205 lines
21 KiB
XML
205 lines
21 KiB
XML
<document ID-DOI="10.1093/zoolinnean/zlac015" ID-GBIF-Dataset="372f8265-c53c-43fa-9448-07d5a58d311f" ID-ISSN="0024-4082" ID-Zenodo-Dep="7037958" ID-ZooBank="78C56409-FCCF-4116-8D8C-FF66B247C56C" approvalRequired="8" approvalRequired_for_taxonomicNames="2" approvalRequired_for_treatments="6" checkinTime="1661767832469" checkinUser="plazi" docAuthor="Kajihara, Hiroshi, Abukawa, Shushi & Chernyshev, Alexei V." docDate="2022" docId="038A8132EE7BED56FF14CC4EFBA8FEEA" docLanguage="en" docName="zlac015.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.6:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="6" docTitle="Oxypolella hiebertae Kajihara & Abukawa & Chernyshev 2022, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="519" masterDocId="FFB3F94AEE6BED46FFE9CE7DFFF7FFBC" masterDocTitle="Exploring the basal topology of the heteronemertean tree of life: establishment of a new family, along with turbotaxonomy of Valenciniidae (Nemertea: Pilidiophora: Heteronemertea)" masterLastPageNumber="548" masterPageNumber="503" pageNumber="519" updateTime="1661954276786" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Exploring the basal topology of the heteronemertean tree of life: establishment of a new family, along with turbotaxonomy of Valenciniidae (Nemertea: Pilidiophora: Heteronemertea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kajihara, Hiroshi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Abukawa, Shushi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chernyshev, Alexei V.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="volume">
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<mods:number>196</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>503</mods:start>
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<mods:end>548</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlac015</mods:identifier>
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<mods:identifier type="GBIF-Dataset">372f8265-c53c-43fa-9448-07d5a58d311f</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7037958</mods:identifier>
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<mods:identifier type="ZooBank">78C56409-FCCF-4116-8D8C-FF66B247C56C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7036915" ID-GBIF-Taxon="199902355" ID-Zenodo-Dep="7036915" LSID="urn:lsid:plazi:treatment:038A8132EE7BED56FF14CC4EFBA8FEEA" httpUri="http://treatment.plazi.org/id/038A8132EE7BED56FF14CC4EFBA8FEEA" lastPageNumber="519" pageId="16" pageNumber="519">
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<subSubSection box="[253,653,563,587]" pageId="16" pageNumber="519" type="nomenclature">
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<paragraph blockId="16.[253,653,563,587]" box="[253,653,563,587]" pageId="16" pageNumber="519">
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<heading box="[253,653,563,587]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="519" reason="2">
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<taxonomicName authority="Kajihara & Abukawa & Chernyshev, 2022" authorityName="Kajihara & Abukawa & Chernyshev" authorityYear="2022" box="[253,547,563,587]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="hiebertae" status="sp. nov.">OXYPOLELLA HIEBERTAE</taxonomicName>
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<emphasis bold="true" box="[554,653,563,587]" pageId="16" pageNumber="519">
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<taxonomicNameLabel box="[554,653,563,587]" pageId="16" pageNumber="519" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="519" type="description">
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<paragraph blockId="16.[401,504,602,627]" box="[401,504,602,627]" pageId="16" pageNumber="519">
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(
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<figureCitation box="[410,496,602,627]" captionStart="Figure 2" captionStartId="11.[168,247,1520,1542]" captionTargetBox="[328,1288,196,1480]" captionTargetId="figure-25@11.[328,1288,196,1480]" captionTargetPageId="11" captionText="Figure 2. A, Eopilidion misakiense gen. et sp. nov., holotype, ICHUM 6303, lateral view, head to the right; B, Oxypolella hiebertae sp. nov., holotype, MIMB 42256; C, Cephalomastax brevis Iwata, 1957, ICHUM 6304; D, E, Valencinura jambio sp. nov., holotype, ICHUM 6305, anterior body fragment (D) and enlargement of intestinal region (E) showing yellowish proboscis (indicated by arrowheads); F, Baseodiscus takakurai Gibson, 1995, ICHUM 6306; G, Baseodiscus profundus sp. nov., holotype, MIMB 42257; H, I, Baseodiscus narusei sp. nov., holotype, ICHUM 6310, anterior end of body, dorsal (H) and ventral (I) views; J, Baseodiscus paracelensis sp. nov., holotype, MIMB 33132; K, Baseodiscus komatsui sp. nov., holotype, NMNS-Ne 1; L, Baseodiscus unicolor Stiasny-Wijnhoff, 1925; M, Baseodiscus giribeti sp. nov., holotype, MCZ IZ-135324; N, Baseodiscus cf. amboinensis (Staub, 1900); O, Baseodiscus hemprichii (Ehrenberg, 1828-1831), specimen from Okinawa, different from any voucher specimens in this study; P, Baseodiscus zebra sp. nov., holotype, RUMF-ZN-00001; Q, Baseodiscus quinquelineatus (Quoy & Gaimard, 1833), ICHUM 6319. Photographs by H. Kajihara (A, C, D, E, F, O), A.V. Chernyshev (B, G, J), T. Naruse (H, I), H. Komatsu (K), G. Giribet (L, M), D. Uyeno (P), and R.Yoshida (Q)." figureDoi="http://doi.org/10.5281/zenodo.7037964" httpUri="https://zenodo.org/record/7037964/files/figure.png" pageId="16" pageNumber="519">FIG. 2B</figureCitation>
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)
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</paragraph>
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<paragraph blockId="16.[145,623,642,664]" box="[145,623,642,664]" pageId="16" pageNumber="519">
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Oxypolellinae gen. sp.:
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<bibRefCitation author="Hiebert TC" box="[407,618,642,664]" pageId="16" pageNumber="519" refId="ref37105" refString="Hiebert TC. 2016. New nemertean diversity discovered in the Northeast Pacific, using surveys of both planktonic larvae and benthic adults. Unpublished D. Phil. Thesis, University of Oregon." type="book" year="2016">Hiebert, 2016: 318</bibRefCitation>
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.
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</paragraph>
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<paragraph blockId="16.[145,762,710,763]" pageId="16" pageNumber="519">
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<emphasis box="[145,479,710,732]" italics="true" pageId="16" pageNumber="519">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. org:act:
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<uuid box="[223,758,741,763]" pageId="16" pageNumber="519">1F5D91AE-5C57-4E9A-BE20-C05D2DA554B4</uuid>
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.
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</paragraph>
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<paragraph blockId="16.[145,762,804,1040]" pageId="16" pageNumber="519">
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<emphasis box="[145,373,804,825]" italics="true" pageId="16" pageNumber="519">Material examined:</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3897229318" collectingDate="2007-05-23" collectionCode="V" country="Vietnam" elevation="3" latitude="15.383333" location="Cu-Lao Re Island" longLatPrecision="1285" longitude="109.083336" pageId="16" pageNumber="519" specimenCode="MIMB 42256" specimenCount="1" typeStatus="holotype">
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Four specimens, all collected by A.
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<collectionCode box="[173,187,835,856]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="519" type="Museum">V</collectionCode>
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. Chernyshev.
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<typeStatus box="[342,443,834,856]" pageId="16" pageNumber="519">Holotype</typeStatus>
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,
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<specimenCode box="[454,602,834,856]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="16" pageNumber="519">MIMB 42256</specimenCode>
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,
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<date box="[612,757,835,856]" pageId="16" pageNumber="519" value="2007-05-23">
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<collectingDate box="[612,757,835,856]" pageId="16" pageNumber="519" value="2007-05-23">23 May 2007</collectingDate>
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</date>
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, among dead corals,
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<elevation box="[383,536,865,887]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="16" pageNumber="519" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">
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<quantity box="[383,457,865,887]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="16" pageNumber="519" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
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depth
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</elevation>
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,
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<location LSID="urn:lsid:plazi:treatment:038A8132EE7BED56FF14CC4EFBA8FEEA:8EFC66FFEE7BED56FDCFCD1CFD0FFCCB" box="[550,760,865,887]" country="Vietnam" latitude="15.383333" longLatPrecision="1285" longitude="109.083336" name="Cu-Lao Re Island" pageId="16" pageNumber="519">Cù-Lao Ré Island</location>
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(
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<geoCoordinate box="[153,250,895,918]" degrees="15" direction="north" minutes="23" orientation="latitude" pageId="16" pageNumber="519" precision="925" value="15.383333">15°23′N</geoCoordinate>
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,
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<geoCoordinate box="[265,375,895,918]" degrees="109" direction="east" minutes="05" orientation="longitude" pageId="16" pageNumber="519" precision="925" value="109.083336">109°05′E</geoCoordinate>
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),
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<collectingCountry box="[398,504,896,918]" name="Vietnam" pageId="16" pageNumber="519">Vietnam</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3897229406" collectingDate="2007-05-19" country="Vietnam" elevation="6" latitude="20.15" location="Bach Long Vi Island" longLatPrecision="1268" longitude="107.73333" pageId="16" pageNumber="519" specimenCount="1">
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<typeStatus box="[517,704,896,918]" isOtherTypeStatus="true" pageId="16" pageNumber="519">Other material</typeStatus>
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:
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<specimenCount pageId="16" pageNumber="519" type="generic">two specimens</specimenCount>
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,
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<date box="[268,406,927,948]" pageId="16" pageNumber="519" value="2007-05-19">
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<collectingDate box="[268,406,927,948]" pageId="16" pageNumber="519" value="2007-05-19">19 May 2007</collectingDate>
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</date>
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, among dead corals,
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<elevation box="[622,757,926,948]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="519" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">
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||
<quantity box="[622,688,926,948]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="519" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">5–7 m</quantity>
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depth
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</elevation>
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,
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||
<location LSID="urn:lsid:plazi:treatment:038A8132EE7BED56FF14CC4EFBA8FEEA:8EFC66FFEE7BED56FF78CDC0FE81FC6F" box="[145,374,957,979]" country="Vietnam" latitude="20.15" longLatPrecision="1268" longitude="107.73333" name="Bach Long Vi Island" pageId="16" pageNumber="519">Bach Long Vi Island</location>
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||
(
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||
<geoCoordinate box="[389,479,956,979]" degrees="20" direction="north" minutes="09" orientation="latitude" pageId="16" pageNumber="519" precision="925" value="20.15">20°09′N</geoCoordinate>
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,
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<geoCoordinate box="[491,593,956,979]" degrees="107" direction="east" minutes="44" orientation="longitude" pageId="16" pageNumber="519" precision="925" value="107.73333">107°44′E</geoCoordinate>
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),
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<collectingCountry box="[612,710,957,979]" name="Vietnam" pageId="16" pageNumber="519">Vietnam</collectingCountry>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3897229402" collectingDate="2010-05-22" country="Vietnam" latitude="12.666667" location="Van Phong Bay" longLatPrecision="1292" longitude="109.333336" pageId="16" pageNumber="519" specimenCount="1">
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||
<specimenCount pageId="16" pageNumber="519" type="generic">one specimen</specimenCount>
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||
,
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||
<date box="[266,417,988,1009]" pageId="16" pageNumber="519" value="2010-05-22">
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<collectingDate box="[266,417,988,1009]" pageId="16" pageNumber="519" value="2010-05-22">22 May 2010</collectingDate>
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</date>
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, intertidal, among calcareous red algae,
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||
<location LSID="urn:lsid:plazi:treatment:038A8132EE7BED56FF14CC4EFBA8FEEA:8EFC66FFEE7BED56FEE8CD86FE5BFBAC" box="[257,428,1018,1040]" country="Vietnam" latitude="12.666667" longLatPrecision="1292" longitude="109.333336" name="Van Phong Bay" pageId="16" pageNumber="519">Van Phong Bay</location>
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||
(
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<geoCoordinate box="[441,531,1017,1040]" degrees="12" direction="north" minutes="40" orientation="latitude" pageId="16" pageNumber="519" precision="925" value="12.666667">12°40′N</geoCoordinate>
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,
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<geoCoordinate box="[541,642,1017,1040]" degrees="109" direction="east" minutes="20" orientation="longitude" pageId="16" pageNumber="519" precision="925" value="109.333336">109°20′E</geoCoordinate>
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),
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<collectingCountry box="[660,759,1018,1040]" name="Vietnam" pageId="16" pageNumber="519">Vietnam</collectingCountry>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph blockId="16.[143,760,1081,1164]" pageId="16" pageNumber="519">
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<emphasis box="[145,256,1081,1102]" italics="true" pageId="16" pageNumber="519">Sequence:</emphasis>
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From the
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<typeStatus box="[376,475,1081,1103]" pageId="16" pageNumber="519">holotype</typeStatus>
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;
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<accessionNumber box="[484,607,1081,1103]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KU748595" pageId="16" pageNumber="519">KU748595</accessionNumber>
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, 16S (524 bp), determined by
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<bibRefCitation author="Hiebert TC" box="[330,507,1112,1134]" pageId="16" pageNumber="519" refId="ref37105" refString="Hiebert TC. 2016. New nemertean diversity discovered in the Northeast Pacific, using surveys of both planktonic larvae and benthic adults. Unpublished D. Phil. Thesis, University of Oregon." type="book" year="2016">Hiebert (2016)</bibRefCitation>
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as Oxypolellinae sp. TCH-2017 isolate E4H7.
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</paragraph>
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<paragraph blockId="16.[145,764,1205,1288]" pageId="16" pageNumber="519">
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<emphasis box="[145,280,1205,1226]" italics="true" pageId="16" pageNumber="519">Etymology:</emphasis>
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The new specific name, a noun in the genitive case, honours Dr Terra C. Hiebert for her outstanding contribution to nemertean systematics.
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</paragraph>
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<paragraph blockId="16.[145,762,1329,1627]" pageId="16" pageNumber="519">
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<emphasis box="[145,275,1330,1351]" italics="true" pageId="16" pageNumber="519">Description:</emphasis>
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Body soft,
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<quantity box="[399,478,1329,1351]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="16" pageNumber="519" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
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long,
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||
<quantity box="[541,604,1329,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="519" unit="mm" value="1.0">1 mm</quantity>
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||
wide, dorsally yellow or lemon yellow, ventrally pale yellowish; head long, narrow, with two thin transverse furrows in front of mouth; eyes and caudal cirrus absent (
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<figureCitation box="[658,745,1421,1443]" captionStart="Figure 2" captionStartId="11.[168,247,1520,1542]" captionTargetBox="[328,1288,196,1480]" captionTargetId="figure-25@11.[328,1288,196,1480]" captionTargetPageId="11" captionText="Figure 2. A, Eopilidion misakiense gen. et sp. nov., holotype, ICHUM 6303, lateral view, head to the right; B, Oxypolella hiebertae sp. nov., holotype, MIMB 42256; C, Cephalomastax brevis Iwata, 1957, ICHUM 6304; D, E, Valencinura jambio sp. nov., holotype, ICHUM 6305, anterior body fragment (D) and enlargement of intestinal region (E) showing yellowish proboscis (indicated by arrowheads); F, Baseodiscus takakurai Gibson, 1995, ICHUM 6306; G, Baseodiscus profundus sp. nov., holotype, MIMB 42257; H, I, Baseodiscus narusei sp. nov., holotype, ICHUM 6310, anterior end of body, dorsal (H) and ventral (I) views; J, Baseodiscus paracelensis sp. nov., holotype, MIMB 33132; K, Baseodiscus komatsui sp. nov., holotype, NMNS-Ne 1; L, Baseodiscus unicolor Stiasny-Wijnhoff, 1925; M, Baseodiscus giribeti sp. nov., holotype, MCZ IZ-135324; N, Baseodiscus cf. amboinensis (Staub, 1900); O, Baseodiscus hemprichii (Ehrenberg, 1828-1831), specimen from Okinawa, different from any voucher specimens in this study; P, Baseodiscus zebra sp. nov., holotype, RUMF-ZN-00001; Q, Baseodiscus quinquelineatus (Quoy & Gaimard, 1833), ICHUM 6319. Photographs by H. Kajihara (A, C, D, E, F, O), A.V. Chernyshev (B, G, J), T. Naruse (H, I), H. Komatsu (K), G. Giribet (L, M), D. Uyeno (P), and R.Yoshida (Q)." figureDoi="http://doi.org/10.5281/zenodo.7037964" httpUri="https://zenodo.org/record/7037964/files/figure.png" pageId="16" pageNumber="519">Fig. 2B</figureCitation>
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). Rhynchopore ventral. Mouth small, pit-like. Rhynchocoel extending length of body. Two regions detected in proboscis of live specimens. Frontal organ present (from CSLSM observation; data not shown). Body wall with two dorsal nerves (from CSLSM observation; data not shown). Living worms with distinctive, permanent mucous tube.
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||
</paragraph>
|
||
<paragraph blockId="16.[145,760,1673,1695]" box="[145,760,1673,1695]" pageId="16" pageNumber="519">
|
||
<emphasis box="[145,286,1673,1694]" italics="true" pageId="16" pageNumber="519">Distribution:</emphasis>
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Known only from
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<collectingCountry box="[487,583,1673,1695]" name="Vietnam" pageId="16" pageNumber="519">Vietnam</collectingCountry>
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(present study).
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</paragraph>
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<paragraph blockId="16.[145,761,1746,1891]" lastBlockId="16.[809,1426,197,342]" pageId="16" pageNumber="519">
|
||
<emphasis box="[145,512,1746,1767]" italics="true" pageId="16" pageNumber="519">
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Remarks:
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<taxonomicName authorityName="Kajihara & Abukawa & Chernyshev" authorityYear="2022" box="[269,512,1746,1767]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="hiebertae">Oxypolella hiebertae</taxonomicName>
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</emphasis>
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is the sixth member described in the genus (cf.
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<bibRefCitation author="Cantell C-E" box="[447,599,1777,1799]" pageId="16" pageNumber="519" pagination="123 - 132" refId="ref34723" refString="Cantell C-E. 2005. On the anatomy of a new species of Oxypolella (Nemertini) and further notes on Oxypolella bergendali Cantell 1975. Journal of Natural History 39: 123 - 132." type="journal article" year="2005">Cantell, 2005</bibRefCitation>
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). It resembles the most recently described species,
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<taxonomicName authority="Cantell, 2005" authorityName="Cantell" authorityYear="2005" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="banyulensis">
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<emphasis italics="true" pageId="16" pageNumber="519">Oxypolella banyulensis</emphasis>
|
||
<bibRefCitation author="Cantell C-E" box="[295,455,1838,1860]" pageId="16" pageNumber="519" pagination="123 - 132" refId="ref34723" refString="Cantell C-E. 2005. On the anatomy of a new species of Oxypolella (Nemertini) and further notes on Oxypolella bergendali Cantell 1975. Journal of Natural History 39: 123 - 132." type="journal article" year="2005">Cantell, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Mediterranean, in having yellowish body colour.
|
||
<taxonomicName authorityName="Kajihara & Abukawa & Chernyshev" authorityYear="2022" box="[524,761,1869,1890]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="hiebertae">
|
||
<emphasis box="[524,761,1869,1890]" italics="true" pageId="16" pageNumber="519">Oxypolella hiebertae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName authorityName="Cantell" authorityYear="2005" box="[950,1115,197,219]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="banyulensis">
|
||
<emphasis box="[950,1115,197,219]" italics="true" pageId="16" pageNumber="519">O. banyulensis</emphasis>
|
||
</taxonomicName>
|
||
in having cephalic furrows as well as in the body coloration; in
|
||
<taxonomicName authorityName="Kajihara & Abukawa & Chernyshev" authorityYear="2022" box="[1235,1372,228,249]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="hiebertae">
|
||
<emphasis box="[1235,1372,228,249]" italics="true" pageId="16" pageNumber="519">O. hiebertae</emphasis>
|
||
</taxonomicName>
|
||
, the body is uniformly yellow, whereas in
|
||
<taxonomicName authorityName="Cantell" authorityYear="2005" box="[1248,1419,258,280]" class="Pilidiophora" family="Lineidae" genus="Oxypolella" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="519" phylum="Nemertea" rank="species" species="banyulensis">
|
||
<emphasis box="[1248,1419,258,280]" italics="true" pageId="16" pageNumber="519">O. banyulensis</emphasis>
|
||
</taxonomicName>
|
||
, the anterior third of the body is white and the posterior two-thirds yellowish white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |