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<mods:title id="BBF4883A3BFBD9ABF362B5122AEDE469">First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)</mods:title>
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<treatment id="039C6B6DFF84FFACD61DC5754C22FDC8" ID-DOI="http://doi.org/10.5281/zenodo.5748097" ID-GBIF-Taxon="190875388" ID-Zenodo-Dep="5748097" LSID="urn:lsid:plazi:treatment:039C6B6DFF84FFACD61DC5754C22FDC8" httpUri="http://treatment.plazi.org/id/039C6B6DFF84FFACD61DC5754C22FDC8" lastPageId="7" lastPageNumber="671" pageId="6" pageNumber="670">
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<paragraph id="8B8ADA7BFF84FFADD61DC5754B6FF9F1" blockId="6.[892,1375,1598,1619]" box="[892,1375,1598,1619]" pageId="6" pageNumber="670">
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<taxonomicName id="4C35A1F8FF84FFADD6E0C5754B6FF9F1" authority="SEBOUI" authorityName="CHINJIENSIS LINDSAY" authorityYear="1988" box="[897,1375,1598,1619]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD6E0C5754A04F9F1" box="[897,1076,1598,1619]" italics="true" pageId="6" pageNumber="670">PARADAKKAMYS</emphasis>
<emphasis id="B9410669FF84FFADD122C50A4AA5F9F3" box="[1091,1173,1601,1617]" italics="true" pageId="6" pageNumber="670">SEBOUI</emphasis>
(JAEGER, 1977B)
</taxonomicName>
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<paragraph id="8B8ADA7BFF84FFADD65BC52F4AD4F8CD" blockId="6.[826,1441,1636,1903]" pageId="6" pageNumber="670">
<bibRefCitation id="EFA4A78AFF84FFADD65BC52F4DD4F9D8" author="Jaeger JJ" box="[826,996,1636,1658]" pageId="6" pageNumber="670" pagination="1 - 166" refId="ref11251" refString="Jaeger JJ. 1977 b. Les rongeurs du Miocene Moyen et Superieur du Maghreb. Palaeovertebrata 8: 1 - 166." type="journal article" year="1977">Jaeger (1977b)</bibRefCitation>
erected this species on the basis of 18 upper molars and 12 lower molars from the early Late Miocene site of Oued Zra, middle-Atlas,
<collectingCountry id="F3229AEBFF84FFADD068C5E94B5AF915" box="[1289,1386,1698,1719]" name="Morocco" pageId="6" pageNumber="670">Morocco</collectingCountry>
. The
<typeStatus id="548E64D9FF84FFADD65BC58B4DABF974" box="[826,923,1728,1750]" pageId="6" pageNumber="670" type="holotype">holotype</typeStatus>
(OZ-20) is a left M2 housed at the SGM. It was originally placed in the genus
<taxonomicName id="4C35A1F8FF84FFADD1A8C5954B5CF951" box="[1225,1388,1758,1779]" class="Mammalia" family="Muridae" genus="Myocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD1A8C5954B5CF951" box="[1225,1388,1758,1779]" italics="true" pageId="6" pageNumber="670">Myocricetodon</emphasis>
</taxonomicName>
, but was later allocated to the genus
<taxonomicName id="4C35A1F8FF84FFADD19BC5B64B42F8B0" authorityName="ZAIANI JAEGER" authorityYear="1977" box="[1274,1394,1789,1810]" class="Mammalia" family="Muridae" genus="Dakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD19BC5B64B42F8B0" box="[1274,1394,1789,1810]" italics="true" pageId="6" pageNumber="670">Dakkamys</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA4A78AFF84FFADD65BC4574DDEF890" author="Lindsay EH" box="[826,1006,1820,1842]" pageId="6" pageNumber="670" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay (1988)</bibRefCitation>
and to
<taxonomicName id="4C35A1F8FF84FFADD13EC4574B3AF893" authorityName="CHINJIENSIS LINDSAY" authorityYear="1988" box="[1119,1290,1820,1841]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD13EC4574B3AF893" box="[1119,1290,1820,1841]" italics="true" pageId="6" pageNumber="670">Paradakkamys</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA4A78AFF84FFADD027C4574DE7F8F2" author="Tong H &amp; Jaeger JJ" pageId="6" pageNumber="670" pagination="51 - 73" refId="ref12202" refString="Tong H, Jaeger JJ. 1993. Muroid rodents from the Middle Miocene Fort Ternan locality (Kenya) and their contribution to the phylogeny of muroids. Palaeontographica, Abteilung A 229: 51 - 73." type="journal article" year="1993">Tong &amp; Jaeger (1993)</bibRefCitation>
. None of these allocations are confirmed by the cladistic analysis (see below).
</paragraph>
<paragraph id="8B8ADA7BFF85FFACD5C9C38D4C22FDC8" blockId="7.[144,759,198,618]" pageId="7" pageNumber="671">
The M1 of
<taxonomicName id="4C35A1F8FF85FFACD451C38D4FEBFF79" authorityName="CHINJIENSIS LINDSAY" authorityYear="1988" box="[304,475,198,219]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="671" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF85FFACD451C38D4FEBFF79" box="[304,475,198,219]" italics="true" pageId="7" pageNumber="671">Paradakkamys</emphasis>
</taxonomicName>
<emphasis id="B9410669FF85FFACD488C38D4C00FF79" box="[489,560,198,219]" italics="true" pageId="7" pageNumber="671">seboui</emphasis>
differs from AJ
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having a well-developed longitudinal crest and an enterostyle that joins the posterior side of the protocone. The M2 of
<emphasis id="B9410669FF85FFACD430C2694F56FE95" box="[337,358,290,311]" italics="true" pageId="7" pageNumber="671">P.</emphasis>
<emphasis id="B9410669FF85FFACD410C2694F88FE95" box="[369,440,290,311]" italics="true" pageId="7" pageNumber="671">seboui</emphasis>
has the anterior arm of the hypocone connected to the posterior arm of the paracone, forming a new longitudinal crest, and a small enterostyle joined with the posterior arm of the protocone. In contrast, AJ11 and AJ12 lack a new longitudinal crest and show an isolated small enterostyle. The morphology of the m1 of
<emphasis id="B9410669FF85FFACD498C2914C3EFE4D" box="[505,526,474,495]" italics="true" pageId="7" pageNumber="671">P.</emphasis>
<emphasis id="B9410669FF85FFACD77DC2924C53FE4C" box="[540,611,473,494]" italics="true" pageId="7" pageNumber="671">seboui</emphasis>
is also very different from that of AJ
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lacking the strong reduction of the anterior part of the tooth and in having a well-developed longitudinal crest, which is lacking in the Arabian specimen.
</paragraph>
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