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<document id="E5B960C3E2908993A04BE14676B5BCB8" ID-CLB-Dataset="37320" ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A800107728FF2CFF14A55F93D2F849" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tylosigma ostreicola Van, 2017, sp. nov." docType="treatment" docVersion="21" lastPageNumber="129" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="128" updateTime="1698435695899" updateUser="ExternalLinkService">
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<mods:title id="DEAC4FCDCD28B455466AC85A549B01BC">Sponges of the Guyana Shelf</mods:title>
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<mods:namePart id="7606966288353D661F95CEDAEA74CFE5">Van, Rob W. M.</mods:namePart>
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<mods:date id="9001098992FE3037DC5AC02DD1DBCC08">2017</mods:date>
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<subSubSection id="C31BE28D7728FFD3FF14A55F9148F9F5" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="8BBEB1067728FFD3FF14A55F9189F9D4" blockId="127.[151,500,1588,1647]" box="[151,500,1588,1614]" pageId="127" pageNumber="128">
<heading id="D0F6066A7728FFD3FF14A55F9189F9D4" bold="true" box="[151,500,1588,1614]" fontSize="11" level="1" pageId="127" pageNumber="128" reason="1">
<emphasis id="B9756D147728FFD3FF14A55F9189F9D4" bold="true" box="[151,500,1588,1614]" pageId="127" pageNumber="128">
<taxonomicName id="4C01CA857728FFD3FF14A55F91F2F9D4" ID-CoL="7CZTR" box="[151,399,1588,1614]" class="Demospongiae" family="Desmacellidae" genus="Tylosigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Desmacellida" pageId="127" pageNumber="128" phylum="Porifera" rank="species" species="ostreicola" status="sp. nov.">
<emphasis id="B9756D147728FFD3FF14A55F91F2F9D4" bold="true" box="[151,399,1588,1614]" italics="true" pageId="127" pageNumber="128">Tylosigma ostreicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246D06F7728FFD3FE14A55F9189F9D4" box="[407,500,1588,1614]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBEB1067728FFD3FF14A53D9148F9F5" blockId="127.[151,500,1588,1647]" box="[151,309,1622,1647]" pageId="127" pageNumber="128">
<figureCitation id="133AAD837728FFD3FF14A53D9173F9F5" box="[151,270,1622,1647]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="127" pageNumber="128">Figures 79</figureCitation>
ae
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D7728FFD3FF14A5F59320F941" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="8BBEB1067728FFD3FF14A5F59320F941" blockId="127.[151,1436,1694,2010]" pageId="127" pageNumber="128">
<materialsCitation id="3B69BB5B7728FFD3FF14A5F59324F941" ID-GBIF-Occurrence="1424726303" collectingDate="1966-05-07" collectionCode="RMNH" collectorName="Snellius O. C. P. S." country="Suriname" elevation="64" latitude="7.28" location="Suriname" longLatPrecision="555" longitude="-56.7933" pageId="127" pageNumber="128" specimenCount="1" typeStatus="holotype">
<emphasis id="B9756D147728FFD3FF14A5F591FCF92D" bold="true" box="[151,385,1694,1719]" pageId="127" pageNumber="128">Material examined.</emphasis>
<typeStatus id="54BA0FA47728FFD3FE0AA5F5918CF92D" box="[393,497,1694,1719]" pageId="127" pageNumber="128" type="holotype">Holotype</typeStatus>
<collectionCode id="ED1029C37728FFD3FE7AA5F49230F92C" box="[505,589,1695,1718]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="127" pageNumber="128">RMNH</collectionCode>
Por. 9955,
<collectingCountry id="F316F1967728FFD3FD53A5F59343F92D" box="[720,830,1694,1719]" name="Suriname" pageId="127" pageNumber="128">Suriname</collectingCountry>
,
<collectorName id="26F4D4D07728FFD3FCCDA5F59464F92D" box="[846,1049,1694,1719]" pageId="127" pageNumber="128">Snellius O.C.P.S.</collectorName>
<collectingCountry id="F316F1967728FFD3FBA5A5F49400F92D" box="[1062,1149,1695,1719]" name="Guyana" pageId="127" pageNumber="128">Guyana</collectingCountry>
Shelf Expedition, station G7,
<geoCoordinate id="EE35D7C17728FFD3FF44A5A8916AF941" box="[199,279,1731,1755]" direction="north" orientation="latitude" pageId="127" pageNumber="128" precision="555" value="7.28">7.28°N</geoCoordinate>
<geoCoordinate id="EE35D7C17728FFD3FE9DA5A991E2F941" box="[286,415,1730,1755]" direction="west" orientation="longitude" pageId="127" pageNumber="128" precision="5" value="-56.7933">56.7933°W</geoCoordinate>
, depth
<quantity id="4CF91CE37728FFD3FE73A5A99256F940" box="[496,555,1730,1755]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="127" pageNumber="128" unit="m" value="64.0">
<elevation id="002C56357728FFD3FE73A5A99256F940" box="[496,555,1730,1755]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="127" pageNumber="128" unit="m" value="64.0">64 m</elevation>
</quantity>
, bottom sand,
<date id="FFBF97C67728FFD3FD4DA5A89324F941" box="[718,857,1730,1755]" pageId="127" pageNumber="128" value="1966-05-07">
<collectingDate id="EFFB6E2E7728FFD3FD4DA5A89324F941" box="[718,857,1730,1755]" pageId="127" pageNumber="128" value="1966-05-07">7 May 1966</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D7728FF2CFF44A58D949BFA41" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128" type="description">
<paragraph id="8BBEB1067728FFD3FF44A58D92B6F8BC" blockId="127.[151,1436,1694,2010]" pageId="127" pageNumber="128">
<emphasis id="B9756D147728FFD3FF44A58D9124F965" bold="true" box="[199,345,1766,1791]" pageId="127" pageNumber="128">Description.</emphasis>
(
<figureCitation id="133AAD837728FFD3FEE4A58D91C4F965" box="[359,441,1766,1791]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="127" pageNumber="128">Fig. 79</figureCitation>
a) Encrusting eroded rims and holes of a dead oyster. Surface hispid-conulose. Colour in alcohol beige. Size several mm2. Consistency soft.
</paragraph>
<paragraph id="8BBEB1067728FFD3FF44A45A9244F8F4" blockId="127.[151,1436,1694,2010]" pageId="127" pageNumber="128">
<emphasis id="B9756D147728FFD3FF44A45A9148F8D0" bold="true" box="[199,309,1841,1866]" pageId="127" pageNumber="128">Skeleton.</emphasis>
(
<figureCitation id="133AAD837728FFD3FEC0A45A91E9F8D0" box="[323,404,1841,1866]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="127" pageNumber="128">Fig. 79</figureCitation>
b) Single tylostyles penetrating the surface, heads embedded in the substratum. Low spicular density. Tissue crowded with sigmas.
</paragraph>
<paragraph id="8BBEB1067728FFD3FF44A41292DCF808" blockId="127.[151,1436,1694,2010]" box="[199,673,1913,1938]" pageId="127" pageNumber="128">
<emphasis id="B9756D147728FFD3FF44A412914DF808" bold="true" box="[199,304,1913,1938]" pageId="127" pageNumber="128">Spicules.</emphasis>
(
<figureCitation id="133AAD837728FFD3FEBCA41291E9F808" box="[319,404,1913,1938]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="127" pageNumber="128">Figs 79</figureCitation>
ce) Tylostyles, sigmas.
</paragraph>
<paragraph id="8BBEB1067728FFD3FF44A4F69396F840" blockId="127.[151,1436,1694,2010]" pageId="127" pageNumber="128">
Tylostyles (
<figureCitation id="133AAD837728FFD3FECBA4F691E3F82C" box="[328,414,1949,1974]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="127" pageNumber="128">Figs 79</figureCitation>
cd), curved, thin, with prominent round tyles, in a large size range, but not clearly divisible in categories, 342
<emphasis id="B9756D147728FFD3FEE8A4A991E8F840" box="[363,405,1986,2010]" italics="true" pageId="127" pageNumber="128">779</emphasis>
<date id="FFBF97C67728FFD3FE27A4A99279F840" box="[420,516,1985,2010]" pageId="127" pageNumber="128" value="1092-10-06">1092 x 6</date>
<emphasis id="B9756D147728FFD3FD90A4A9924BF840" box="[531,566,1986,2010]" italics="true" pageId="127" pageNumber="128">9.1</emphasis>
13 µm (diameter of tyles 8.515 µm).
</paragraph>
<caption id="DF7EE18E77D7FF2CFF14A62393F1FA01" httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="128" pageNumber="129" targetBox="[151,1436,193,1330]" targetPageId="128">
<paragraph id="8BBEB10677D7FF2CFF14A62393F1FA01" blockId="128.[151,1436,1352,1435]" pageId="128" pageNumber="129">
<emphasis id="B9756D1477D7FF2CFF14A6239162FAC7" bold="true" box="[151,287,1352,1374]" pageId="128" pageNumber="129">FIGURE 79.</emphasis>
<taxonomicName id="4C01CA8577D7FF2CFEA7A623918BFAC4" box="[292,502,1352,1374]" class="Demospongiae" family="Desmacellidae" genus="Tylosigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Desmacellida" pageId="128" pageNumber="129" phylum="Porifera" rank="species" species="ostreicola" status="sp. nov.">
<emphasis id="B9756D1477D7FF2CFEA7A623918BFAC4" box="[292,502,1352,1374]" italics="true" pageId="128" pageNumber="129">Tylosigma ostreicola</emphasis>
</taxonomicName>
<emphasis id="B9756D1477D7FF2CFE7FA6239236FAC7" bold="true" box="[508,587,1352,1373]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A246D06F77D7FF2CFE7FA6239236FAC7" box="[508,587,1352,1373]" pageId="128" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, ce, SEM images of spicules, c, large tylostyle, c1, details of c, d, small tylostyle, d1, details of d, e, sigma.
</paragraph>
</caption>
<paragraph id="8BBEB10677D7FF2CFF44A6A9949BFA41" blockId="128.[151,1436,1474,2003]" box="[199,1254,1474,1499]" pageId="128" pageNumber="129">
Sigmas (
<figureCitation id="133AAD8377D7FF2CFEAAA6A99101FA41" box="[297,380,1474,1499]" captionStart="FIGURE 79" captionStartId="128.[151,250,1352,1374]" captionTargetBox="[151,1436,193,1330]" captionTargetId="figure@128.[151,1436,193,1330]" captionTargetPageId="128" captionText="FIGURE 79. Tylosigma ostreicola sp. nov., a, habitus (arrow) of holotype RMNH Por. 9955 (scale bar = 1 cm), b, cross section of skeleton showing single styles erect on the substratum and scattered sigmas in the tissue, c e, SEM images of spicules, c, large tylostyle, c 1, details of c, d, small tylostyle, d 1, details of d, e, sigma." httpUri="https://zenodo.org/record/273030/files/figure.png" pageId="128" pageNumber="129">Fig. 79</figureCitation>
e), with incurved apices, one of which may be faintly rugose, 16
<emphasis id="B9756D1477D7FF2CFBD4A6A894F5FA41" box="[1111,1160,1475,1499]" italics="true" pageId="128" pageNumber="129">21.8</emphasis>
27 µm.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77D7FF2CFF44A68D938DFA65" box="[199,1008,1510,1535]" pageId="128" pageNumber="129" type="distribution">
<paragraph id="8BBEB10677D7FF2CFF44A68D938DFA65" blockId="128.[151,1436,1474,2003]" box="[199,1008,1510,1535]" pageId="128" pageNumber="129">
<emphasis id="B9756D1477D7FF2CFF44A68D918FFA65" bold="true" box="[199,498,1510,1535]" pageId="128" pageNumber="129">Distribution and ecology.</emphasis>
<collectingCountry id="F316F19677D7FF2CFE7AA68C922DFA65" box="[505,592,1511,1535]" name="Guyana" pageId="128" pageNumber="129">Guyana</collectingCountry>
Shelf, on a dead shell at
<quantity id="4CF91CE377D7FF2CFCE9A68D93DEFA64" box="[874,931,1510,1535]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="128" pageNumber="129" unit="m" value="64.0">64 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77D7FF2CFF44A561924DF9DD" pageId="128" pageNumber="129" type="etymology">
<paragraph id="8BBEB10677D7FF2CFF44A561924DF9DD" blockId="128.[151,1436,1474,2003]" pageId="128" pageNumber="129">
<emphasis id="B9756D1477D7FF2CFF44A5619133F9B9" bold="true" box="[199,334,1546,1571]" pageId="128" pageNumber="129">Etymology.</emphasis>
The name ostrea (L.) means oyster, and suffix cola (Gr.) means living on or in, together the compound name refers to its habitat.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77D7FF2CFF44A53993D2F849" pageId="128" pageNumber="129" type="discussion">
<paragraph id="8BBEB10677D7FF2CFF44A539912EF8D9" blockId="128.[151,1436,1474,2003]" pageId="128" pageNumber="129">
<emphasis id="B9756D1477D7FF2CFF44A5399146F9F1" bold="true" box="[199,315,1618,1643]" pageId="128" pageNumber="129">Remarks.</emphasis>
The new species was compared to a slide of
<taxonomicName id="4C01CA8577D7FF2CFCDAA53994FDF9F1" box="[857,1152,1618,1643]" class="Demospongiae" family="Desmacellidae" genus="Tylosigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Desmacellida" pageId="128" pageNumber="129" phylum="Porifera" rank="species" species="campechianum">
<emphasis id="B9756D1477D7FF2CFCDAA53994FDF9F1" box="[857,1152,1618,1643]" italics="true" pageId="128" pageNumber="129">Tylosigma campechianum</emphasis>
</taxonomicName>
(originally
<taxonomicName id="4C01CA8577D7FF2CFA88A5399151F915" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="128" pageNumber="129" phylum="Porifera" rank="species" species="campechiana">
<emphasis id="B9756D1477D7FF2CFA88A5399151F915" italics="true" pageId="128" pageNumber="129">Hymedesmia campechiana</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EF90CCF777D7FF2CFEE9A51C927AF915" author="Topsent" box="[362,519,1655,1679]" pageId="128" pageNumber="129" refString="Topsent, E. (1889) Quelques spongiaires du Banc de Campeche et de la Pointe- - Pitre. Memoires de la Societ zoologique de France, 2, 30 - 52." type="journal article" year="1889">Topsent 1889</bibRefCitation>
, p. 14, fig. 8c) in the Paris Museum, registered as MNHN D.T. 1844. The new species differs from it by the lack of a separate small category of tylostyles three/four times as short as the larger tylostyles, and a separate category of very small thick sigmas. The short tylostyles are also provided with a few microspines on the head, which is lacking in the present material. These differences appear too great to assume variability and thus I propose here a new species.
<bibRefCitation id="EF90CCF777D7FF2CFD50A46D9382F885" author="Alcolado" box="[723,1023,1798,1823]" pageId="128" pageNumber="129" refString="Alcolado, P. M. &amp; Gotera, G. G. (1986) Nuevas adiciones a la fauna de poriferos de Cuba [New additions to the poriferan fauna of Cuba]. Poeyana, 331, 1 - 19." type="journal article" year="1986">Alcolado &amp; Gotera (1986)</bibRefCitation>
report
<taxonomicName id="4C01CA8577D7FF2CFBD0A46C9564F885" box="[1107,1305,1798,1823]" class="Demospongiae" family="Desmacellidae" genus="Tylosigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Desmacellida" pageId="128" pageNumber="129" phylum="Porifera" rank="species" species="campechianum">
<emphasis id="B9756D1477D7FF2CFBD0A46C9564F885" box="[1107,1305,1798,1823]" italics="true" pageId="128" pageNumber="129">T. campechianum</emphasis>
</taxonomicName>
from
<collectingCountry id="F316F19677D7FF2CFAE2A46D95E1F885" box="[1377,1436,1798,1823]" name="Cuba" pageId="128" pageNumber="129">Cuba</collectingCountry>
(as
<taxonomicName id="4C01CA8577D7FF2CFF3DA441913FF8D9" box="[190,322,1834,1859]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="128" pageNumber="129" phylum="Porifera" rank="genus">
<emphasis id="B9756D1477D7FF2CFF3DA441913FF8D9" box="[190,322,1834,1859]" italics="true" pageId="128" pageNumber="129">Desmacella</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBEB10677D7FF2CFF44A42593D2F849" blockId="128.[151,1436,1474,2003]" pageId="128" pageNumber="129">
The two species are of uncertain affiliation, as the tylostyles remind of the genera
<taxonomicName id="4C01CA8577D7FF2CFBF5A4249501F8FD" box="[1142,1404,1870,1895]" class="Demospongiae" family="Raspailiidae" genus="Eurypon" kingdom="Animalia" order="Poecilosclerida" pageId="128" pageNumber="129" phylum="Porifera" rank="genus">
<emphasis id="B9756D1477D7FF2CFBF5A42494ABF8FC" box="[1142,1238,1871,1894]" italics="true" pageId="128" pageNumber="129">Eurypon</emphasis>
(Raspailiidae)
</taxonomicName>
or
<taxonomicName id="4C01CA8577D7FF2CFF14A4199278F811" box="[151,517,1906,1931]" class="Demospongiae" family="Suberitidae" genus="Prosuberites" kingdom="Animalia" order="Hadromerida" pageId="128" pageNumber="129" phylum="Porifera" rank="genus">
<emphasis id="B9756D1477D7FF2CFF14A4199158F811" box="[151,293,1906,1931]" italics="true" pageId="128" pageNumber="129">Prosuberites</emphasis>
(Hymerhabdiidae)
</taxonomicName>
, but these genera do not have sigmas or other sigmiform microscleres. A combination of tylostyles and sigmas is found in the family
<taxonomicName id="4C01CA8577D7FF2CFCF9A4FD9462F835" box="[890,1055,1942,1967]" class="Demospongiae" family="Desmacellidae" kingdom="Animalia" order="Poecilosclerida" pageId="128" pageNumber="129" phylum="Porifera" rank="family">Desmacellidae</taxonomicName>
and thus I propose to reassign
<taxonomicName id="4C01CA8577D7FF2CFF14A4D19175F849" box="[151,264,1978,2003]" class="Demospongiae" family="Desmacellidae" genus="Tylosigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Desmacellida" pageId="128" pageNumber="129" phylum="Porifera" rank="genus">
<emphasis id="B9756D1477D7FF2CFF14A4D19175F849" box="[151,264,1978,2003]" italics="true" pageId="128" pageNumber="129">Tylosigma</emphasis>
</taxonomicName>
to that family, until molecular analysis will have been done.
</paragraph>
</subSubSection>
</treatment>
</document>