tibiae (
<figureCitation box="[806,936,330,357]" captionStart="FIGURES 75–83" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1418,181,1821]" captionTargetId="figure-23@20.[151,1436,181,1821]" captionTargetPageId="20" captionText="FIGURES 75–83. Teloganodes (Dudgeodes) rasus sp. n., larval exuviae of legs (holotype). 75–76, fore leg and its fragment; 77, middle leg; 78–82, hind leg and its fragments; 83, claw." figureDoi="http://doi.org/10.5281/zenodo.7673069" httpUri="https://zenodo.org/record/7673069/files/figure.png" pageId="24" pageNumber="551">Figs 75–82</figureCitation>
). Male imago of
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[1122,1261,331,357]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[1122,1143,331,356]" italics="true" pageId="24" pageNumber="551">T.</emphasis>
(
<emphasis box="[1157,1185,331,356]" italics="true" pageId="24" pageNumber="551">D.</emphasis>
)
<emphasis box="[1200,1261,331,356]" italics="true" pageId="24" pageNumber="551">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1266,1329,331,357]" pageId="24" pageNumber="551">
<taxonomicNameLabel box="[1266,1329,331,357]" pageId="24" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to the sympatric species
<taxonomicName box="[435,635,367,393]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="species" species="celebensis" subGenus="Dudgeodes">
<emphasis box="[435,456,367,392]" italics="true" pageId="24" pageNumber="551">T.</emphasis>
(
<emphasis box="[474,502,367,392]" italics="true" pageId="24" pageNumber="551">D.</emphasis>
)
<emphasis box="[520,635,367,393]" italics="true" pageId="24" pageNumber="551">celebensis</emphasis>
</taxonomicName>
in wings and legs coloration, shape of hind wing and shape of penis; differs by coloration of abdomen, which in
<taxonomicName authority="Kluge, 2023" authorityName="Kluge" authorityYear="2023" box="[639,780,403,429]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="species" species="rasus" status="sp. n." subGenus="Dudgeodes">
<emphasis box="[639,660,403,428]" italics="true" pageId="24" pageNumber="551">T.</emphasis>
(
<emphasis box="[676,703,403,428]" italics="true" pageId="24" pageNumber="551">D.</emphasis>
)
<emphasis box="[719,780,403,428]" italics="true" pageId="24" pageNumber="551">rasus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[787,851,403,429]" pageId="24" pageNumber="551">
<taxonomicNameLabel box="[787,851,403,429]" pageId="24" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has terga III–VI lighter than terga VII–IX (
<figureCitation box="[1339,1423,403,429]" captionStart="FIGURES 84–92" captionStartId="21.[151,264,1841,1865]" captionTargetBox="[151,1431,181,1808]" captionTargetId="figure-15@21.[151,1436,181,1817]" captionTargetPageId="21" captionText="FIGURES 84–92. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 84, head and thorax of imago; 85, subimaginal exuviae of abdomen; 86, imaginal abdomen; 87–88, two portions of imaginal cercus; 89–90, subimaginal exuviae of fore and hind legs; 91–92, imaginal fore and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7673071" httpUri="https://zenodo.org/record/7673071/files/figure.png" pageId="24" pageNumber="551">Fig. 86</figureCitation>
), in contrast to uniformly dark terga I–X in
<taxonomicName box="[622,817,439,465]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="551" phylum="Arthropoda" rank="species" species="celebensis" subGenus="Dudgeodes">
<emphasis box="[622,643,439,464]" italics="true" pageId="24" pageNumber="551">T.</emphasis>
(
<emphasis box="[659,686,439,464]" italics="true" pageId="24" pageNumber="551">D.</emphasis>
)
<emphasis box="[702,817,439,465]" italics="true" pageId="24" pageNumber="551">celebensis</emphasis>
</taxonomicName>
(
<figureCitation box="[833,917,439,465]" captionStart="FIGURES 31–39" captionStartId="13.[151,264,1840,1864]" captionTargetBox="[150,1438,181,1817]" captionTargetId="figure-15@13.[150,1438,181,1817]" captionTargetPageId="13" captionText="FIGURES 31–39. Teloganodes (Dudgeodes) celebensis. 31, male imago; 32, abdomen of male imago; 33, head and thorax of female imago; 34, fore leg of female imago; 35–36, fore legs of male imagines; 37, middle leg of male imago; 38–39, exuviae of fore and middle legs of male subimago (32–39 with equal magnification)." figureDoi="http://doi.org/10.5281/zenodo.7673055" httpUri="https://zenodo.org/record/7673055/files/figure.png" pageId="24" pageNumber="551">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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