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<document id="60649E16178D4DC70D5946D6EAE4A0BB" ID-DOI="10.5281/zenodo.5372938" ID-ISSN="0374-1036" ID-Zenodo-Dep="5372938" ID-ZooBank="FC0EF6ED-15E6-437A-94D3-25F3D8D98734" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630108704444" checkinUser="marcus" docAuthor="Vávra, Jiří Ch." docDate="2015" docId="03A46A7DFFFF0C63FC45899DFDB1FC00" docLanguage="en" docName="ActaEntMusNatPra.2015.55.1.85-103.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philomessor (Attumbrinus) henroti Ruzicka 1996" docType="treatment" docVersion="4" lastPageNumber="102" masterDocId="FF9D1205FFEE0C72FD018806FFE4FFAE" masterDocTitle="Review of the subgenus Attumbrinus (Coleoptera: Leiodidae: Cholevinae: Philomessor), with description of Philomessor lackneri sp. nov. from Morocco" masterLastPageNumber="103" masterPageNumber="85" pageNumber="102" updateTime="1699154510538" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C431FE907369D571EAC67705B456EFE3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C5E80696A16BA766C4D8C4B6FB983BF6">Review of the subgenus Attumbrinus (Coleoptera: Leiodidae: Cholevinae: Philomessor), with description of Philomessor lackneri sp. nov. from Morocco</mods:title>
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<mods:namePart id="099F21546638405D075A1CB2C60A4EF5">Vávra, Jiří Ch.</mods:namePart>
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<mods:title id="14CE61A5A6FA4AEA2F8DD79B8F55BEF2">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DF1EC5A673BF83DC8F73C2A9A2414EC2">2015</mods:date>
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<mods:number id="833F1EC6CD417D49DA5B6E1F83D5CCBE">55</mods:number>
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<mods:identifier id="CDDCB304EF2637785D0E885299F35507" type="DOI">10.5281/zenodo.5372938</mods:identifier>
<mods:identifier id="538B118E5860BD14D555DDA3D37FC564" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="BA069BA727A62D9E25FD5D8ADEAE8427" type="Zenodo-Dep">5372938</mods:identifier>
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<treatment id="03A46A7DFFFF0C63FC45899DFDB1FC00" ID-DOI="http://doi.org/10.5281/zenodo.5332547" ID-Zenodo-Dep="5332547" LSID="urn:lsid:plazi:treatment:03A46A7DFFFF0C63FC45899DFDB1FC00" httpUri="http://treatment.plazi.org/id/03A46A7DFFFF0C63FC45899DFDB1FC00" lastPageNumber="102" pageId="17" pageNumber="102">
<subSubSection id="C31788E0FFFF0C63FC45899DFC89FE1D" box="[324,877,411,435]" pageId="17" pageNumber="102" type="nomenclature">
<paragraph id="8BB2DB6BFFFF0C63FC45899DFC89FE1D" blockId="17.[324,877,411,435]" box="[324,877,411,435]" pageId="17" pageNumber="102">
<heading id="D0FA6C07FFFF0C63FC45899DFC89FE1D" bold="true" box="[324,877,411,435]" centered="true" fontSize="8" level="2" pageId="17" pageNumber="102" reason="3">
<taxonomicName id="4C0DA0E8FFFF0C63FC45899DFC89FE1D" ID-CoL="4G9BC" authority="Ruzicka, 1996" authorityName="Ruzicka" authorityYear="1996" box="[324,877,411,435]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="102" phylum="Arthropoda" rank="species" species="henroti" subGenus="Attumbrinus">
<emphasis id="B9790779FFFF0C63FC45899DFC89FE1D" bold="true" box="[324,877,411,435]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FC45899DFE28FE1D" bold="true" box="[324,460,411,435]" italics="true" pageId="17" pageNumber="102">Philomessor</emphasis>
(
<emphasis id="B9790779FFFF0C63FCDD899DFD83FE1D" bold="true" box="[476,615,411,435]" italics="true" pageId="17" pageNumber="102">Attumbrinus</emphasis>
)
<emphasis id="B9790779FFFF0C63FF76899DFD23FE1D" bold="true" box="[631,711,411,435]" italics="true" pageId="17" pageNumber="102">henroti</emphasis>
<bibRefCitation id="EF9CA69AFFFF0C63FFCF899AFC89FE1D" author="RUZICKA J." box="[718,877,412,435]" pageId="17" pageNumber="102" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">Růžička, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFFF0C63FF0389BCFD54FE63" box="[514,688,442,461]" pageId="17" pageNumber="102" type="description">
<paragraph id="8BB2DB6BFFFF0C63FF0389BCFD54FE63" blockId="17.[113,688,442,496]" box="[514,688,442,461]" pageId="17" pageNumber="102">
(
<figureCitation id="1336C7EEFFFF0C63FF0989BCFDAEFE63" box="[520,586,442,461]" captionStart="Figs 30–32" captionStartId="11.[113,150,1515,1534]" captionTargetBox="[113,1088,222,1430]" captionTargetId="figure-11@11.[113,1088,222,1452]" captionTargetPageId="11" captionText="Figs 30–32. General habitus of types with original labels, males. 30 – Philomessor (Attumbrinus) cloueti (Portevin, 1907), holotype; 31 – P. (A.) bolivari Jeannel, 1936, lectotype; 32 – P. (A.) henroti Růžička, 1996, holotype. An asterisk ‘*’ denotes upper and under side of the same label." figureDoi="http://doi.org/10.5281/zenodo.5372950" httpUri="https://zenodo.org/record/5372950/files/figure.png" pageId="17" pageNumber="102">Figs 32</figureCitation>
,
<figureCitation id="1336C7EEFFFF0C63FF5289BCFD6EFE63" box="[595,650,442,461]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="17" pageNumber="102">39, 40</figureCitation>
,
<figureCitation id="1336C7EEFFFF0C63FF9289BCFD4DFE63" box="[659,681,442,461]" captionStart="Fig" captionStartId="16.[151,182,1564,1583]" captionTargetBox="[152,1122,1132,1537]" captionTargetId="figure-95@16.[151,1127,1132,1540]" captionTargetPageId="16" captionText="Fig. 59. Distribution of Philomessor (Attumbrinus) Jeannel, 1936 in the northwest of Africa." figureDoi="http://doi.org/10.5281/zenodo.5372960" httpUri="https://zenodo.org/record/5372960/files/figure.png" pageId="17" pageNumber="102">59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFFF0C63FD7089DBFE35FE5E" box="[113,465,477,496]" pageId="17" pageNumber="102" type="reference_group">
<paragraph id="8BB2DB6BFFFF0C63FD7089DBFE35FE5E" blockId="17.[113,688,442,496]" box="[113,465,477,496]" pageId="17" pageNumber="102">
<treatmentCitationGroup id="AB1DFC45FFFF0C63FD7089DBFE35FE5E" box="[113,465,477,496]" pageId="17" pageNumber="102">
<treatmentCitation id="0AACFD7AFFFF0C63FD7089DBFE29FE5E" author="RUZICKA J." box="[113,461,477,496]" page="624" pageId="17" pageNumber="102" year="1996">
<taxonomicName id="4C0DA0E8FFFF0C63FD7089DBFE29FE5E" ID-CoL="4G9BC" authority="Ruzicka, 1996: 624" authorityName="Ruzicka" authorityPageNumber="624" authorityYear="1996" box="[113,461,477,496]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="102" phylum="Arthropoda" rank="species" species="henroti">
<emphasis id="B9790779FFFF0C63FD7089DBFEFAFE5E" box="[113,286,477,496]" italics="true" pageId="17" pageNumber="102">Philomessor henroti</emphasis>
<bibRefCitation id="EF9CA69AFFFF0C63FC2589DBFE29FE5E" author="RUZICKA J." box="[292,461,477,496]" pageId="17" pageNumber="102" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">Růžička, 1996: 624</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFFF0C63FD708A00FD32FCA0" pageId="17" pageNumber="102" type="materials_examined">
<paragraph id="8BB2DB6BFFFF0C63FD708A00FBCDFDB7" blockId="17.[113,1089,518,671]" box="[113,1065,518,537]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FD708A00FF0EFDB7" bold="true" box="[113,234,518,537]" pageId="17" pageNumber="102">
<typeStatus id="54B665C9FFFF0C63FD708A00FF7AFDB7" box="[113,158,518,537]" pageId="17" pageNumber="102">Type</typeStatus>
locality.
</emphasis>
‘Steppen bei Chichaoua, Marokko’ [
<collectingCountry id="F31A9BFBFFFF0C63FF258A00FD91FDB7" box="[548,629,518,537]" name="Morocco" pageId="17" pageNumber="102">Morocco</collectingCountry>
,
<collectingRegion id="49C91589FFFF0C63FF7C8A00FC53FDB7" box="[637,951,518,537]" country="Morocco" name="Marrakech-Tensift-Al Haouz" pageId="17" pageNumber="102">Marrakesh-Tensift-El Haouz Region</collectingRegion>
, Chichaoua].
</paragraph>
<paragraph id="8BB2DB6BFFFF0C63FD708A27FE97FD31" blockId="17.[113,1089,518,671]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FD708A27FEB6FD9A" bold="true" box="[113,338,545,564]" pageId="17" pageNumber="102">Type material examined.</emphasis>
<typeStatus id="54B665C9FFFF0C63FC568A27FE4EFD9C" box="[343,426,545,564]" pageId="17" pageNumber="102" type="holotype">HOLOTYPE</typeStatus>
(
<figureCitation id="1336C7EEFFFF0C63FCB78A27FE12FD9A" box="[438,502,545,564]" captionStart="Figs 30–32" captionStartId="11.[113,150,1515,1534]" captionTargetBox="[113,1088,222,1430]" captionTargetId="figure-11@11.[113,1088,222,1452]" captionTargetPageId="11" captionText="Figs 30–32. General habitus of types with original labels, males. 30 – Philomessor (Attumbrinus) cloueti (Portevin, 1907), holotype; 31 – P. (A.) bolivari Jeannel, 1936, lectotype; 32 – P. (A.) henroti Růžička, 1996, holotype. An asterisk ‘*’ denotes upper and under side of the same label." figureDoi="http://doi.org/10.5281/zenodo.5372950" httpUri="https://zenodo.org/record/5372950/files/figure.png" pageId="17" pageNumber="102">Fig. 32</figureCitation>
): J (
<collectionCode id="ED1C43AEFFFF0C63FF1E8A27FD81FD9A" box="[543,613,545,564]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="102">MNHN</collectionCode>
), ‘Steppen bei Chichaoua /
<collectingCountry id="F31A9BFBFFFF0C63FE4C8A27FC79FD9A" box="[845,925,545,564]" name="Morocco" pageId="17" pageNumber="102">Marokko</collectingCountry>
, lg. H. Franz [printed] / Sp 927 [underside of the label, handwritten] // MUSEUM PARIS [printed] / coll. / H. Henrot [handwritten by Henrot] //
<taxonomicName id="4C0DA0E8FFFF0C63FDE68A50FEB4FDC7" box="[231,336,598,617]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="102" phylum="Arthropoda" rank="genus">Philomessor</taxonomicName>
/ Franzi [handwritten over habitus sketch in red ink and text Attumbrinus
<taxonomicNameLabel id="A24ABA02FFFF0C63FED38A50FBF2FDC7" box="[978,1046,598,617]" pageId="17" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
– all probably by Henroťs manuscriptum] //
<taxonomicName id="4C0DA0E8FFFF0C63FCBE8A77FD4CFD2A" authority="" authorityName="Bolivari Jeannel" authorityYear="1936" box="[447,680,625,644]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="102" phylum="Arthropoda" rank="subGenus" subGenus="Attumbrinus">Philomessor (Attumbrinus)</taxonomicName>
/ henroti
<taxonomicNameLabel id="A24ABA02FFFF0C63FFF88A77FCC3FD2A" box="[761,807,625,644]" pageId="17" pageNumber="102" rank="species">sp. n.</taxonomicNameLabel>
J /
<typeStatus id="54B665C9FFFF0C63FE4A8A77FC2EFD2A" box="[843,970,625,644]" pageId="17" pageNumber="102" type="holotype">HOLOTYPUS</typeStatus>
/ Jan Růžička det. 1995 [red label, printed]’.
</paragraph>
<paragraph id="8BB2DB6BFFFF0C63FD708AB1FD32FCA0" blockId="17.[113,1089,695,942]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FD708AB1FF5CFD60" bold="true" box="[113,184,695,718]" pageId="17" pageNumber="102">Notes.</emphasis>
This species is known only from the
<typeStatus id="54B665C9FFFF0C63FF3D8AB1FD8EFD60" box="[572,618,695,718]" pageId="17" pageNumber="102">type</typeStatus>
specimen, described and figured by
<bibRefCitation id="EF9CA69AFFFF0C63FEE98AB1FF58FD40" author="RUZICKA J." pageId="17" pageNumber="102" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">
<subScript id="1789D92EFFFF0C63FEE98AB1FBA5FD63" attach="right" box="[1000,1089,695,718]" fontSize="7" pageId="17" pageNumber="102">RŮŽIČKA</subScript>
(1996)
</bibRefCitation>
. Photographs of the whole habitus (
<figureCitation id="1336C7EEFFFF0C63FF388AD1FD6DFD40" box="[569,649,727,750]" captionStart="Figs 30–32" captionStartId="11.[113,150,1515,1534]" captionTargetBox="[113,1088,222,1430]" captionTargetId="figure-11@11.[113,1088,222,1452]" captionTargetPageId="11" captionText="Figs 30–32. General habitus of types with original labels, males. 30 – Philomessor (Attumbrinus) cloueti (Portevin, 1907), holotype; 31 – P. (A.) bolivari Jeannel, 1936, lectotype; 32 – P. (A.) henroti Růžička, 1996, holotype. An asterisk ‘*’ denotes upper and under side of the same label." figureDoi="http://doi.org/10.5281/zenodo.5372950" httpUri="https://zenodo.org/record/5372950/files/figure.png" pageId="17" pageNumber="102">Fig. 32</figureCitation>
), posterior pronotal corners (
<figureCitation id="1336C7EEFFFF0C63FEBF8AD1FBE8FD40" box="[958,1036,727,750]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="17" pageNumber="102">Fig. 40</figureCitation>
) and the pubescence of elytra (
<figureCitation id="1336C7EEFFFF0C63FC838AF1FE34FCA0" box="[386,464,759,782]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="17" pageNumber="102">Fig. 39</figureCitation>
) are supplemented here.
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFFF0C63FD708B11FE38FC20" pageId="17" pageNumber="102" type="biology_ecology">
<paragraph id="8BB2DB6BFFFF0C63FD708B11FE38FC20" blockId="17.[113,1089,695,942]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FD708B11FF29FC80" bold="true" box="[113,205,791,814]" pageId="17" pageNumber="102">Biology.</emphasis>
Unknown. According to the personal communication of the collector Herbert Franz (Mödling,
<collectingCountry id="F31A9BFBFFFF0C63FDE38B31FED6FCE0" box="[226,306,823,846]" name="Austria" pageId="17" pageNumber="102">Austria</collectingCountry>
)
<bibRefCitation id="EF9CA69AFFFF0C63FC418B31FE02FCE0" author="RUZICKA J." box="[320,486,823,846]" pageId="17" pageNumber="102" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">
<subScript id="1789D92EFFFF0C63FC418B31FE7CFCE3" attach="right" box="[320,408,823,846]" fontSize="7" pageId="17" pageNumber="102">RŮŽIČKA</subScript>
(1996)
</bibRefCitation>
wrote that the
<typeStatus id="54B665C9FFFF0C63FF878B31FD50FCE0" box="[646,692,823,846]" pageId="17" pageNumber="102">type</typeStatus>
specimen was collected west of Chichaoua on
<date id="FFB3FDABFFFF0C63FDE58B51FEB0FCC0" box="[228,340,855,878]" pageId="17" pageNumber="102" value="1963-04-13">13.iv.1963</date>
under a stone in the steppe and supposed that, like all his congeners, it is also a myrmecophilous species.
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFFF0C63FD708B91FDB1FC00" box="[113,597,919,942]" pageId="17" pageNumber="102" type="distribution">
<paragraph id="8BB2DB6BFFFF0C63FD708B91FDB1FC00" blockId="17.[113,1089,695,942]" box="[113,597,919,942]" pageId="17" pageNumber="102">
<emphasis id="B9790779FFFF0C63FD708B91FEE0FC00" bold="true" box="[113,260,919,942]" pageId="17" pageNumber="102">Distribution.</emphasis>
<collectingCountry id="F31A9BFBFFFF0C63FC0B8B91FE8BFC00" box="[266,367,919,942]" name="Morocco" pageId="17" pageNumber="102">Morocco</collectingCountry>
: Chichaoua (
<figureCitation id="1336C7EEFFFF0C63FCF88B91FDADFC00" box="[505,585,919,942]" captionStart="Fig" captionStartId="16.[151,182,1564,1583]" captionTargetBox="[152,1122,1132,1537]" captionTargetId="figure-95@16.[151,1127,1132,1540]" captionTargetPageId="16" captionText="Fig. 59. Distribution of Philomessor (Attumbrinus) Jeannel, 1936 in the northwest of Africa." figureDoi="http://doi.org/10.5281/zenodo.5372960" httpUri="https://zenodo.org/record/5372960/files/figure.png" pageId="17" pageNumber="102">Fig. 59</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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