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<document ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A8001077E0FF14FF14A3B292F2FB8A" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5ce3343f" docStyleName="zootaxa.2013.monograph" docTitle="Epipolasis tubulata Van, 2017, sp. nov." docType="treatment" docVersion="19" lastPageNumber="185" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="184" updateTime="1645784257244" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Sponges of the Guyana Shelf</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Van, Rob W. M.</mods:namePart>
</mods:name>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2017</mods:date>
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<mods:number>1</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.272951</mods:identifier>
<mods:identifier type="GBIF-Dataset">e2c88f4c-3ac2-45f9-95e4-99b75561a081</mods:identifier>
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<mods:identifier type="ZooBank">6D68A019-6F63-4AA4-A8B3-92D351F1F69B</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5698750" ID-GBIF-Taxon="127419806" ID-Zenodo-Dep="5698750" LSID="urn:lsid:plazi:treatment:03A8001077E0FF14FF14A3B292F2FB8A" httpUri="http://treatment.plazi.org/id/03A8001077E0FF14FF14A3B292F2FB8A" lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184">
<subSubSection pageId="183" pageNumber="184" type="nomenclature">
<paragraph blockId="183.[151,485,217,277]" box="[151,485,217,243]" pageId="183" pageNumber="184">
<heading bold="true" box="[151,485,217,243]" fontSize="11" level="1" pageId="183" pageNumber="184" reason="1">
<emphasis bold="true" box="[151,485,217,243]" pageId="183" pageNumber="184">
<taxonomicName box="[151,385,217,243]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="183" pageNumber="184" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
<emphasis bold="true" box="[151,385,217,243]" italics="true" pageId="183" pageNumber="184">Epipolasis tubulata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[391,485,217,243]" pageId="183" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="183.[151,485,217,277]" box="[151,322,252,277]" pageId="183" pageNumber="184">
<figureCitation box="[151,282,252,277]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="183" pageNumber="184">Figures 115</figureCitation>
ag
</paragraph>
</subSubSection>
<subSubSection lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184" type="materials_examined">
<paragraph blockId="183.[151,1436,318,379]" pageId="183" pageNumber="184">
<materialsCitation ID-GBIF-Occurrence="1424726530" collectingDate="1966-05-10" collectionCode="RMNH" collectorName="Snellius O. C. P. S." country="Suriname" elevation="68" latitude="7.26" location="Suriname" longLatPrecision="555" longitude="-56.6667" pageId="183" pageNumber="184" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[151,386,318,343]" pageId="183" pageNumber="184">Material examined.</emphasis>
<typeStatus box="[396,500,318,343]" pageId="183" pageNumber="184">Holotype</typeStatus>
<collectionCode box="[509,593,319,342]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="183" pageNumber="184">RMNH</collectionCode>
Por. 9965 (larger specimen),
<collectingCountry box="[935,1045,318,343]" name="Suriname" pageId="183" pageNumber="184">Suriname</collectingCountry>
,
<collectorName box="[1063,1266,318,343]" pageId="183" pageNumber="184">Snellius O.C.P.S.</collectorName>
<collectingCountry box="[1281,1368,319,343]" name="Guyana" pageId="183" pageNumber="184">Guyana</collectingCountry>
Shelf Expedition, station G56,
<geoCoordinate box="[432,512,354,379]" direction="north" orientation="latitude" pageId="183" pageNumber="184" precision="555" value="7.26">7.26°N</geoCoordinate>
<geoCoordinate box="[519,648,354,379]" direction="west" orientation="longitude" pageId="183" pageNumber="184" precision="5" value="-56.6667">56.6667°W</geoCoordinate>
, depth
<quantity box="[728,829,354,379]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="183" pageNumber="184" unit="m" value="67.5" valueMax="68.0" valueMin="67.0">
<elevation box="[728,829,354,379]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="183" pageNumber="184" unit="m" value="67.5" valueMax="68.0" valueMin="67.0">6768 m</elevation>
</quantity>
, Agassiz trawl,
<date box="[1008,1161,354,379]" pageId="183" pageNumber="184" value="1966-05-10">
<collectingDate box="[1008,1161,354,379]" pageId="183" pageNumber="184" value="1966-05-10">10 May 1966</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="183" pageNumber="184" targetBox="[151,1436,423,1886]" targetPageId="183">
<paragraph blockId="183.[151,1437,1908,2022]" pageId="183" pageNumber="184">
<emphasis bold="true" box="[151,302,1908,1930]" pageId="183" pageNumber="184">FIGURE 115.</emphasis>
<taxonomicName box="[311,512,1908,1930]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="183" pageNumber="184" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
<emphasis box="[311,512,1908,1930]" italics="true" pageId="183" pageNumber="184">
<taxonomicName box="[311,418,1908,1930]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="183" pageNumber="184" phylum="Porifera" rank="genus">Epipolasis</taxonomicName>
<taxonomicName box="[428,512,1908,1930]" pageId="183" pageNumber="184" rank="species" species="tubulata" status="sp. nov.">tubulata</taxonomicName>
</emphasis>
</taxonomicName>
<emphasis bold="true" box="[521,605,1909,1930]" pageId="183" pageNumber="184">
<taxonomicNameLabel box="[521,605,1909,1930]" pageId="183" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, a, habitus of holotype RMNH Por. 9965, a1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, dg, SEM images of the spicules, d, large oxeote style, d1, details of d, e, small oxeote style, e1, details of e, f, trichodragma, g, individual raphide, g1, detail of hooked end of g.
</paragraph>
</caption>
<paragraph blockId="184.[151,1437,151,1040]" box="[199,1005,151,176]" pageId="184" pageNumber="185">
<materialsCitation ID-GBIF-Occurrence="1424726531" box="[199,1005,151,176]" collectingDate="1966-05-10" collectionCode="RMNH" collectorName="Snellius O. C. P. S." country="Suriname" elevation="68" latitude="7.26" location="Suriname" longLatPrecision="555" longitude="-56.6667" pageId="184" pageNumber="185" specimenCount="1" typeStatus="paratype">
<typeStatus box="[199,297,152,176]" pageId="184" pageNumber="185">Paratype</typeStatus>
<collectionCode box="[304,388,152,175]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="184" pageNumber="185">RMNH</collectionCode>
Por. 10549 (smaller specimen), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="184" pageNumber="185" type="description">
<paragraph blockId="184.[151,1437,151,1040]" pageId="184" pageNumber="185">
<emphasis bold="true" box="[199,345,188,213]" pageId="184" pageNumber="185">Description.</emphasis>
(
<figureCitation box="[358,452,188,213]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Fig. 115</figureCitation>
a) Sponge forms low thick-walled tubes, diameter of the larger tube
<quantity box="[1209,1287,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="184" pageNumber="185" unit="cm" value="1.5">1.5 cm</quantity>
, height
<quantity box="[1374,1432,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="184" pageNumber="185" unit="cm" value="2.0">2 cm</quantity>
. Largest tube opening
<quantity box="[400,467,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="184" pageNumber="185" unit="mm" value="8.0">8 mm</quantity>
in diameter. Surface smooth, but irregular around the oscules. Color greyish brown. Consistency compressible. The smaller
<typeStatus box="[597,693,261,285]" pageId="184" pageNumber="185">paratype</typeStatus>
(
<figureCitation box="[708,805,260,285]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Fig. 115</figureCitation>
a1) is similar.
</paragraph>
<paragraph blockId="184.[151,1437,151,1040]" pageId="184" pageNumber="185">
<emphasis bold="true" box="[199,309,295,320]" pageId="184" pageNumber="185">Skeleton.</emphasis>
The surface (
<figureCitation box="[463,558,295,320]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Fig. 115</figureCitation>
b) is a halichondrioid tangential crust of intercrossing spicule tracts and single spicules, 210 spicules in diameter. This is carried by choanosomal spicule tracts (
<figureCitation box="[1135,1237,332,357]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Fig. 115</figureCitation>
c), likewise 210 spicules in cross section, surrounded by a largely confused mass of single spicules and numerous single raphides, partially arranged in thin trichodragmas.
</paragraph>
<paragraph blockId="184.[151,1437,151,1040]" box="[199,808,439,464]" pageId="184" pageNumber="185">
<emphasis bold="true" box="[199,304,439,464]" pageId="184" pageNumber="185">Spicules.</emphasis>
(
<figureCitation box="[319,416,439,464]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Figs 115</figureCitation>
dg) Oxeote styles, trichodragmas.
</paragraph>
<paragraph blockId="184.[151,1437,151,1040]" pageId="184" pageNumber="185">
Oxeotes, style-like, smooth, slightly fusiform, usually somewhat curved, with a faint subapical constriction, in two distinct size categories, (1) larger (
<figureCitation box="[615,723,511,536]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Figs 115</figureCitation>
d,d1), 462
<emphasis box="[851,893,511,536]" italics="true" pageId="184" pageNumber="185">601</emphasis>
618
<date box="[960,1013,512,536]" pageId="184" pageNumber="185" value="2012-10">x 12</date>
<emphasis box="[1028,1077,511,536]" italics="true" pageId="184" pageNumber="185">16.3</emphasis>
19 µm, and (2) smaller (
<figureCitation captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Figs 115</figureCitation>
e,e1), 192
<emphasis box="[317,359,548,573]" italics="true" pageId="184" pageNumber="185">286</emphasis>
324
<date box="[421,456,548,573]" pageId="184" pageNumber="185">x 5</date>
<emphasis box="[471,506,548,572]" italics="true" pageId="184" pageNumber="185">7.4</emphasis>
9 µm.
</paragraph>
<paragraph blockId="184.[151,1437,151,1040]" pageId="184" pageNumber="185">
Raphides, partially in thin trichodragmas (
<figureCitation box="[708,810,583,608]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">Fig. 115</figureCitation>
f), but many raphides (
<figureCitation box="[1092,1194,583,608]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">
Fig.
<quantity box="[1152,1194,583,608]" metricMagnitude="-1" metricUnit="kg" metricValue="1.15" pageId="184" pageNumber="185" unit="g" value="115.0">115</quantity>
</figureCitation>
g) are strewn singly throughout the skeleton; provided with a lanceolate hook at both ends (
<figureCitation box="[985,1087,620,645]" captionStart="FIGURE 115" captionStartId="183.[151,250,1908,1930]" captionTargetBox="[151,1436,423,1886]" captionTargetId="figure@183.[151,1436,423,1886]" captionTargetPageId="183" captionText="FIGURE 115. Epipolasis tubulata sp. nov., a, habitus of holotype RMNH Por. 9965, a 1, habitus of paratype RMNH Por. 10549 (scale bar = 1 cm), b, light microscopic image of tangential halichondrioid ectosomal skeleton, c, ditto of cross section of peripheral skeleton, d g, SEM images of the spicules, d, large oxeote style, d 1, details of d, e, small oxeote style, e 1, details of e, f, trichodragma, g, individual raphide, g 1, detail of hooked end of g." httpUri="https://zenodo.org/record/273065/files/figure.png" pageId="184" pageNumber="185">
Fig.
<quantity box="[1041,1087,620,645]" metricMagnitude="-1" metricUnit="kg" metricValue="1.15" pageId="184" pageNumber="185" unit="g" value="115.0">115</quantity>
</figureCitation>
g1), 76
<emphasis box="[1178,1206,620,644]" italics="true" pageId="184" pageNumber="185">87</emphasis>
114 x 0.30.4 µm; trichodragmas up to 20 µm in diameter.
</paragraph>
</subSubSection>
<subSubSection box="[199,875,692,717]" pageId="184" pageNumber="185" type="distribution">
<paragraph blockId="184.[151,1437,151,1040]" box="[199,875,692,717]" pageId="184" pageNumber="185">
<emphasis bold="true" box="[199,498,692,717]" pageId="184" pageNumber="185">Distribution and ecology.</emphasis>
<collectingCountry box="[505,592,692,716]" name="Guyana" pageId="184" pageNumber="185">Guyana</collectingCountry>
Shelf, at
<quantity box="[700,799,692,717]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="184" pageNumber="185" unit="m" value="67.5" valueMax="68.0" valueMin="67.0">6768 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection box="[199,915,727,752]" pageId="184" pageNumber="185" type="etymology">
<paragraph blockId="184.[151,1437,151,1040]" box="[199,915,727,752]" pageId="184" pageNumber="185">
<emphasis bold="true" box="[199,334,727,752]" pageId="184" pageNumber="185">Etymology.</emphasis>
The name refers to the tubular shape of the sponge.
</paragraph>
</subSubSection>
<subSubSection pageId="184" pageNumber="185" type="discussion">
<paragraph blockId="184.[151,1437,151,1040]" pageId="184" pageNumber="185">
<materialsCitation ID-GBIF-Occurrence="1424726533" collectingDate="1600-10-29" location="Central West" pageId="184" pageNumber="185" specimenCount="1" stateProvince="Central West" typeStatus="holotype">
<emphasis bold="true" box="[199,315,764,789]" pageId="184" pageNumber="185">Remarks.</emphasis>
The genus assignment is tentative. At first the species was assigned to the axinellid genus
<taxonomicName box="[151,279,801,824]" class="Demospongiae" family="Axinellidae" genus="Dragmaxia" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="genus">
<emphasis box="[151,279,801,824]" italics="true" pageId="184" pageNumber="185">Dragmaxia</emphasis>
</taxonomicName>
(cf. above), because of the combination of smooth stylote spicules with trichodragmas. However, the clear division into a larger and a smaller category of megascleres added to the tangential halichondrioid ectosomal skeleton, precludes membership of the genus
<taxonomicName box="[664,791,873,896]" class="Demospongiae" family="Axinellidae" genus="Dragmaxia" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="genus">
<emphasis box="[664,791,873,896]" italics="true" pageId="184" pageNumber="185">Dragmaxia</emphasis>
</taxonomicName>
. So far,
<taxonomicName box="[888,1006,871,896]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="genus">
<emphasis box="[888,1006,871,896]" italics="true" pageId="184" pageNumber="185">Epipolasis</emphasis>
</taxonomicName>
has been limited to species with large and small oxeas, although stylote modifications are reported in the
<typeStatus box="[918,966,909,933]" pageId="184" pageNumber="185">type</typeStatus>
species,
<taxonomicName authority="Wilson, 1925" authorityName="Wilson" authorityYear="1925" class="Demospongiae" family="Halichondriidae" genus="Spongosorites" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="suluensis">
<emphasis box="[1071,1340,908,933]" italics="true" pageId="184" pageNumber="185">Spongosorites suluensis</emphasis>
<bibRefCitation author="Wilson" pageId="184" pageNumber="185" refString="Wilson, H. V. (1925) Silicious and horny sponges collected by the U. S. Fisheries Steamer ' Albatross' during the Philippine Expedition, 1907 - 10. In: Contributions to the biology of the Philippine Archipelago and adjacent regions. Bulletin of the United States National Museum, 100 (2, part 4), 273 - 532." type="journal article" year="1925">Wilson, 1925</bibRefCitation>
</taxonomicName>
. There is one other
<taxonomicName box="[437,555,943,968]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="genus">
<emphasis box="[437,555,943,968]" italics="true" pageId="184" pageNumber="185">Epipolasis</emphasis>
</taxonomicName>
in the
<collectingRegion box="[636,781,943,968]" isEnumeration="true" pageId="184" pageNumber="185">Central West</collectingRegion>
Atlantic,
<taxonomicName authority="Diaz, Pomponi &amp; Van Soest, 1993" authorityName="Diaz, Pomponi &amp; Van Soest" authorityYear="1993" box="[895,1432,943,968]" class="Demospongiae" family="Halichondriidae" genus="Epipolasis" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="profunda">
<emphasis box="[895,1029,943,968]" italics="true" pageId="184" pageNumber="185">E. profunda</emphasis>
<bibRefCitation author="Diaz" box="[1038,1432,943,968]" pageId="184" pageNumber="185" refString="Diaz, M. C., Pomponi, S. A. &amp; Van Soest, R. W. M. (1993) A systematic revision of the central West Atlantic Halichondrida (Demospongiae, Porifera). Part III: Description of valid species. In: Uriz, M. - J. &amp; Rutzler, K. (Eds.), Recent Advances in Ecology and Systematics of Sponges. Scientia Marina, 57 (4), 283 - 306." type="journal article" year="1993">Díaz, Pomponi &amp; Van Soest, 1993</bibRefCitation>
</taxonomicName>
, but this has much larger spicules (up to
<date box="[602,715,980,1005]" pageId="184" pageNumber="185" value="1600-10-29">
<collectingDate box="[602,715,980,1005]" pageId="184" pageNumber="185" value="1600-10-29">1600 x 29</collectingDate>
</date>
µm), which are predominantly proper oxeas; trichodragmas are only half as long as those of the new species.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>