treatments-xml/data/03/98/4C/03984C68BB57F571E8A1F8FDBEA49411.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="2D5408587A7442DF5748B484268A815F">New species and records of heterostigmatic mites (Acari: Prostigmata: Heterostigmata) phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from northwestern Iran</mods:title>
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<mods:namePart id="C59187027D83E6A1C406E0173DFF1568">Sobhi, Mohammad</mods:namePart>
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<mods:namePart id="CB746CD02498F9F6C24BA81CE89E5427">Hajiqanbar, Hamidreza</mods:namePart>
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<mods:namePart id="B48839EC7C45165589348A03146E68CA">Mortazavi, Azim</mods:namePart>
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<treatment id="03984C68BB57F571E8A1F8FDBEA49411" ID-DOI="http://doi.org/10.5281/zenodo.6051995" ID-GBIF-Taxon="131876701" ID-Zenodo-Dep="6051995" LSID="urn:lsid:plazi:treatment:03984C68BB57F571E8A1F8FDBEA49411" httpUri="http://treatment.plazi.org/id/03984C68BB57F571E8A1F8FDBEA49411" lastPageId="6" lastPageNumber="433" pageId="5" pageNumber="432">
<subSubSection id="C32BAEF5BB57F571E8A1F8FDBFFA96D3" lastPageId="6" lastPageNumber="433" pageId="5" pageNumber="432" type="nomenclature">
<paragraph id="8B8EFD7EBB57F572E8A1F8FDBEF99149" blockId="5.[151,767,1845,1871]" box="[151,767,1845,1871]" pageId="5" pageNumber="432">
<heading id="D0C64A12BB57F572E8A1F8FDBEF99149" bold="true" box="[151,767,1845,1871]" fontSize="11" level="1" pageId="5" pageNumber="432" reason="1">
<taxonomicName id="4C3186FDBB57F572E8A1F8FDBEF99149" authority="Mahunka, 1970" authorityName="Mahunka" authorityYear="1970" box="[151,767,1845,1871]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="B945216CBB57F572E8A1F8FDBEF99149" bold="true" box="[151,767,1845,1871]" pageId="5" pageNumber="432">
<emphasis id="B945216CBB57F572E8A1F8FDBE279149" bold="true" box="[151,545,1845,1871]" italics="true" pageId="5" pageNumber="432">Pseudopygmephorellus szekessyi</emphasis>
(
<bibRefCitation id="EFA0808FBB57F572EA07F8FDBEF19149" author="Mahunka" box="[561,759,1845,1871]" pageId="5" pageNumber="432" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Hungaricae, 16, 137 - 174." type="journal article" year="1970">Mahunka, 1970</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8EFD7EBB57F571E8A1F8B3BFFA96D3" blockId="5.[151,1437,1915,2012]" lastBlockId="6.[151,1436,151,536]" lastPageId="6" lastPageNumber="433" pageId="5" pageNumber="432">
This species was described from
<collectingCountry id="F326BDEEBB57F572EA13F8B4BE819192" box="[549,647,1916,1940]" name="Hungary" pageId="5" pageNumber="432">Hungary</collectingCountry>
from litter in the nest of
<taxonomicName id="4C3186FDBB57F572EBF2F8B5B8979192" authority="Schrank" authorityName="Schrank" box="[964,1169,1915,1940]" class="Mammalia" family="Muridae" genus="Microtus" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="B945216CBB57F572EBF2F8B5B82E9192" box="[964,1064,1917,1940]" italics="true" pageId="5" pageNumber="432">Microtus</emphasis>
Schrank
</taxonomicName>
(
<taxonomicName id="4C3186FDBB57F572EC93F8B3B9169192" authorityName="Bowditch" authorityYear="1821" box="[1189,1296,1915,1940]" class="Mammalia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="432" phylum="Chordata" rank="order">Rodentia</taxonomicName>
:
<taxonomicName id="4C3186FDBB57F572ED29F8B3B9929192" box="[1311,1428,1915,1940]" class="Mammalia" family="Cricetidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="432" phylum="Chordata" rank="family">Cricetidae</taxonomicName>
) (
<bibRefCitation id="EFA0808FBB57F572E8A8F857BD4A91BE" author="Mahunka" box="[158,332,1951,1976]" pageId="5" pageNumber="432" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Hungaricae, 16, 137 - 174." type="journal article" year="1970">Mahunka 1970</bibRefCitation>
).
<bibRefCitation id="EFA0808FBB57F572E969F857BDF591BE" author="Kaluz" box="[351,499,1951,1976]" pageId="5" pageNumber="432" refString="Kaluz, S. (1992) Some aspects of mite (Acarina) occurrence on beetles (Coleoptera). Pracovisk aSlovenska Entomologicka Spoloˇ cnost ', 9, 45 - 50." type="journal article" year="1992">Kaluz (1992)</bibRefCitation>
reported it from
<collectingCountry id="F326BDEEBB57F572EA87F868BF0D91BE" box="[689,779,1952,1976]" name="Norway" pageId="5" pageNumber="432">Norway</collectingCountry>
as phoretic on
<taxonomicName id="4C3186FDBB57F572EB82F868B90C91BE" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[948,1290,1951,1976]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="vernalis">
<emphasis id="B945216CBB57F572EB82F868B88F91BE" box="[948,1161,1951,1976]" italics="true" pageId="5" pageNumber="432">Geotrupes vernalis</emphasis>
(Linnaeus)
</taxonomicName>
(
<taxonomicName id="4C3186FDBB57F572ED2EF857B99F91BE" box="[1304,1433,1951,1976]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C3186FDBB57F572E8A1F80BBD2191DA" box="[151,295,1987,2012]" class="Insecta" family="Geotrupidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="432" phylum="Arthropoda" rank="family">Geotrupidae</taxonomicName>
). This species has a wider distribution and until now, it has been found several times from the Palaearctic (including
<collectingCountry id="F326BDEEBB54F571E9A2FF50BDC596B6" box="[404,451,152,176]" name="Iran" pageId="6" pageNumber="433">Iran</collectingCountry>
) and Nearctic regions in association with geotrupid and trogid beetles (
<bibRefCitation id="EFA0808FBB54F571ECDFFF5FB99196B6" author="Mahunka" box="[1257,1431,151,176]" pageId="6" pageNumber="433" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Hungaricae, 16, 137 - 174." type="journal article" year="1970">Mahunka 1970</bibRefCitation>
;
<bibRefCitation id="EFA0808FBB54F571E8A1FF74BD1A96D2" author="Kaluz" box="[151,284,188,213]" pageId="6" pageNumber="433" refString="Kaluz, S. (1992) Some aspects of mite (Acarina) occurrence on beetles (Coleoptera). Pracovisk aSlovenska Entomologicka Spoloˇ cnost ', 9, 45 - 50." type="journal article" year="1992">Kaluz 1992</bibRefCitation>
;
<bibRefCitation id="EFA0808FBB54F571E911FF74BDB296D2" author="Masan" box="[295,436,188,212]" pageId="6" pageNumber="433" refString="Masan, P. (1993) Mites (Acarina) associated with species of Trox (Coleoptera: Scarabaeidae). European Journal of Entomology, 90, 359 - 364." type="journal article" year="1993">Mašan 1993</bibRefCitation>
; Khaustov 2008;
<bibRefCitation id="EFA0808FBB54F571EA4FFF74BF0F96D2" author="Philips" box="[633,777,188,213]" pageId="6" pageNumber="433" refString="Philips, J. R. (2009) The mite (Acarina) fauna of trogid beetles (Coleoptera: Trogidae). International Journal of Acarology, 36, 1 - 17." type="journal article" year="2009">Philips 2009</bibRefCitation>
;
<bibRefCitation id="EFA0808FBB54F571EB23FF74BFF696D3" author="Katlav" box="[789,1008,188,213]" pageId="6" pageNumber="433" refString="Katlav, A., Hajiqanbar, H. &amp; Talebi, A. A. (2015 b) Pseudopygmephorellus mazandaranicus sp. nov. (Acari: Heterostigmata: Pygmephoridae), phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from Iran. Zootaxa, 3919 (1), 100 - 110." type="journal article" year="2015" yearSuffix="b">
Katlav
<emphasis id="B945216CBB54F571EB50FF75BF9896D3" box="[870,926,188,213]" italics="true" pageId="6" pageNumber="433">et al.</emphasis>
2015b
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32BAEF5BB54F571E8F1FF17BE1B9746" pageId="6" pageNumber="433" type="materials_examined">
<paragraph id="8B8EFD7EBB54F571E8F1FF17BE1B9746" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="433">
<materialsCitation id="3B59F723BB54F571E8F1FF17BE1B9746" ID-GBIF-Occurrence="1563872047" collectingDate="2016-05-14" collectorName="M. Sobhi." country="Iran" county="Material" latitude="38.23" location="Northwestern Iran" longLatPrecision="706" longitude="47.41" municipality="One" pageId="6" pageNumber="433" specimenCount="1" stateProvince="Ardabil province">
<emphasis id="B945216CBB54F571E8F1FF17BDB696FE" bold="true" box="[199,432,223,248]" pageId="6" pageNumber="433">
<collectingCounty id="62EF85F2BB54F571E8F1FF17BD2996FE" box="[199,303,223,248]" pageId="6" pageNumber="433">Material</collectingCounty>
examined.
</emphasis>
<collectingMunicipality id="6BEA6704BB54F571E981FF28BDE096FE" box="[439,486,224,248]" pageId="6" pageNumber="433">One</collectingMunicipality>
female,
<collectingCountry id="F326BDEEBB54F571EA70FF17BE7D96FE" box="[582,635,223,248]" name="Iran" pageId="6" pageNumber="433">
<emphasis id="B945216CBB54F571EA70FF17BE7D96FE" bold="true" box="[582,635,223,248]" pageId="6" pageNumber="433">Iran</emphasis>
</collectingCountry>
,
<location id="8EEEABA5BB54F571EABFFF17BF5E96F1" LSID="urn:lsid:plazi:treatment:03984C68BB57F571E8A1F8FDBEA49411:8EEEABA5BB54F571EABFFF17BF5E96F1" box="[649,856,223,248]" country="Iran" county="Material" latitude="38.23" longLatPrecision="706" longitude="47.41" municipality="One" name="Northwestern Iran" pageId="6" pageNumber="433" stateProvince="Ardabil province">Northwestern Iran</location>
,
<collectingRegion id="49F5339CBB54F571EB52FF17B82396FE" box="[868,1061,223,248]" country="Iran" name="Ardabil" pageId="6" pageNumber="433">Ardabil province</collectingRegion>
, forest park, vicinity of
<location id="8EEEABA5BB54F571ED07FF17BCD3971B" LSID="urn:lsid:plazi:treatment:03984C68BB57F571E8A1F8FDBEA49411:8EEEABA5BB54F571ED07FF17BCD3971B" country="Iran" county="Material" latitude="38.23" longLatPrecision="706" longitude="47.41" municipality="One" name="Meshginshahr" pageId="6" pageNumber="433" stateProvince="Ardabil province">Meshginshahr</location>
,
<geoCoordinate id="EE059BB9BB54F571E8D4FECCBD45971A" box="[226,323,260,284]" direction="north" orientation="latitude" pageId="6" pageNumber="433" precision="555" value="38.23">38.23°N</geoCoordinate>
,
<geoCoordinate id="EE059BB9BB54F571E966FECCBDAB971A" box="[336,429,260,284]" direction="east" orientation="longitude" pageId="6" pageNumber="433" precision="555" value="47.41">47.41°E</geoCoordinate>
, phoretic
<collectedFrom id="382DBE83BB54F571EA29FECDBF3C971A" box="[543,826,260,285]" pageId="6" pageNumber="433">
on
<taxonomicName id="4C3186FDBB54F571EA75FECCBF3C971A" authority="Say" authorityName="Say" authorityYear="1825" box="[579,826,260,285]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B945216CBB54F571EA75FECCBF01971A" box="[579,775,260,285]" italics="true" pageId="6" pageNumber="433">Aphodius vittatus</emphasis>
Say
</taxonomicName>
</collectedFrom>
(
<taxonomicName id="4C3186FDBB54F571EB7CFECCBFCD971B" box="[842,971,260,285]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C3186FDBB54F571EBE1FECCB86B971B" authorityName="Latreille" authorityYear="1802" box="[983,1133,260,285]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
) collected in dung pads of cow,
<collectingDate id="EFCB2256BB54F571E8E4FEE0BD6C9746" box="[210,362,295,320]" pageId="6" pageNumber="433" value="2016-05-14">
<date id="FF8FDBBEBB54F571E8E4FEE0BD6C9746" box="[210,362,295,320]" pageId="6" pageNumber="433" value="2016-05-14">14 May 2016</date>
</collectingDate>
, coll.
<collectorName id="26C498A8BB54F571E99BFEE0BE1B9746" box="[429,541,295,320]" pageId="6" pageNumber="433">M. Sobhi.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32BAEF5BB54F571E8F1FE84BEA49411" pageId="6" pageNumber="433" type="discussion">
<paragraph id="8B8EFD7EBB54F571E8F1FE84BEA49411" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="433">
<emphasis id="B945216CBB54F571E8F1FE84BD3D9763" bold="true" box="[199,315,332,357]" pageId="6" pageNumber="433">Remarks.</emphasis>
This species was transferred by
<bibRefCitation id="EFA0808FBB54F571EA8AFE84BF8D9763" author="Khaustov" box="[700,907,332,357]" pageId="6" pageNumber="433" refString="Khaustov, A. A. (2008 b) A new genus of mites of the family Pygmephoridae (Acari, Heterostigmata) from Russia. Zoologichesky Zhournal, 87, 891 - 896. [in Russian]" type="journal article" year="2008" yearSuffix="b">Khaustov (2008b)</bibRefCitation>
from the genus
<taxonomicName id="4C3186FDBB54F571EC78FE84B9059763" authorityName="Cross &amp; Moser" authorityYear="1971" box="[1102,1283,332,357]" class="Arachnida" family="Siteroptidae" genus="Pygmephorellus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="B945216CBB54F571EC78FE84B9059763" box="[1102,1283,332,357]" italics="true" pageId="6" pageNumber="433">Pygmephorellus</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4C3186FDBB54F571E8A1FEA7BD9D978E" authorityName="Khaustov" authorityYear="2008" box="[151,411,367,392]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="B945216CBB54F571E8A1FEA7BD9D978E" box="[151,411,367,392]" italics="true" pageId="6" pageNumber="433">Pseudopygmephorellus</emphasis>
</taxonomicName>
. The genus
<taxonomicName id="4C3186FDBB54F571EA13FEA7BF2F978E" authorityName="Khaustov" authorityYear="2008" box="[549,809,367,392]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="B945216CBB54F571EA13FEA7BF2F978E" box="[549,809,367,392]" italics="true" pageId="6" pageNumber="433">Pseudopygmephorellus</emphasis>
</taxonomicName>
includes six species among which only two species are associated with scarabaeid beetles as follows:
<taxonomicName id="4C3186FDBB54F571EA87FE5DB8CB97AB" authority="Katlav &amp; Hajiqanbar, 2015" authorityName="Katlav &amp; Hajiqanbar" authorityYear="2015" box="[689,1229,404,429]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="mazandaranicus">
<emphasis id="B945216CBB54F571EA87FE5DBF8F97AB" box="[689,905,404,429]" italics="true" pageId="6" pageNumber="433">P. mazandaranicus</emphasis>
Katlav &amp; Hajiqanbar, 2015
</taxonomicName>
phoretic on three different scarabaeid dung beetles,
<taxonomicName id="4C3186FDBB54F571EA17FE7FBEBE97D6" authorityName="Latreille" authorityYear="1802" box="[545,696,439,464]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="B945216CBB54F571EA17FE7FBEBE97D6" box="[545,696,439,464]" italics="true" pageId="6" pageNumber="433">Onthophagus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C3186FDBB54F571EAC4FE7FB85B97D6" authority="Kugelann" authorityName="Kugelann" authorityYear="1792" box="[754,1117,439,464]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B945216CBB54F571EAC4FE7FBFD497D6" box="[754,978,439,464]" italics="true" pageId="6" pageNumber="433">Aphodius depressus</emphasis>
(Kugelann)
</taxonomicName>
and
<taxonomicName id="4C3186FDBB54F571ECAEFE70BD8197F3" authority="Duftschmid (Katlav et al. 2015b)" authorityName="Duftschmidt" authorityYear="1805" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B945216CBB54F571ECAEFE70B90897D6" box="[1176,1294,440,464]" italics="true" pageId="6" pageNumber="433">A. varians</emphasis>
Duftschmid (
<bibRefCitation id="EFA0808FBB54F571E8A8FE14BD8697F3" author="Katlav" box="[158,384,476,501]" pageId="6" pageNumber="433" refString="Katlav, A., Hajiqanbar, H. &amp; Talebi, A. A. (2015 b) Pseudopygmephorellus mazandaranicus sp. nov. (Acari: Heterostigmata: Pygmephoridae), phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from Iran. Zootaxa, 3919 (1), 100 - 110." type="journal article" year="2015" yearSuffix="b">
Katlav
<emphasis id="B945216CBB54F571E8C5FE15BD2B97F3" box="[243,301,476,501]" italics="true" pageId="6" pageNumber="433">et al.</emphasis>
2015b
</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="4C3186FDBB54F571E9F3FE15BE4197F3" authorityName="Mahunka" authorityYear="1970" box="[453,583,476,501]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="B945216CBB54F571E9F3FE15BE4197F3" box="[453,583,476,501]" italics="true" pageId="6" pageNumber="433">P. szekessyi</emphasis>
</taxonomicName>
phoretic on
<taxonomicName id="4C3186FDBB54F571EAEEFE14BF9A97F2" authorityName="Say" authorityYear="1825" box="[728,924,476,501]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B945216CBB54F571EAEEFE14BF9A97F2" box="[728,924,476,501]" italics="true" pageId="6" pageNumber="433">Aphodius vittatus</emphasis>
</taxonomicName>
(current study). The association between the mite
<taxonomicName id="4C3186FDBB54F571E8F9FDC9BD56941E" authorityName="Mahunka" authorityYear="1970" box="[207,336,511,536]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="B945216CBB54F571E8F9FDC9BD56941E" box="[207,336,511,536]" italics="true" pageId="6" pageNumber="433">P. szekessyi</emphasis>
</taxonomicName>
and beetles
<taxonomicName id="4C3186FDBB54F571E9EDFDC8BE4B941E" authorityName="Say" authorityYear="1825" box="[475,589,512,536]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B945216CBB54F571E9EDFDC8BE4B941E" box="[475,589,512,536]" italics="true" pageId="6" pageNumber="433">A. vittatus</emphasis>
</taxonomicName>
is new.
</paragraph>
</subSubSection>
</treatment>
</document>