927 lines
142 KiB
XML
927 lines
142 KiB
XML
<document id="450EA46FDDADD1595F718F3B93A3F866" ID-CLB-Dataset="25413" ID-DOI="10.11646/zootaxa.4674.2.2" ID-GBIF-Dataset="10b5d8f0-30c1-4227-b17b-fb8d865ddd06" ID-ISSN="1175-5326" ID-Zenodo-Dep="3458925" ID-ZooBank="B5BA6E39-C9F3-486E-863F-5496DD879CAF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569310351136" checkinUser="plazi" docAuthor="Pati, S. K., Sujila, P. S. & Devi, A. R. Sudha" docDate="2019" docId="C55F87B1E03A755DFF3FF8F5FDF4CBAB" docLanguage="en" docName="zootaxa.4674.2.2.pdf" docOrigin="Zootaxa 4674 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcithelphusa tumpikkai Pati & Sujila & Devi 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="212" masterDocId="3966FFC9E03E7554FFA8FFC1FF99CC09" masterDocTitle="Description of a new species of freshwater crab of the genus Arcithelphusa Pati & Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India" masterLastPageNumber="214" masterPageNumber="203" pageNumber="207" updateTime="1698756317107" updateUser="ExternalLinkService">
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<mods:mods id="3A41D8AF1CB028EA75F3DC08A00011E3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E229B63E7C98B34CB1A81C633645DE90">
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<mods:title id="8AF632D23652374538C7F668CBEA6EB3">Description of a new species of freshwater crab of the genus Arcithelphusa Pati & Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India</mods:title>
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</mods:titleInfo>
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<mods:name id="3953B31A80611DDB68F2AC400394660A" type="personal">
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<mods:role id="4D38B3F3B821A93F3679EB12BCE6E885">
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<mods:roleTerm id="384D3C5D6CBCF8B062EE9591D5E5FC3F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="48B02B04FB4EC583A193B82099DAAD09">Pati, S. K.</mods:namePart>
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<mods:name id="4C503A642ECC67AA5547E0B6911EC3FB" type="personal">
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<mods:roleTerm id="9F94BAACB25F2BDB68F8DD488C75F110">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0B04A239973B7D20702C8F8EB9FD0B56">Sujila, P. S.</mods:namePart>
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</mods:name>
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<mods:name id="0AB0D7368127796872A8913831668313" type="personal">
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<mods:roleTerm id="56C78033D0D5434735196A7084E0703C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D6A21A2EB0F27EB1B184B0CE41A83613">Devi, A. R. Sudha</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AD8E40EFA53A3DEC5AA4DBF88FA19177">text</mods:typeOfResource>
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<mods:title id="1B35A57F1F0D91A229A690CED275363F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0DCEC2BCB242B6E48117152F75A1D958">
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<mods:date id="5A2AAA81FF3CE6BDF66D05E827F1560A">2019</mods:date>
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<mods:number id="527E01BA2D9489928A704E8350878ADD">2019-09-24</mods:number>
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<mods:number id="ACA96F02963F0F5B18C2D4199E0C3113">4674</mods:number>
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<mods:detail id="7815E1970C0F717A42174590C872ADBB" type="issue">
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<mods:number id="346C227000A804BE8F4D9D1A85DB2B76">2</mods:number>
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<mods:classification id="0038AD499428035A7DEDAF2A328BF39E">journal article</mods:classification>
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<mods:identifier id="36F9411073873B1102F69DBB02FB566C" type="CLB-Dataset">25413</mods:identifier>
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<mods:identifier id="04DB0CAE5BACE6531783A051F3E44FB6" type="DOI">10.11646/zootaxa.4674.2.2</mods:identifier>
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</mods:mods>
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<treatment id="C55F87B1E03A755DFF3FF8F5FDF4CBAB" ID-GBIF-Taxon="159398119" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB" httpUri="http://treatment.plazi.org/id/C55F87B1E03A755DFF3FF8F5FDF4CBAB" lastPageId="9" lastPageNumber="212" pageId="4" pageNumber="207">
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<subSubSection id="05EC652CE03A7550FF3FF8F5FDB0CB47" box="[151,553,1844,1871]" pageId="4" pageNumber="207" type="nomenclature">
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<paragraph id="4D4936A7E03A7550FF3FF8F5FDB0CB47" blockId="4.[151,553,1844,1906]" box="[151,553,1844,1871]" pageId="4" pageNumber="207">
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<heading id="160181CBE03A7550FF3FF8F5FDB0CB47" bold="true" box="[151,553,1844,1871]" fontSize="11" level="1" pageId="4" pageNumber="207" reason="1">
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<emphasis id="7F82EAB5E03A7550FF3FF8F5FDB0CB47" bold="true" box="[151,553,1844,1871]" pageId="4" pageNumber="207">
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<taxonomicName id="8AF64D24E03A7550FF3FF8F5FE5DCB46" ID-CoL="8KDBG" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[151,452,1844,1871]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
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||
<emphasis id="7F82EAB5E03A7550FF3FF8F5FE5DCB46" bold="true" box="[151,452,1844,1871]" italics="true" pageId="4" pageNumber="207">Arcithelphusa tumpikkai</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="64B157CEE03A7550FE63F8F5FDB0CB47" box="[459,553,1844,1870]" pageId="4" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE03A7550FF3FF899FE6FCB7B" box="[151,502,1879,1906]" pageId="4" pageNumber="207" type="description">
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<paragraph id="4D4936A7E03A7550FF3FF899FE6FCB7B" blockId="4.[151,553,1844,1906]" box="[151,502,1879,1906]" pageId="4" pageNumber="207">
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(
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<figureCitation id="D5CD2A22E03A7550FF37F899FF74CB7B" box="[159,237,1880,1906]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="4" pageNumber="207">Figs. 2</figureCitation>
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||
A–C, 3A–L, 4A–F, 5A)
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="05EC652CE03A7551FF3FF85BFB28C83E" lastPageId="5" lastPageNumber="208" pageId="4" pageNumber="207" type="materials_examined">
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<paragraph id="4D4936A7E03A7550FF3FF85BFF60CBF5" blockId="4.[151,1436,1945,2044]" pageId="4" pageNumber="207">
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<emphasis id="7F82EAB5E03A7550FF3FF85BFEDFCBBD" bold="true" box="[151,326,1945,1972]" pageId="4" pageNumber="207">Type material.</emphasis>
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<materialsCitation id="FD9E3CFAE03A7550FEE5F858FF60CBF5" ID-GBIF-Occurrence="2417904840" collectingDate="2017-09-06" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" elevation="792" latitude="11.869" location="Makkimala" longLatPrecision="77" longitude="75.948" municipality="Wayanad district" pageId="4" pageNumber="207" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
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||
<typeStatus id="924D8805E03A7550FEE5F858FE23CBBA" box="[333,442,1945,1971]" pageId="4" pageNumber="207" type="holotype">Holotype</typeStatus>
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||
: adult male (
|
||
<quantity id="8A0E9B42E03A7550FDE2F85BFD27CBBD" box="[586,702,1946,1972]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="4" pageNumber="207" unit="mm" value="19.4">19.40 mm</quantity>
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||
cw,
|
||
<quantity id="8A0E9B42E03A7550FD5AF85BFCFCCBBD" box="[754,869,1946,1972]" metricMagnitude="-2" metricUnit="m" metricValue="1.393" pageId="4" pageNumber="207" unit="mm" value="13.93">13.93 mm</quantity>
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||
cl,
|
||
<quantity id="8A0E9B42E03A7550FC27F858FB98CBBD" box="[911,1025,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" pageId="4" pageNumber="207" unit="mm" value="12.25">12.25 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03A7550FB9AF858FB0FCBBD" box="[1074,1174,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="4.63" pageId="4" pageNumber="207" unit="mm" value="4.63">4.63 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03A7550FB7BF858FACFCBBA" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03A7550FB7BF858FACFCBBA" box="[1235,1366,1945,1971]" country="India" latitude="11.869" longLatPrecision="77" longitude="75.948" municipality="Wayanad district" name="Makkimala" pageId="4" pageNumber="207" stateProvince="Kerala">Makkimala</location>
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||
,
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||
<collectingMunicipality id="AD2DACDDE03A7550FAC8F85BFEBFCBDE" pageId="4" pageNumber="207">Wayanad district</collectingMunicipality>
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,
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<collectingRegion id="8F32F845E03A7550FE9AF87CFEE6CBDE" box="[306,383,1981,2007]" country="India" name="Kerala" pageId="4" pageNumber="207">Kerala</collectingRegion>
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,
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<collectingCountry id="35E17637E03A7550FE24F87CFE5FCBDE" box="[396,454,1981,2007]" name="India" pageId="4" pageNumber="207">India</collectingCountry>
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(
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<geoCoordinate id="28C25060E03A7550FE7FF87CFDDFCBDE" box="[471,582,1981,2007]" degrees="11.869" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="11.869">11.869°N</geoCoordinate>
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,
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<geoCoordinate id="28C25060E03A7550FDFBF87CFD59CBD1" box="[595,704,1981,2008]" degrees="75.948" direction="east" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="75.948">75.948°E</geoCoordinate>
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; altitude
|
||
<quantity id="8A0E9B42E03A7550FC81F87FFCECCBD1" box="[809,885,1982,2008]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="4" pageNumber="207" unit="m" value="792.0">
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<elevation id="C6DBD194E03A7550FC81F87FFCECCBD1" box="[809,885,1982,2008]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="4" pageNumber="207" unit="m" value="792.0">792 m</elevation>
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</quantity>
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),
|
||
<date id="39481067E03A7550FC23F87CFBC4CBD1" box="[907,1117,1981,2008]" pageId="4" pageNumber="207" value="2017-09-06">
|
||
<collectingDate id="290CE98FE03A7550FC23F87CFBC4CBD1" box="[907,1117,1981,2008]" pageId="4" pageNumber="207" value="2017-09-06">6 September 2017</collectingDate>
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||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03A7550FB0BF87FFA8ECBDE" box="[1187,1303,1981,2008]" pageId="4" pageNumber="207">P.S. Sujila</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03A7550FA8FF87FFA05CBD1" box="[1319,1436,1982,2008]" pageId="4" pageNumber="207">ZSI-WRC</collectionCode>
|
||
C.1816).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03B7551FF6FFF58FED9CD4A" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
<materialsCitation id="FD9E3CFAE03B7551FF6FFF58FEA4CD4A" ID-GBIF-Occurrence="2417904835" collectingDate="2018-09-16" collectionCode="ZSI-WRC, ZSI-, WRC" collectorName="P. S. Sujila" country="India" elevation="729" latitude="11.838" location="Varayal" longLatPrecision="77" longitude="75.901" municipality="Wayanad district" pageId="5" pageNumber="208" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
|
||
<typeStatus id="924D8805E03B7551FF6FFF58FEA1CCBA" box="[199,312,153,179]" pageId="5" pageNumber="208" type="paratype">Paratypes</typeStatus>
|
||
: male (
|
||
<quantity id="8A0E9B42E03B7551FE2FFF58FE63CCBA" box="[391,506,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.909" pageId="5" pageNumber="208" unit="mm" value="19.09">19.09 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FD84FF58FD07CCBA" box="[556,670,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.402" pageId="5" pageNumber="208" unit="mm" value="14.02">14.02 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FD6EFF58FCAECCBA" box="[710,823,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.221" pageId="5" pageNumber="208" unit="mm" value="12.21">12.21 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FCCDFF58FC50CCBA" box="[869,969,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="4.87" pageId="5" pageNumber="208" unit="mm" value="4.87">4.87 mm</quantity>
|
||
fw) and female (
|
||
<quantity id="8A0E9B42E03B7551FB2EFF58FB60CCBA" box="[1158,1273,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.018" pageId="5" pageNumber="208" unit="mm" value="20.18">20.18 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FA83FF58FA04CCBA" box="[1323,1437,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.418" pageId="5" pageNumber="208" unit="mm" value="14.18">14.18 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FF13FF7CFEB7CCDE" box="[187,302,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.223" pageId="5" pageNumber="208" unit="mm" value="12.23">12.23 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FEC8FF7DFE5CCCDE" box="[352,453,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" pageId="5" pageNumber="208" unit="mm" value="4.75">4.75 mm</quantity>
|
||
fw), same collection data as holotype (
|
||
<collectionCode id="2BE7AE62E03B7551FC2CFF7CFC60CCDE" box="[900,1017,189,215]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1817); male (
|
||
<quantity id="8A0E9B42E03B7551FB1BFF7DFABECCDE" box="[1203,1319,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.8579999999999999" pageId="5" pageNumber="208" unit="mm" value="18.58">18.58 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FAF5FF7DFF5ACCF2" metricMagnitude="-2" metricUnit="m" metricValue="1.385" pageId="5" pageNumber="208" unit="mm" value="13.85">13.85 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FF45FF20FEC6CCF2" box="[237,351,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="5" pageNumber="208" unit="mm" value="11.8">11.80 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FE27FF20FE6DCCF2" box="[399,500,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.72" pageId="5" pageNumber="208" unit="mm" value="4.72">4.72 mm</quantity>
|
||
fw) and female (
|
||
<quantity id="8A0E9B42E03B7551FD1EFF21FCB3CCF2" box="[694,810,224,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.795" pageId="5" pageNumber="208" unit="mm" value="17.95">17.95 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FCF7FF20FC48CCF2" box="[863,977,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.317" pageId="5" pageNumber="208" unit="mm" value="13.17">13.17 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FC54FF20FBF7CCF2" box="[1020,1134,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.081" pageId="5" pageNumber="208" unit="mm" value="10.81">10.81 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FB37FF20FA9ACCF2" box="[1183,1283,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.46" pageId="5" pageNumber="208" unit="mm" value="4.46">4.46 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03B7551FAE8FF20FA01CCF2" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FAE8FF20FA01CCF2" box="[1344,1432,225,251]" country="India" latitude="11.838" longLatPrecision="77" longitude="75.901" municipality="Wayanad district" name="Varayal" pageId="5" pageNumber="208" stateProvince="Kerala">Varayal</location>
|
||
,
|
||
<collectingMunicipality id="AD2DACDDE03B7551FF3FFEC4FEC1CD16" box="[151,344,261,287]" pageId="5" pageNumber="208">Wayanad district</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="8F32F845E03B7551FECDFEC4FE2BCD16" box="[357,434,261,287]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
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||
,
|
||
<collectingCountry id="35E17637E03B7551FE17FEC4FE60CD16" box="[447,505,261,287]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
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(
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<geoCoordinate id="28C25060E03B7551FDA1FEC4FDE0CD16" box="[521,633,261,287]" degrees="11.838" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.838">11.838°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FD2DFEC5FD6BCD16" box="[645,754,260,287]" degrees="75.901" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="75.901">75.901°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FCF4FEC4FC3ECD16" box="[860,935,261,287]" metricMagnitude="2" metricUnit="m" metricValue="7.29" pageId="5" pageNumber="208" unit="m" value="729.0">
|
||
<elevation id="C6DBD194E03B7551FCF4FEC4FC3ECD16" box="[860,935,261,287]" metricMagnitude="2" metricUnit="m" metricValue="7.29" pageId="5" pageNumber="208" unit="m" value="729.0">729 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FC15FEC4FB04CD16" box="[957,1181,261,287]" pageId="5" pageNumber="208" value="2018-09-16">
|
||
<collectingDate id="290CE98FE03B7551FC15FEC4FB04CD16" box="[957,1181,261,287]" pageId="5" pageNumber="208" value="2018-09-16">16 September 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FB4BFEC4FACECD16" box="[1251,1367,261,287]" pageId="5" pageNumber="208">P.S. Sujila</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FACFFEC4FA05CD16" box="[1383,1436,261,287]" pageId="5" pageNumber="208">ZSI-</collectionCode>
|
||
<collectionCode id="2BE7AE62E03B7551FF3FFEE8FF4ECD4A" box="[151,215,297,323]" country="Japan" httpUri="http://grbio.org/cool/i1ct-fvht" name="Wildlife Research Center of Kyoto University" pageId="5" pageNumber="208">WRC</collectionCode>
|
||
C.1818)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03B7551FF6FFE8DFDB7CF1E" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
<materialsCitation id="FD9E3CFAE03B7551FF6FFE8DFF6FCDA6" ID-GBIF-Occurrence="2417904842" collectingDate="2017-09-06" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" elevation="792" latitude="11.869" location="Makkimala" longLatPrecision="77" longitude="75.948" municipality="Wayanad district" pageId="5" pageNumber="208" specimenCount="2" specimenCount-male="2" stateProvince="Kerala">
|
||
<emphasis id="7F82EAB5E03B7551FF6FFE8DFE1ACD6E" bold="true" box="[199,387,332,359]" pageId="5" pageNumber="208">Other material.</emphasis>
|
||
<specimenCount id="5BF0FD2EE03B7551FE23FE8CFE7ACD6E" box="[395,483,333,359]" pageId="5" pageNumber="208" type="male">2 males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FE5BFE8CFD2CCD6E" box="[499,693,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.9555" metricValueMax="1.98" metricValueMin="1.9309999999999998" pageId="5" pageNumber="208" unit="mm" value="19.555" valueMax="19.8" valueMin="19.31">19.31–19.80 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FD44FE8CFC35CD6E" box="[748,940,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.4285" metricValueMax="1.439" metricValueMin="1.418" pageId="5" pageNumber="208" unit="mm" value="14.285" valueMax="14.39" valueMin="14.18">14.18–14.39 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FC70FE8CFB0ECD6E" box="[984,1175,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.1195" metricValueMax="1.126" metricValueMin="1.113" pageId="5" pageNumber="208" unit="mm" value="11.195" valueMax="11.26" valueMin="11.13">11.13–11.26 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FB61FE8DFAF4CD6E" box="[1225,1389,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.08" metricValueMin="4.92" pageId="5" pageNumber="208" unit="mm" value="5.0" valueMax="5.08" valueMin="4.92">4.92–5.08 mm</quantity>
|
||
fw) and female (
|
||
<quantity id="8A0E9B42E03B7551FE8BFEB0FE0ECD82" box="[291,407,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.888" pageId="5" pageNumber="208" unit="mm" value="18.88">18.88 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FE63FEB0FDA7CD82" box="[459,574,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.337" pageId="5" pageNumber="208" unit="mm" value="13.37">13.37 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FDC0FEB0FD40CD82" box="[616,729,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.199" pageId="5" pageNumber="208" unit="mm" value="11.99">11.99 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FCA1FEB0FCF7CD82" box="[777,878,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="4.61" pageId="5" pageNumber="208" unit="mm" value="4.61">4.61 mm</quantity>
|
||
fw), same collection data as holotype (
|
||
<collectionCode id="2BE7AE62E03B7551FA8FFEB0FA05CD82" box="[1319,1436,369,395]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1819)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="FD9E3CFAE03B7551FEA9FE54FC7BCDDA" ID-GBIF-Occurrence="2417904837" collectingDate="2017-09-13" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" elevation="792" latitude="11.869" location="Makkimala" longLatPrecision="77" longitude="75.948" municipality="Wayanad district" pageId="5" pageNumber="208" specimenCount="3" specimenCount-male="3" stateProvince="Kerala" typeStatus="holotype">
|
||
<specimenCount id="5BF0FD2EE03B7551FEA9FE54FECFCDA6" box="[257,342,405,431]" pageId="5" pageNumber="208" type="male">3 males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FECDFE54FDBCCDA6" box="[357,549,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.973" metricValueMax="2.102" metricValueMin="1.844" pageId="5" pageNumber="208" unit="mm" value="19.73" valueMax="21.02" valueMin="18.44">18.44–21.02 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FDF0FE55FC8FCDA6" box="[600,790,404,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.4905" metricValueMax="1.579" metricValueMin="1.402" pageId="5" pageNumber="208" unit="mm" value="14.905" valueMax="15.79" valueMin="14.02">14.02–15.79 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FC96FE54FC64CDA6" box="[830,1021,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.1995" metricValueMax="1.302" metricValueMin="1.097" pageId="5" pageNumber="208" unit="mm" value="11.995" valueMax="13.02" valueMin="10.97">10.97–13.02 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FB83FE54FB57CDA6" box="[1067,1230,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="4.875" metricValueMax="4.89" metricValueMin="4.86" pageId="5" pageNumber="208" unit="mm" value="4.875" valueMax="4.89" valueMin="4.86">4.86–4.89 mm</quantity>
|
||
fw), same locality data as
|
||
<typeStatus id="924D8805E03B7551FF46FE78FECACDDA" box="[238,339,441,467]" pageId="5" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
,
|
||
<date id="39481067E03B7551FEF7FE78FDA7CDDA" box="[351,574,441,467]" pageId="5" pageNumber="208" value="2017-09-13">
|
||
<collectingDate id="290CE98FE03B7551FEF7FE78FDA7CDDA" box="[351,574,441,467]" pageId="5" pageNumber="208" value="2017-09-13">13 September 2017</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FD2CFE78FD6ECDDA" box="[644,759,441,467]" pageId="5" pageNumber="208">P.S. Sujila</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FCAFFE78FCE5CDDA" box="[775,892,441,467]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1820)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="FD9E3CFAE03B7551FC58FE78FB03CE12" ID-GBIF-Occurrence="2417904839" collectingDate="2017-09-13" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" elevation="860" latitude="11.859" location="Puthiyidam" longLatPrecision="77" longitude="75.958" pageId="5" pageNumber="208" specimenCount="2" specimenCount-male="2" stateProvince="Kerala">
|
||
<specimenCount id="5BF0FD2EE03B7551FC58FE78FBDECDDA" box="[1008,1095,441,467]" pageId="5" pageNumber="208" type="male">2 males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FBFFFE78FA80CDDA" box="[1111,1305,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.891" metricValueMax="1.978" metricValueMin="1.8039999999999998" pageId="5" pageNumber="208" unit="mm" value="18.91" valueMax="19.78" valueMin="18.04">18.04–19.78 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FAE7FE78FE94CDFE" metricMagnitude="-2" metricUnit="m" metricValue="1.3745" metricValueMax="1.431" metricValueMin="1.318" pageId="5" pageNumber="208" unit="mm" value="13.745" valueMax="14.31" valueMin="13.18">13.18– 14.31 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FE95FE1CFE64CDFE" box="[317,509,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.148" metricValueMax="1.194" metricValueMin="1.1019999999999999" pageId="5" pageNumber="208" unit="mm" value="11.48" valueMax="11.94" valueMin="11.02">11.02–11.94 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FD9BFE1DFD43CDFE" box="[563,730,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" metricValueMax="5.04" metricValueMin="4.58" pageId="5" pageNumber="208" unit="mm" value="4.81" valueMax="5.04" valueMin="4.58">4.58–5.04 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03B7551FCB5FE1CFC38CDFE" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FCB5FE1CFC38CDFE" box="[797,929,477,503]" country="India" latitude="11.859" longLatPrecision="77" longitude="75.958" name="Puthiyidam" pageId="5" pageNumber="208" stateProvince="Kerala">Puthiyidam</location>
|
||
, Wayanad district,
|
||
<collectingRegion id="8F32F845E03B7551FB28FE1CFB54CDFE" box="[1152,1229,477,503]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E03B7551FB74FE1CFA8FCDFE" box="[1244,1302,477,503]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
||
(
|
||
<geoCoordinate id="28C25060E03B7551FA81FE1DFA01CDFE" box="[1321,1432,476,503]" degrees="11.859" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.859">11.859°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FF3FFDC1FE9DCE12" box="[151,260,512,539]" degrees="75.958" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="75.958">75.958°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FEC6FDC0FE23CE12" box="[366,442,513,539]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="5" pageNumber="208" unit="m" value="860.0">
|
||
<elevation id="C6DBD194E03B7551FEC6FDC0FE23CE12" box="[366,442,513,539]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="5" pageNumber="208" unit="m" value="860.0">860 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FE78FDC0FD29CE12" box="[464,688,513,539]" pageId="5" pageNumber="208" value="2017-09-13">
|
||
<collectingDate id="290CE98FE03B7551FE78FDC0FD29CE12" box="[464,688,513,539]" pageId="5" pageNumber="208" value="2017-09-13">13 September 2017</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FD5FFDC0FCF3CE12" box="[759,874,513,539]" pageId="5" pageNumber="208">P.S. Sujila</collectorName>
|
||
<emphasis id="7F82EAB5E03B7551FCDCFDC3FC3ECE12" box="[884,935,513,539]" italics="true" pageId="5" pageNumber="208">et al</emphasis>
|
||
. (
|
||
<collectionCode id="2BE7AE62E03B7551FC16FDC0FBAACE12" box="[958,1075,513,539]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1821)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="FD9E3CFAE03B7551FB0FFDC0FCF6CE49" ID-GBIF-Occurrence="2423311247" pageId="5" pageNumber="208">
|
||
<specimenCount id="5BF0FD2EE03B7551FB0FFDC0FB66CE12" box="[1191,1279,513,539]" pageId="5" pageNumber="208" type="male">3 males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FAB8FDC1FF5ACE36" metricMagnitude="-2" metricUnit="m" metricValue="1.363" metricValueMax="1.545" metricValueMin="1.181" pageId="5" pageNumber="208" unit="mm" value="13.629999999999999" valueMax="15.45" valueMin="11.81">11.81–15.45 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FF5EFDE5FE3FCE36" box="[246,422,548,575]" metricMagnitude="-2" metricUnit="m" metricValue="1.032" metricValueMax="1.165" metricValueMin="0.899" pageId="5" pageNumber="208" unit="mm" value="10.32" valueMax="11.65" valueMin="8.99">8.99–11.65 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FE66FDE4FDE9CE36" box="[462,624,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="7.975" metricValueMax="9.21" metricValueMin="6.74" pageId="5" pageNumber="208" unit="mm" value="7.975" valueMax="9.21" valueMin="6.74">6.74–9.21 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FD37FDE5FCD8CE36" box="[671,833,548,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.6799999999999997" metricValueMax="3.84" metricValueMin="3.52" pageId="5" pageNumber="208" unit="mm" value="3.6799999999999997" valueMax="3.84" valueMin="3.52">3.52–3.84 mm</quantity>
|
||
fw)
|
||
</materialsCitation>
|
||
and
|
||
<materialsCitation id="FD9E3CFAE03B7551FC0CFDE4FA0ECE6A" ID-GBIF-Occurrence="2423311246" collectingDate="2018-09-30" collectingDateMax="2018-10-26" collectingDateMin="2018-09-30" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" county="Wayanad district" elevation="753" latitude="11.869" location="Kappikalam" longLatPrecision="77" longitude="75.942" municipality="Kambamala" pageId="5" pageNumber="208" specimenCount="2" specimenCount-female="2" stateProvince="Kerala" typeStatus="paratype">
|
||
<specimenCount id="5BF0FD2EE03B7551FC0CFDE4FB89CE36" box="[932,1040,549,575]" pageId="5" pageNumber="208" type="female">2 females</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FBB6FDE5FB46CE36" box="[1054,1247,548,575]" metricMagnitude="-2" metricUnit="m" metricValue="1.6515" metricValueMax="1.7449999999999999" metricValueMin="1.558" pageId="5" pageNumber="208" unit="mm" value="16.515" valueMax="17.45" valueMin="15.58">15.58–17.45 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FAB9FDE4FF5ACE6A" metricMagnitude="-2" metricUnit="m" metricValue="1.2395" metricValueMax="1.283" metricValueMin="1.196" pageId="5" pageNumber="208" unit="mm" value="12.395" valueMax="12.83" valueMin="11.96">11.96–12.83 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FF47FD89FE3BCE6A" box="[239,418,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="9.955" metricValueMax="10.33" metricValueMin="9.58" pageId="5" pageNumber="208" unit="mm" value="9.955" valueMax="10.33" valueMin="9.58">9.58–10.33 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FE7DFD88FDE0CE6A" box="[469,633,585,611]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.71" metricValueMin="3.99" pageId="5" pageNumber="208" unit="mm" value="4.35" valueMax="4.71" valueMin="3.99">3.99–4.71 mm</quantity>
|
||
fw), same collection data as preceding
|
||
<typeStatus id="924D8805E03B7551FBE8FD88FB32CE6A" box="[1088,1195,585,611]" pageId="5" pageNumber="208" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FB14FD88FAA8CE6A" box="[1212,1329,585,611]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1822)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="FD9E3CFAE03B7551FF3FFDACFCD2CEC6" ID-GBIF-Occurrence="2423311249" collectingDate="2018-09-30" collectingDateMax="2018-10-26" collectingDateMin="2018-09-30" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" county="Wayanad district" elevation="753" latitude="11.869" location="Kappikalam" longLatPrecision="77" longitude="75.942" municipality="Kambamala" pageId="5" pageNumber="208" specimenCount="1" specimenCount-female="2" stateProvince="Kerala">
|
||
male (
|
||
<quantity id="8A0E9B42E03B7551FF75FDACFEC8CE8E" box="[221,337,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="2.0629999999999997" pageId="5" pageNumber="208" unit="mm" value="20.63">20.63 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FE2DFDADFE6ECE8E" box="[389,503,620,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.544" pageId="5" pageNumber="208" unit="mm" value="15.44">15.44 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FD88FDACFD0ACE8E" box="[544,659,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.2269999999999999" pageId="5" pageNumber="208" unit="mm" value="12.27">12.27 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FD6AFDADFCBECE8E" box="[706,807,620,647]" metricMagnitude="-3" metricUnit="m" metricValue="5.39" pageId="5" pageNumber="208" unit="mm" value="5.39">5.39 mm</quantity>
|
||
fw) and female (
|
||
<quantity id="8A0E9B42E03B7551FC4EFDADFBC3CE8E" box="[998,1114,620,647]" metricMagnitude="-2" metricUnit="m" metricValue="2.335" pageId="5" pageNumber="208" unit="mm" value="23.35">23.35 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FB26FDACFA98CE8E" box="[1166,1281,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.669" pageId="5" pageNumber="208" unit="mm" value="16.69">16.69 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FA82FDACFA05CE8E" box="[1322,1436,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.4609999999999999" pageId="5" pageNumber="208" unit="mm" value="14.61">14.61 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FF16FD51FEB8CEA2" box="[190,289,656,683]" metricMagnitude="-3" metricUnit="m" metricValue="5.72" pageId="5" pageNumber="208" unit="mm" value="5.72">5.72 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03B7551FEF2FD50FE7FCEA2" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FEF2FD50FE7FCEA2" box="[346,486,657,683]" country="India" county="Wayanad district" latitude="11.869" longLatPrecision="77" longitude="75.942" municipality="Kambamala" name="Kappikalam" pageId="5" pageNumber="208" stateProvince="Kerala">Kappikalam</location>
|
||
, near
|
||
<location id="4829607CE03B7551FD8DFD50FD20CEA2" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FD8DFD50FD20CEA2" box="[549,697,657,683]" country="India" county="Wayanad district" latitude="11.869" longLatPrecision="77" longitude="75.942" municipality="Kambamala" name="Thalappuzha" pageId="5" pageNumber="208" stateProvince="Kerala">Thalappuzha</location>
|
||
,
|
||
<location id="4829607CE03B7551FD6AFD50FC19CEA2" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FD6AFD50FC19CEA2" box="[706,896,657,683]" country="India" county="Wayanad district" latitude="11.869" longLatPrecision="77" longitude="75.942" municipality="Kambamala" name="Wayanad district" pageId="5" pageNumber="208" stateProvince="Kerala">Wayanad district</location>
|
||
,
|
||
<collectingRegion id="8F32F845E03B7551FC22FD50FC4ECEA2" box="[906,983,657,683]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E03B7551FC49FD50FB82CEA2" box="[993,1051,657,683]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
||
(
|
||
<geoCoordinate id="28C25060E03B7551FB80FD50FB01CEA2" box="[1064,1176,657,683]" degrees="11.844" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.844">11.844°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FB0AFD51FA96CEA2" box="[1186,1295,656,683]" degrees="75.940" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="75.94">75.940°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FADAFD51FF29CEC6" metricMagnitude="2" metricUnit="m" metricValue="7.529999999999999" pageId="5" pageNumber="208" unit="m" value="753.0">
|
||
<elevation id="C6DBD194E03B7551FADAFD51FF29CEC6" metricMagnitude="2" metricUnit="m" metricValue="7.529999999999999" pageId="5" pageNumber="208" unit="m" value="753.0">753 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FF6EFD74FE3FCEC6" box="[198,422,693,719]" pageId="5" pageNumber="208" value="2018-09-30">
|
||
<collectingDate id="290CE98FE03B7551FF6EFD74FE3FCEC6" box="[198,422,693,719]" pageId="5" pageNumber="208" value="2018-09-30">30 September 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FE43FD74FDC6CEC6" box="[491,607,693,719]" pageId="5" pageNumber="208">P.S. Sujila</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FDC7FD74FD7DCEC6" box="[623,740,693,719]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1823)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="FD9E3CFAE03B7551FCF0FD74FDB2CF1E" ID-GBIF-Occurrence="2423311248" collectingDate="2018-10-26" collectingDateMax="2018-10-26" collectingDateMin="2018-09-30" collectionCode="ZSI-WRC" collectorName="P. S. Sujila" country="India" county="Wayanad district" elevation="816" latitude="11.869" location="Kambamala" longLatPrecision="77" longitude="75.942" municipality="Kambamala" pageId="5" pageNumber="208" specimenCount="1" specimenCount-female="2" stateProvince="Kerala">
|
||
female (
|
||
<quantity id="8A0E9B42E03B7551FC1CFD74FBB3CEC6" box="[948,1066,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="2.023" pageId="5" pageNumber="208" unit="mm" value="20.23">20.23 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FBC8FD74FB4ACEC6" box="[1120,1235,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.484" pageId="5" pageNumber="208" unit="mm" value="14.84">14.84 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FB57FD75FAEACEC6" box="[1279,1395,692,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.355" pageId="5" pageNumber="208" unit="mm" value="13.55">13.55 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FF3FFD19FF62CEFA" box="[151,251,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="5.03" pageId="5" pageNumber="208" unit="mm" value="5.03">5.03 mm</quantity>
|
||
fw),
|
||
<collectingMunicipality id="AD2DACDDE03B7551FE9DFD18FE26CEFA" box="[309,447,729,755]" pageId="5" pageNumber="208">Kambamala</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="A4284E2BE03B7551FE61FD18FD11CEFA" box="[457,648,729,755]" pageId="5" pageNumber="208">Wayanad district</collectingCounty>
|
||
,
|
||
<collectingRegion id="8F32F845E03B7551FD3AFD18FD46CEFA" box="[658,735,729,755]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E03B7551FD42FD18FCBDCEFA" box="[746,804,729,755]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
||
(
|
||
<geoCoordinate id="28C25060E03B7551FC9AFD18FC38CEFA" box="[818,929,729,755]" degrees="11.869" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.869">11.869°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FC04FD19FB80CEFA" box="[940,1049,728,755]" degrees="75.942" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="75.942">75.942°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FBD5FD18FB5ECEFA" box="[1149,1223,729,755]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="5" pageNumber="208" unit="m" value="816.0">
|
||
<elevation id="C6DBD194E03B7551FBD5FD18FB5ECEFA" box="[1149,1223,729,755]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="5" pageNumber="208" unit="m" value="816.0">816 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FB72FD18FA01CEFA" box="[1242,1432,729,755]" pageId="5" pageNumber="208" value="2018-10-26">
|
||
<collectingDate id="290CE98FE03B7551FB72FD18FA01CEFA" box="[1242,1432,729,755]" pageId="5" pageNumber="208" value="2018-10-26">26 October 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FF67FD3CFED8CF1E" box="[207,321,765,791]" pageId="5" pageNumber="208">P.S. Sujila</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FEF8FD3CFE5CCF1E" box="[336,453,765,791]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1824)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03B7551FF6FFCE1FB28C83E" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
<emphasis id="7F82EAB5E03B7551FF6FFCE1FE4FCF32" bold="true" box="[199,470,800,827]" pageId="5" pageNumber="208">Comparative material.</emphasis>
|
||
<taxonomicName id="8AF64D24E03B7551FE74FCE0FD33CF32" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[476,682,801,827]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E03B7551FE74FCE0FD33CF32" box="[476,682,801,827]" italics="true" pageId="5" pageNumber="208">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<materialsCitation id="FD9E3CFAE03B7551FD10FCE0FD56CFAE" ID-GBIF-Occurrence="2417904836" collectingDate="2011-06-25" collectionCode="ZSI-WRC" collectorName="M. K. Smija" country="India" county="Wayanad district" elevation="767" latitude="11.823" location="Ondayangadi" longLatPrecision="77" longitude="76.023" municipality="Mananthavady" pageId="5" pageNumber="208" specimenCount="3" specimenCount-male="3" stateProvince="Kerala" typeStatus="holotype">
|
||
<typeStatus id="924D8805E03B7551FD10FCE0FC83CF32" box="[696,794,801,827]" pageId="5" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
male (
|
||
<quantity id="8A0E9B42E03B7551FCCBFCE0FC4FCF32" box="[867,982,801,827]" metricMagnitude="-2" metricUnit="m" metricValue="3.082" pageId="5" pageNumber="208" unit="mm" value="30.82">30.82 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FBAEFCE0FBEECF32" box="[1030,1143,801,827]" metricMagnitude="-2" metricUnit="m" metricValue="2.493" pageId="5" pageNumber="208" unit="mm" value="24.93">24.93 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FB35FCE0FA97CF32" box="[1181,1294,801,827]" metricMagnitude="-2" metricUnit="m" metricValue="2.282" pageId="5" pageNumber="208" unit="mm" value="22.82">22.82 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FA92FCE1FA04CF32" box="[1338,1437,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" pageId="5" pageNumber="208" unit="mm" value="7.65">7.65 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03B7551FF63FC84FEFACF56" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FF63FC84FEFACF56" box="[203,355,837,863]" country="India" county="Wayanad district" latitude="11.823" longLatPrecision="77" longitude="76.023" municipality="Mananthavady" name="Ondayangadi" pageId="5" pageNumber="208" stateProvince="Kerala">Ondayangadi</location>
|
||
,
|
||
<quantity id="8A0E9B42E03B7551FEC6FC85FE3FCF56" box="[366,422,836,863]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="208" unit="km" value="5.0">5 km</quantity>
|
||
north east of
|
||
<collectingMunicipality id="AD2DACDDE03B7551FD95FC84FD7CCF56" box="[573,741,837,863]" pageId="5" pageNumber="208">Mananthavady</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="A4284E2BE03B7551FD46FC84FC34CF56" box="[750,941,837,863]" pageId="5" pageNumber="208">Wayanad district</collectingCounty>
|
||
,
|
||
<collectingRegion id="8F32F845E03B7551FC1FFC84FB9DCF56" box="[951,1028,837,863]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E03B7551FBA7FC84FBD0CF56" box="[1039,1097,837,863]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
||
(
|
||
<geoCoordinate id="28C25060E03B7551FBFFFC84FB5FCF56" box="[1111,1222,837,863]" degrees="11.823" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.823">11.823°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FB79FC84FAA7CF56" box="[1233,1342,837,863]" degrees="76.023" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="76.023">76.023°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FF3FFCA8FF7BCF8A" box="[151,226,873,899]" metricMagnitude="2" metricUnit="m" metricValue="7.67" pageId="5" pageNumber="208" unit="m" value="767.0">
|
||
<elevation id="C6DBD194E03B7551FF3FFCA8FF7BCF8A" box="[151,226,873,899]" metricMagnitude="2" metricUnit="m" metricValue="7.67" pageId="5" pageNumber="208" unit="m" value="767.0">767 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FF5EFCA9FE16CF8A" box="[246,399,872,899]" pageId="5" pageNumber="208" value="2011-06-25">
|
||
<collectingDate id="290CE98FE03B7551FF5EFCA9FE16CF8A" box="[246,399,872,899]" pageId="5" pageNumber="208" value="2011-06-25">25 June 2011</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FE7BFCA8FDCECF8A" box="[467,599,873,899]" pageId="5" pageNumber="208">M.K. Smija</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FDCEFCA8FD42CF8A" box="[614,731,873,899]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1188);
|
||
<specimenCount id="5BF0FD2EE03B7551FCE4FCA8FB92CF8A" box="[844,1035,873,899]" pageId="5" pageNumber="208" type="male" typeStatus="paratype">3 paratype males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FBB2FCA8FB45CF8A" box="[1050,1244,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="2.9545" metricValueMax="3.223" metricValueMin="2.686" pageId="5" pageNumber="208" unit="mm" value="29.544999999999998" valueMax="32.23" valueMin="26.86">26.86–32.23 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FAB8FCA9FF5ACFAE" metricMagnitude="-2" metricUnit="m" metricValue="2.332" metricValueMax="2.521" metricValueMin="2.143" pageId="5" pageNumber="208" unit="mm" value="23.32" valueMax="25.21" valueMin="21.43">21.43–25.21 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FF41FC4CFE3ECFAE" box="[233,423,909,935]" metricMagnitude="-2" metricUnit="m" metricValue="2.1225" metricValueMax="2.323" metricValueMin="1.922" pageId="5" pageNumber="208" unit="mm" value="21.225" valueMax="23.23" valueMin="19.22">19.22–23.23 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FE7BFC4DFDECCFAE" box="[467,629,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="7.225" metricValueMax="7.59" metricValueMin="6.86" pageId="5" pageNumber="208" unit="mm" value="7.225" valueMax="7.59" valueMin="6.86">6.86–7.59 mm</quantity>
|
||
fw) and
|
||
</materialsCitation>
|
||
<materialsCitation id="FD9E3CFAE03B7551FD7DFC4CFC58CFE6" ID-GBIF-Occurrence="2417904834" collectingDate="2011-06-25" collectionCode="ZSI-WRC" collectorName="M. K. Smija" country="India" county="Wayanad district" elevation="767" latitude="11.823" location="Ondayangadi" longLatPrecision="77" longitude="76.023" municipality="Mananthavady" pageId="5" pageNumber="208" specimenCount="5" specimenCount-male="5" stateProvince="Kerala" typeStatus="paratype">
|
||
<typeStatus id="924D8805E03B7551FD7DFC4CFCACCFAE" box="[725,821,909,935]" pageId="5" pageNumber="208" type="paratype">paratype</typeStatus>
|
||
female (
|
||
<quantity id="8A0E9B42E03B7551FC3CFC4DFB9ECFAE" box="[916,1031,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="2.685" pageId="5" pageNumber="208" unit="mm" value="26.85">26.85 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FB9FFC4DFB31CFAE" box="[1079,1192,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="2.1149999999999998" pageId="5" pageNumber="208" unit="mm" value="21.15">21.15 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FB66FC4DFAA6CFAE" box="[1230,1343,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="1.954" pageId="5" pageNumber="208" unit="mm" value="19.54">19.54 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FAC3FC4CFF5ACFC2" metricMagnitude="-3" metricUnit="m" metricValue="6.86" pageId="5" pageNumber="208" unit="mm" value="6.86">6.86 mm</quantity>
|
||
fw), same collection data as holotype, coll.
|
||
<collectorName id="E0035371E03B7551FD17FC70FCDDCFC2" box="[703,836,945,971]" pageId="5" pageNumber="208">M.K. Smija</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FCFCFC70FC50CFC2" box="[852,969,945,971]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1189);
|
||
<specimenCount id="5BF0FD2EE03B7551FB95FC71FB67CFC2" box="[1085,1278,944,971]" pageId="5" pageNumber="208" type="male" typeStatus="paratype">5 paratype males</specimenCount>
|
||
(
|
||
<quantity id="8A0E9B42E03B7551FAA6FC70FF5ACFE6" metricMagnitude="-2" metricUnit="m" metricValue="2.6625" metricValueMax="2.903" metricValueMin="2.4219999999999997" pageId="5" pageNumber="208" unit="mm" value="26.625" valueMax="29.03" valueMin="24.22">24.22–29.03 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FF5DFC14FE2ACFE6" box="[245,435,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="2.0795" metricValueMax="2.226" metricValueMin="1.9329999999999998" pageId="5" pageNumber="208" unit="mm" value="20.795" valueMax="22.26" valueMin="19.33">19.33–22.26 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FE71FC15FD0ECFE6" box="[473,663,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.8094999999999999" metricValueMax="1.95" metricValueMin="1.669" pageId="5" pageNumber="208" unit="mm" value="18.095" valueMax="19.5" valueMin="16.69">16.69–19.50 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FD6CFC15FCFFCFE6" box="[708,870,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="6.245" metricValueMax="6.87" metricValueMin="5.62" pageId="5" pageNumber="208" unit="mm" value="6.245" valueMax="6.87" valueMin="5.62">5.62–6.87 mm</quantity>
|
||
fw) and
|
||
</materialsCitation>
|
||
<materialsCitation id="FD9E3CFAE03B7551FC6FFC14FB35C83E" ID-GBIF-Occurrence="2417904838" collectingDate="2013-11-16" collectionCode="ZSI-WRC" collectorName="Ammini" country="India" county="Wayanad district" elevation="760" latitude="11.825" location="Ondayangadi" longLatPrecision="77" longitude="76.03" municipality="Mananthavady" pageId="5" pageNumber="208" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
|
||
<typeStatus id="924D8805E03B7551FC6FFC14FBBECFE6" box="[967,1063,981,1007]" pageId="5" pageNumber="208" type="paratype">paratype</typeStatus>
|
||
female (
|
||
<quantity id="8A0E9B42E03B7551FB20FC14FB63CFE6" box="[1160,1274,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="3.206" pageId="5" pageNumber="208" unit="mm" value="32.06">32.06 mm</quantity>
|
||
cw,
|
||
<quantity id="8A0E9B42E03B7551FA84FC14FA04CFE6" box="[1324,1437,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="2.464" pageId="5" pageNumber="208" unit="mm" value="24.64">24.64 mm</quantity>
|
||
cl,
|
||
<quantity id="8A0E9B42E03B7551FF13FC38FEB6C81A" box="[187,303,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="2.263" pageId="5" pageNumber="208" unit="mm" value="22.63">22.63 mm</quantity>
|
||
ch,
|
||
<quantity id="8A0E9B42E03B7551FEC9FC38FE5EC81A" box="[353,455,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="7.09" pageId="5" pageNumber="208" unit="mm" value="7.09">7.09 mm</quantity>
|
||
fw),
|
||
<location id="4829607CE03B7551FDAEFC38FD07C81A" LSID="urn:lsid:plazi:treatment:C55F87B1E03A755DFF3FF8F5FDF4CBAB:4829607CE03B7551FDAEFC38FD07C81A" box="[518,670,1017,1043]" country="India" county="Wayanad district" latitude="11.825" longLatPrecision="77" longitude="76.03" municipality="Mananthavady" name="Ondayangadi" pageId="5" pageNumber="208" stateProvince="Kerala">Ondayangadi</location>
|
||
,
|
||
<quantity id="8A0E9B42E03B7551FD03FC39FD7FC81A" box="[683,742,1016,1043]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="208" unit="km" value="5.0">5 km</quantity>
|
||
north east of
|
||
<collectingMunicipality id="AD2DACDDE03B7551FC2FFC38FBB6C81A" box="[903,1071,1017,1043]" pageId="5" pageNumber="208">Mananthavady</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="A4284E2BE03B7551FB93FC38FB65C81A" box="[1083,1276,1017,1043]" pageId="5" pageNumber="208">Wayanad district</collectingCounty>
|
||
,
|
||
<collectingRegion id="8F32F845E03B7551FAA0FC38FACCC81A" box="[1288,1365,1017,1043]" country="India" name="Kerala" pageId="5" pageNumber="208">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E03B7551FACBFC38FA04C81A" box="[1379,1437,1017,1043]" name="India" pageId="5" pageNumber="208">India</collectingCountry>
|
||
(
|
||
<geoCoordinate id="28C25060E03B7551FF37FBDDFE96C83E" box="[159,271,1052,1079]" degrees="11.825" direction="north" orientation="latitude" pageId="5" pageNumber="208" precision="55" value="11.825">11.825°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28C25060E03B7551FEB2FBDCFE1EC83E" box="[282,391,1053,1079]" degrees="76.030" direction="east" orientation="longitude" pageId="5" pageNumber="208" precision="55" value="76.03">76.030°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="8A0E9B42E03B7551FE45FBDCFDAEC83E" box="[493,567,1053,1079]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="208" unit="m" value="760.0">
|
||
<elevation id="C6DBD194E03B7551FE45FBDCFDAEC83E" box="[493,567,1053,1079]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="208" unit="m" value="760.0">760 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="39481067E03B7551FDE3FBDCFCBFC83E" box="[587,806,1053,1079]" pageId="5" pageNumber="208" value="2013-11-16">
|
||
<collectingDate id="290CE98FE03B7551FDE3FBDCFCBFC83E" box="[587,806,1053,1079]" pageId="5" pageNumber="208" value="2013-11-16">16 November 2013</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0035371E03B7551FCC0FBDCFC5CC83E" box="[872,965,1053,1079]" pageId="5" pageNumber="208">Ammini</collectorName>
|
||
(
|
||
<collectionCode id="2BE7AE62E03B7551FC7BFBDCFBD1C83E" box="[979,1096,1053,1079]" pageId="5" pageNumber="208">ZSI-WRC</collectionCode>
|
||
C.1190)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE03B7551FF6FFB81FB9AC9EE" pageId="5" pageNumber="208" type="diagnosis">
|
||
<paragraph id="4D4936A7E03B7551FF6FFB81FB9AC9EE" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
<emphasis id="7F82EAB5E03B7551FF6FFB81FEDBC852" bold="true" box="[199,322,1088,1115]" pageId="5" pageNumber="208">Diagnosis.</emphasis>
|
||
Carapace small in adults (<
|
||
<quantity id="8A0E9B42E03B7551FD28FB80FD57C852" box="[640,718,1089,1115]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="208" unit="mm" value="24.0">24 mm</quantity>
|
||
cw), transversely ovate, relatively broad (cw/cl = 1.3–1.4), relatively low (ch/cw = 0.6); posterior margin of carapace gently sinuous; cervical grooves shallow, with short course; H-shaped groove barely visible (
|
||
<figureCitation id="D5CD2A22E03B7551FDAAFB48FD1CC8AA" box="[514,645,1161,1187]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="5" pageNumber="208">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03B7551FD38FB48FD2DC8AA" box="[656,692,1161,1187]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">3A</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03B7551FD17FB48FCDCC8AA" box="[703,837,1161,1187]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="5" pageNumber="208">4A, B, D, E</figureCitation>
|
||
). Third maxilliped exopod reaching nearly half length of merus, with short flagellum, reaching half width of merus (
|
||
<figureCitation id="D5CD2A22E03B7551FCCBFB6CFC25C8CE" box="[867,956,1197,1223]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">Fig. 3B</figureCitation>
|
||
). Male pleonal somite 6 as long as telson (
|
||
<figureCitation id="D5CD2A22E03B7551FF37FB10FE9BC8E2" box="[159,258,1233,1259]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="5" pageNumber="208">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03B7551FEA6FB10FEABC8E2" box="[270,306,1233,1259]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">3D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03B7551FE96FB10FEFAC8E2" box="[318,355,1233,1259]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="5" pageNumber="208">4C</figureCitation>
|
||
). Female pleon narrowly ovate; pleonal somite 6 much broader than long, as long as telson (
|
||
<figureCitation id="D5CD2A22E03B7551FAC7FB10FF20C906" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="5" pageNumber="208">Fig. 4F</figureCitation>
|
||
). G1 relatively slender, with sharply bent terminal segment; terminal segment relatively long, ca. 0.4× length of subterminal segment, medially curved outwards at angle of 90° along longitudinal axis, relatively slender proximally, tapering distally, tip truncated, upcurved, resembling uplifted trunk of elephant, dorsal flap distinct but low, broad; subterminal segment relatively slender, outer margin with distinct hump at basal half, inner margin sinuous (
|
||
<figureCitation id="D5CD2A22E03B7551FF08FA44FF79C996" box="[160,224,1413,1439]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">Fig. 3</figureCitation>
|
||
E–G, J). G2 short, ca. 0.6–0.7× length of G1 length; distal segment very short, ca. 0.2× length of basal segment (
|
||
<figureCitation id="D5CD2A22E03B7551FF75FA68FEC1C9CA" box="[221,344,1449,1475]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">Fig. 3H, K</figureCitation>
|
||
). Vulvae orbicular, large, occupying nearly half length of thoracic sternite 6, situated on submedial part of thoracic sternite 6 adjacent to margin with thoracic sternite 5 (
|
||
<figureCitation id="D5CD2A22E03B7551FC37FA0CFC6FC9EE" box="[927,1014,1485,1511]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="5" pageNumber="208">Fig. 3L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE03B7553FF6FFA31FECCC996" lastPageId="7" lastPageNumber="210" pageId="5" pageNumber="208" type="description">
|
||
<paragraph id="4D4936A7E03B7551FF6FFA31FD2CCB9E" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
<emphasis id="7F82EAB5E03B7551FF6FFA31FDBCCA02" bold="true" box="[199,549,1520,1547]" pageId="5" pageNumber="208">
|
||
Description of male
|
||
<typeStatus id="924D8805E03B7551FE1FFA30FDB9CA02" box="[439,544,1521,1547]" pageId="5" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Carapace transversely ovate, broader than long (cw/cl = 1.4), very deep (ch/cw = 0.6); dorsal surface arched, smooth except for antero- and postero-lateral surfaces; anterolateral surface inflated in frontal view; anterolateral margins short, convex, each with 4 or 5 short, cristate lines, curving into epibranchial region; posterolateral margins long, with oblique striae, converging posteriorly, joining gently sinuous posterior carapace margin; front strongly deflexed, almost rectangular, longitudinally broad, surface almost flat; frontal margin almost straight, narrow (fw/cw = 0.25); epigastric cristae only visible as 2 very low, broad, oval protuberances; postorbital cristae indistinct; external orbital angle indistinct; epibranchial tooth indistinct; postorbital region gently concave; branchial regions highly inflated; cervical grooves shallow, with short course; mesogastric groove shallow, narrow, long, bifurcated posteriorly; H-shaped groove barely visible; subhepatic region rugose; suborbital region rugose; suborbital margin concave, lined with very low, rounded granules; pterygostomial region rugose anteriorly; frontal median triangle incomplete, lateral margins indiscernible; epistome posterior margin with triangular median lobe and sinuous lateral lobes (
|
||
<figureCitation id="D5CD2A22E03B7551FE5AF8BCFDA6CB9E" box="[498,575,1917,1943]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="5" pageNumber="208">Figs. 2</figureCitation>
|
||
A–C, 3A).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03B7551FF6FF860FAEECBD6" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="208">
|
||
Eyes small, filling up most of orbital space; eye stalk short, stout; cornea moderately large, pigmented. Antennae very short, reaching up to proximal quarter length of eye stalk; antennules long, folded in narrow fossae (
|
||
<figureCitation id="D5CD2A22E03B7551FABCF804FAF3CBD6" box="[1300,1386,1989,2015]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="5" pageNumber="208">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1989662FE0387552FF3FF8DEFDC0CBE2" ID-DOI="http://doi.org/10.5281/zenodo.3458933" ID-Zenodo-Dep="3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="6" pageNumber="209" startId="6.[151,250,1823,1848]" targetBox="[255,1346,170,1792]" targetPageId="6">
|
||
<paragraph id="4D4936A7E0387552FF3FF8DEFDC0CBE2" blockId="6.[151,1437,1823,2027]" pageId="6" pageNumber="209">
|
||
<emphasis id="7F82EAB5E0387552FF3FF8DEFE8DCB31" bold="true" box="[151,276,1823,1848]" pageId="6" pageNumber="209">FIGURE 3.</emphasis>
|
||
<taxonomicName id="8AF64D24E0387552FEB4F8DEFD8CCB3E" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[284,533,1823,1847]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="209" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E0387552FEB4F8DEFD8CCB3E" box="[284,533,1823,1847]" italics="true" pageId="6" pageNumber="209">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E0387552FDB4F8DEFDF7CB3E" bold="true" box="[540,622,1823,1847]" pageId="6" pageNumber="209">
|
||
<taxonomicNameLabel id="64B157CEE0387552FDB4F8DEFDF7CB3E" box="[540,622,1823,1847]" pageId="6" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L);
|
||
<taxonomicName id="8AF64D24E0387552FD2DF885FCD9CB52" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[645,832,1859,1883]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="209" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E0387552FD2DF885FCD9CB52" box="[645,832,1859,1883]" italics="true" pageId="6" pageNumber="209">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1
|
||
<emphasis id="7F82EAB5E0387552FAFEF84DFA0FCBAA" box="[1366,1430,1932,1955]" italics="true" pageId="6" pageNumber="209">in situ</emphasis>
|
||
; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D4936A7E0397553FF6FFF58FAA6CD4A" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
Mandibular palp 2 segmented; terminal segment bilobed; anterior lobe narrow, long; posterior lobe broad, short, nearly half of anterior lobe. First, second maxillipeds each with long flagellum on exopod. Third maxillipeds cover most of buccal cavity when closed; ischium subrectangular, longer than broad, surface uneven, pitted, with deep, oblique submedial groove; merus subquadrate, slightly broader than long; exopod slender, longer than ischium, reaching nearly half length of merus, with short flagellum, reaching half width of merus (
|
||
<figureCitation id="D5CD2A22E0397553FB28FEE8FA9ACD4A" box="[1152,1283,297,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Figs. 2B, C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FAA6FEE8FAABCD4A" box="[1294,1330,297,323]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFE8CFEB1CDFE" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
Chelipeds almost smooth, asymmetrical, right chela larger (
|
||
<figureCitation id="D5CD2A22E0397553FCDEFE8CFC22CD6E" box="[886,955,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Fig. 2</figureCitation>
|
||
A–C). Major cheliped with multiple blunt teeth of different size on each finger, distinct gape when fingers closed; dactylus gently curved, longer than upper margin of palm, smooth; palm as long as high, inner and outer surfaces smooth; carpus slightly rugose on outer surface, inner distal major tooth short, stout, blunt, sub-basal granule distinct, low; merus rugose on outer surface (
|
||
<figureCitation id="D5CD2A22E0397553FF08FE1CFF7BCDFE" box="[160,226,477,503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Fig. 2</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFDC0FB0CCE8E" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
Ambulatory legs slender, long; p3 longest, nearly twice length of carapace; p5 shortest; merus (p2–p5) slender, ca. 3 times long as broad, slightly rugose externally, lacking chitinous spines on margins; p3 propodus ca. 2.5 times long as broad, with few short, sharp, chitinous spines along posterior margin; dactylus (p2–p5) longer than propodus, with short, sharp, chitinous spines on both anterior and posterior margins (
|
||
<figureCitation id="D5CD2A22E0397553FBA5FDACFBD6CE8E" box="[1037,1103,621,647]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Fig. 2</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFD50FE11CF32" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
Thoracic sternites smooth, scattered with short setae throughout; suture between thoracic sternites s2/s3 visible as shallow, broad groove, not reaching edge of sternum; suture between thoracic sternites s3/s4 deep, broad, reaching edge of sternum; suture between thoracic sternites s4/s5, s5/s6, s6/s7, s7/s8 shallow, medially interrupted (
|
||
<figureCitation id="D5CD2A22E0397553FF37FD3CFE9BCF1E" box="[159,258,765,791]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FEA6FD3CFEDECF1E" box="[270,327,765,791]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">3C, I</figureCitation>
|
||
). Pleonal locking mechanism with low, anteriorly directed tubercles on each antero-submedial part of sternite 5 (
|
||
<figureCitation id="D5CD2A22E0397553FE84FCE0FEE3CF32" box="[300,378,801,827]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">
|
||
Fig.
|
||
<date id="39481067E0397553FEC9FCE0FEE3CF32" box="[353,378,801,827]" pageId="7" pageNumber="210">3I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFC84FE3AC83E" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
Sternopleonal cavity deep, long, reaching imaginary line joining medial part of cheliped coxae (
|
||
<figureCitation id="D5CD2A22E0397553FB59FC84FAD0CF56" box="[1265,1353,837,863]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Fig. 2C</figureCitation>
|
||
). Pleon narrowly triangular, lateral margins concave; pleonal somites 1–6 trapezoidal, progressive longer; pleonal somite 1 with convex lateral margins; pleonal somite 2 with distally divergent lateral margins; pleonal somite 3 broadest among pleonal somites, with distally convergent lateral margins; pleonal somites 4, 5 distinctly broad, with distally convergent lateral margins; pleonal somite 6 broader than long, as long as telson, with slightly convex lateral margins (
|
||
<figureCitation id="D5CD2A22E0397553FF7FFC38FEA2C81A" box="[215,315,1017,1043]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FEE1FC38FEF6C81A" box="[329,367,1017,1043]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">3D</figureCitation>
|
||
). Telson narrow, almost as long as broad at base, with strongly concave lateral margins; apex rounded (
|
||
<figureCitation id="D5CD2A22E0397553FEAAFBDCFEFDC83E" box="[258,356,1053,1079]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FED8FBDCFE0FC83E" box="[368,406,1053,1079]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFB80FD6FC93A" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
G1 relatively slender, straight except for sharply bent terminal segment, i.e., medially curved outwards at angle of 90° along longitudinal axis, tip not reaching pleonal locking structure; flexible zone large, with small demarcation of joint between terminal and subterminal segments; terminal segment relatively long, ca. 0.4× length of subterminal segment, relatively slender proximally, tapering distally, tip truncated, upcurved, resembling uplifted trunk of elephant, proximal half with distinct but low, broad dorsal flap; subterminal segment relatively slender, outer margin with distinct hump at basal half, inner margin sinuous (
|
||
<figureCitation id="D5CD2A22E0397553FCA9FB34FCDBC906" box="[769,834,1269,1295]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">Fig. 3</figureCitation>
|
||
E–G). G2 short, ca. 0.7× length of G1; distal segment very short, ca. 0.2× length of basal segment (
|
||
<figureCitation id="D5CD2A22E0397553FD27FAD8FD70C93A" box="[655,745,1305,1331]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">Fig. 3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FFAFDFECCC996" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
<emphasis id="7F82EAB5E0397553FF6FFAFDFECCC95E" bold="true" box="[199,341,1340,1367]" pageId="7" pageNumber="210">Live colour.</emphasis>
|
||
The dorsal surface of the carapace is purplish brown anteriorly and light brown posteriorly (
|
||
<figureCitation id="D5CD2A22E0397553FAC7FAFCFF24C972" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="7" pageNumber="210">Fig. 5A</figureCitation>
|
||
). The lateral and ventral surfaces are light brown (
|
||
<figureCitation id="D5CD2A22E0397553FD4FFAA0FCD9C972" box="[743,832,1376,1403]" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="7" pageNumber="210">Fig. 5A</figureCitation>
|
||
). Chelipeds and ambulatory legs are brown to purplish brown (
|
||
<figureCitation id="D5CD2A22E0397553FF46FA44FED1C996" box="[238,328,1412,1439]" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="7" pageNumber="210">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE0397553FF6FFA68FAC7CA26" pageId="7" pageNumber="210" type="etymology">
|
||
<paragraph id="4D4936A7E0397553FF6FFA68FE1ACA02" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
<emphasis id="7F82EAB5E0397553FF6FFA68FED7C9CA" bold="true" box="[199,334,1449,1475]" pageId="7" pageNumber="210">Etymology.</emphasis>
|
||
The species name ‘tumpikkai’ is derived from the Malayalam language for elephant trunk, referring to the G1 terminal segment that resembles an uplifted trunk of an elephant, the state animal of
|
||
<collectingRegion id="8F32F845E0397553FB44FA0CFAA3C9EE" box="[1260,1338,1485,1511]" country="India" name="Kerala" pageId="7" pageNumber="210">Kerala</collectingRegion>
|
||
. Used as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E0397553FF6FF9D4FAC7CA26" blockId="7.[151,1437,153,2016]" box="[199,1374,1557,1583]" pageId="7" pageNumber="210">The new species is popular among the tribal people of Wayanad as ‘Kunjan Njandu’ meaning small crab.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE039755DFF6FF9F8FDA3CB50" lastPageId="9" lastPageNumber="212" pageId="7" pageNumber="210" type="discussion">
|
||
<paragraph id="4D4936A7E0397553FF6FF9F8FDAACB7A" blockId="7.[151,1437,153,2016]" pageId="7" pageNumber="210">
|
||
<emphasis id="7F82EAB5E0397553FF6FF9F8FEA2CA5A" bold="true" box="[199,315,1593,1619]" pageId="7" pageNumber="210">Remarks.</emphasis>
|
||
The adult male
|
||
<typeStatus id="924D8805E0397553FE46F9F8FDC0CA5A" box="[494,601,1593,1619]" pageId="7" pageNumber="210" type="paratype">paratypes</typeStatus>
|
||
resemble the
|
||
<typeStatus id="924D8805E0397553FD5CF9F8FCCFCA5A" box="[756,854,1593,1619]" pageId="7" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
in overall carapace morphology and gonopod structure. The following differences, however, are noticed: the carapace is relatively less broad (cw/cl = 1.3) in the smaller male
|
||
<typeStatus id="924D8805E0397553FE98F940FE09CA92" box="[304,400,1665,1691]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
(ZSI-WRC C.1818) (
|
||
<figureCitation id="D5CD2A22E0397553FD2FF940FD79CA92" box="[647,736,1665,1691]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="7" pageNumber="210">Fig. 4A</figureCitation>
|
||
) [vs. relatively more broad (cw/cl = 1.4) in the
|
||
<typeStatus id="924D8805E0397553FB55F940FAC6CA92" box="[1277,1375,1665,1691]" pageId="7" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="D5CD2A22E0397553FAC7F940FF22CAB6" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Fig. 2A</figureCitation>
|
||
) and the larger male
|
||
<typeStatus id="924D8805E0397553FE0AF965FD90CAB6" box="[418,521,1700,1727]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
]; the suture between the thoracic sternites s2/s3 is relatively deep in the larger male
|
||
<typeStatus id="924D8805E0397553FF3FF908FF6ECAEA" box="[151,247,1737,1763]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
(ZSI-WRC C.1817) [vs. shallow in the
|
||
<typeStatus id="924D8805E0397553FD1BF908FC8CCAEA" box="[691,789,1737,1763]" pageId="7" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
and the smaller male
|
||
<typeStatus id="924D8805E0397553FBAFF908FBFECAEA" box="[1031,1127,1737,1763]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="D5CD2A22E0397553FBDEF908FB4ECAEA" box="[1142,1239,1737,1763]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="7" pageNumber="210">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FB4AF908FA9CCAEA" box="[1250,1285,1737,1763]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">3C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E0397553FAB8F909FAAFCAEA" box="[1296,1334,1736,1763]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="7" pageNumber="210">4C</figureCitation>
|
||
)]; the G2 is relatively short, ca. 0.6 times the length of the G
|
||
<quantity id="8A0E9B42E0397553FD75F92CFC9FCB0E" box="[733,774,1773,1799]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="210" unit="in" value="1.0">1 in</quantity>
|
||
the larger male
|
||
<typeStatus id="924D8805E0397553FC68F92CFBB9CB0E" box="[960,1056,1773,1799]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
[vs. relatively long, ca. 0.7 times the length of the G
|
||
<quantity id="8A0E9B42E0397553FED8F8D0FE01CB22" box="[368,408,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="210" unit="in" value="1.0">1 in</quantity>
|
||
the
|
||
<typeStatus id="924D8805E0397553FE61F8D0FDB2CB22" box="[457,555,1809,1835]" pageId="7" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
and the smaller male
|
||
<typeStatus id="924D8805E0397553FC8DF8D0FC1CCB22" box="[805,901,1809,1835]" pageId="7" pageNumber="210" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="D5CD2A22E0397553FC3EF8D0FBD7CB22" box="[918,1102,1808,1835]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="7" pageNumber="210">Fig. 3E, H, J, K</figureCitation>
|
||
)]. The adult female
|
||
<typeStatus id="924D8805E0397553FA99F8D0FA05CB22" box="[1329,1436,1809,1835]" pageId="7" pageNumber="210" type="paratype">paratypes</typeStatus>
|
||
(ZSI-WRC C.1817, 1818) are similar to the
|
||
<typeStatus id="924D8805E0397553FD2CF8F4FD7FCB46" box="[644,742,1845,1871]" pageId="7" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
in the non-sexual characters except for the comparatively less asymmetrical chelipeds (
|
||
<figureCitation id="D5CD2A22E0397553FE06F898FE69CB7A" box="[430,496,1881,1907]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="7" pageNumber="210">Fig. 4</figureCitation>
|
||
D–F).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E039755CFF6FF8BCFDAACD6E" blockId="7.[151,1437,153,2016]" lastBlockId="8.[151,1437,153,359]" lastPageId="8" lastPageNumber="211" pageId="7" pageNumber="210">
|
||
<taxonomicName id="8AF64D24E0397553FF6FF8BCFE40CB9E" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[199,473,1917,1943]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E0397553FF6FF8BCFE40CB9E" box="[199,473,1917,1943]" italics="true" pageId="7" pageNumber="210">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E0397553FE49F8BCFDA2CB9E" bold="true" box="[481,571,1917,1943]" pageId="7" pageNumber="210">
|
||
<taxonomicNameLabel id="64B157CEE0397553FE49F8BCFDA2CB9E" box="[481,571,1917,1943]" pageId="7" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="8AF64D24E0397553FD11F8BCFC13CB9E" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[697,906,1917,1943]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E0397553FD11F8BCFC13CB9E" box="[697,906,1917,1943]" italics="true" pageId="7" pageNumber="210">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
and species of
|
||
<taxonomicName id="8AF64D24E0397553FB91F8BCFB63CB9E" authorityName="Alcock" authorityYear="1909" box="[1081,1274,1917,1943]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E0397553FB91F8BCFB63CB9E" box="[1081,1274,1917,1943]" italics="true" pageId="7" pageNumber="210">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
mainly due to the presence of a relatively deep carapace (ch/cw = 0.5–0.7), a strongly convex dorsal surface, the highly inflated branchial regions, a narrow frontal margin (ca. 0.25 times the carapace width), an incomplete frontal median tri- angle, the prominent and triangular median lobe on the epistome posterior margin, a narrowly triangular male pleon, and a short G2, with a short distal segment (ca. 0.2–0.3 times the length of the basal segment) (
|
||
<figureCitation id="D5CD2A22E036755CFB60FF7CFA8DCCDE" box="[1224,1300,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2</figureCitation>
|
||
A–F, 3D, E, H, J, K, O, P, 4A–E; see
|
||
<bibRefCitation id="29674B56E036755CFE0FFF20FDECCCF2" author="Bahir, M. M. & Yeo, D. C. J." box="[423,629,225,251]" pageId="8" pageNumber="211" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: figs. 4A, D, 5A–C;
|
||
<bibRefCitation id="29674B56E036755CFCF1FF20FB9DCCF2" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[857,1028,225,251]" pageId="8" pageNumber="211" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E036755CFC23FF23FC22CCF2" box="[907,955,225,251]" italics="true" pageId="8" pageNumber="211">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 11a–c, h, 12a, d, 13a–c, h, 14a, d). Both
|
||
<taxonomicName id="8AF64D24E036755CFF51FEC4FE1FCD16" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[249,390,261,287]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E036755CFF51FEC4FE1FCD16" box="[249,390,261,287]" italics="true" pageId="8" pageNumber="211">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E036755CFE25FEC4FE7FCD16" bold="true" box="[397,486,261,287]" pageId="8" pageNumber="211">
|
||
<taxonomicNameLabel id="64B157CEE036755CFE25FEC4FE7FCD16" box="[397,486,261,287]" pageId="8" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="8AF64D24E036755CFDB4FEC4FD72CD16" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[540,747,261,287]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E036755CFDB4FEC4FD72CD16" box="[540,747,261,287]" italics="true" pageId="8" pageNumber="211">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the species of
|
||
<taxonomicName id="8AF64D24E036755CFB10FEC4FAE0CD16" authorityName="Alcock" authorityYear="1909" box="[1208,1401,261,287]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E036755CFB10FEC4FAE0CD16" box="[1208,1401,261,287]" italics="true" pageId="8" pageNumber="211">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
by a suite of characters, which includes anterolateral margins of the carapace, third maxilliped exopod, chelipeds, and G1 (see Remarks for
|
||
<taxonomicName id="8AF64D24E036755CFE2FFE8CFDBACD6E" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[391,547,333,359]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E036755CFE2FFE8CFDBACD6E" box="[391,547,333,359]" italics="true" pageId="8" pageNumber="211">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="1989662FE036755CFF3FFA4DFC70C9C1" ID-DOI="http://doi.org/10.5281/zenodo.3458935" ID-Zenodo-Dep="3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211" startId="8.[151,250,1420,1445]" targetBox="[151,1435,382,1367]" targetPageId="8">
|
||
<paragraph id="4D4936A7E036755CFF3FFA4DFC70C9C1" blockId="8.[151,1437,1420,1481]" pageId="8" pageNumber="211">
|
||
<emphasis id="7F82EAB5E036755CFF3FFA4DFE8BC9AC" bold="true" box="[151,274,1420,1445]" pageId="8" pageNumber="211">FIGURE 4.</emphasis>
|
||
<taxonomicName id="8AF64D24E036755CFEB0FA4DFD96C9AD" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[280,527,1420,1444]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E036755CFEB0FA4DFD96C9AD" box="[280,527,1420,1444]" italics="true" pageId="8" pageNumber="211">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E036755CFDBCFA4DFDFDC9AD" bold="true" box="[532,612,1420,1444]" pageId="8" pageNumber="211">
|
||
<taxonomicNameLabel id="64B157CEE036755CFDBCFA4DFDFDC9AD" box="[532,612,1420,1444]" pageId="8" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D4936A7E036755CFF6FF9DAFD83CBC8" blockId="8.[151,1437,1563,1985]" pageId="8" pageNumber="211">
|
||
In addition to the above-mentioned similarities, the new species shares the following prominent characters with
|
||
<taxonomicName id="8AF64D24E036755CFF3FF9FEFEFECA50" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[151,359,1599,1625]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E036755CFF3FF9FEFEFECA50" box="[151,359,1599,1625]" italics="true" pageId="8" pageNumber="211">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
: the epigastric cristae are only visible as two low and broad protuberances (
|
||
<figureCitation id="D5CD2A22E036755CFB1EF9FEFAA5CA50" box="[1206,1340,1599,1625]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2A, D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFAEFF9FEFA17CA50" box="[1351,1422,1599,1625]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4A, D</figureCitation>
|
||
); the postorbital cristae, the external orbital angle, and the epibranchial tooth all are indistinct (
|
||
<figureCitation id="D5CD2A22E036755CFB1AF9A2FAA3CA74" box="[1202,1338,1635,1661]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2A, D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFAEEF9A2FA17CA74" box="[1350,1422,1635,1661]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4A, D</figureCitation>
|
||
); the suture between the male thoracic sternites s2/s3 is shallow to deep and broad, which never reaches the edge of the sternum (
|
||
<figureCitation id="D5CD2A22E036755CFE86F96AFE29CACC" box="[302,432,1707,1733]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFE15F96AFE46CACC" box="[445,479,1707,1733]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="8" pageNumber="211">3C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFE45F96AFD8BCACC" box="[493,530,1707,1733]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4C</figureCitation>
|
||
); the suture between the male thoracic sternites s3/s4 is deep and broad, which reaches the edge of the sternum (
|
||
<figureCitation id="D5CD2A22E036755CFDAAF90EFDE7CAE0" box="[514,638,1743,1769]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFD20F90EFD32CAE0" box="[648,683,1743,1769]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="8" pageNumber="211">3C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFD1EF90EFD42CAE0" box="[694,731,1743,1769]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4C</figureCitation>
|
||
); the male sternopleonal cavity extends up to the imaginary line joining the medial part of the cheliped coxae (
|
||
<figureCitation id="D5CD2A22E036755CFD3FF932FC8DCB04" box="[663,788,1779,1805]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFCB7F932FCDACB04" box="[799,835,1779,1805]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4C</figureCitation>
|
||
); the male pleonal somite 6 is trapezoidal and broader than long (
|
||
<figureCitation id="D5CD2A22E036755CFEB9F8D6FE09CB38" box="[273,400,1815,1841]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="8" pageNumber="211">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFE34F8D6FE58CB38" box="[412,449,1815,1841]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="8" pageNumber="211">3D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E036755CFE65F8D6FE6BCB38" box="[461,498,1815,1841]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="8" pageNumber="211">4C</figureCitation>
|
||
); the medial portion of the G1 terminal segment is distinctly curved outwards (
|
||
<figureCitation id="D5CD2A22E036755CFAC7F8D6FE89CB5C" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="8" pageNumber="211">Fig. 3E, F, J, O</figureCitation>
|
||
).
|
||
<taxonomicName id="8AF64D24E036755CFE8DF8FAFDAFCB5C" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[293,566,1851,1877]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E036755CFE8DF8FAFDAFCB5C" box="[293,566,1851,1877]" italics="true" pageId="8" pageNumber="211">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E036755CFD96F8FAFD01CB5C" bold="true" box="[574,664,1851,1877]" pageId="8" pageNumber="211">
|
||
<taxonomicNameLabel id="64B157CEE036755CFD96F8FAFD01CB5C" box="[574,664,1851,1877]" pageId="8" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is nevertheless differentiated from
|
||
<taxonomicName id="8AF64D24E036755CFB8DF8FAFB6FCB5C" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[1061,1270,1851,1877]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E036755CFB8DF8FAFB6FCB5C" box="[1061,1270,1851,1877]" italics="true" pageId="8" pageNumber="211">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
by the characters in the carapace, third maxilliped exopod, male pleonal somite 6, and G1 (
|
||
<tableCitation id="0074031CE036755CFBA9F89EFBCCCB70" box="[1025,1109,1887,1913]" captionStart="TABLE 1" captionStartId="10.[151,239,154,178]" captionTargetPageId="10" captionText="TABLE 1. Comparison between A. tumpikkai sp. nov. and A. cochleariformis." httpUri="http://table.plazi.org/id/1989662FE034755EFF3FFF5BFC44CCB8" pageId="8" pageNumber="211" tableUuid="1989662FE034755EFF3FFF5BFC44CCB8">Table 1</tableCitation>
|
||
). Moreover, the size of adult specimens of
|
||
<taxonomicName id="8AF64D24E036755CFE98F842FE24CB94" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[304,445,1923,1949]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E036755CFE98F842FE24CB94" box="[304,445,1923,1949]" italics="true" pageId="8" pageNumber="211">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E036755CFE6CF842FD84CB94" bold="true" box="[452,541,1923,1949]" pageId="8" pageNumber="211">
|
||
<taxonomicNameLabel id="64B157CEE036755CFE6CF842FD84CB94" box="[452,541,1923,1949]" pageId="8" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is apparently smaller (cw
|
||
<quantity id="8A0E9B42E036755CFCEAF843FB9DCB94" box="[834,1028,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" metricValueMax="2.335" metricValueMin="1.7449999999999999" pageId="8" pageNumber="211" unit="mm" value="20.4" valueMax="23.35" valueMin="17.45">17.45–23.35 mm</quantity>
|
||
; n = 17) than that of
|
||
<taxonomicName id="8AF64D24E036755CFB42F842FF27CBC9" authorityName="Pati & Sudha Devi" authorityYear="2015" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="211" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E036755CFB42F842FF27CBC9" italics="true" pageId="8" pageNumber="211">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
(cw
|
||
<quantity id="8A0E9B42E036755CFF5EF866FE21CBC8" box="[246,440,1959,1985]" metricMagnitude="-2" metricUnit="m" metricValue="2.8225" metricValueMax="3.223" metricValueMin="2.4219999999999997" pageId="8" pageNumber="211" unit="mm" value="28.224999999999998" valueMax="32.23" valueMin="24.22">24.22–32.23 mm</quantity>
|
||
; n = 11).
|
||
</paragraph>
|
||
<caption id="1989662FE037755DFF3FFBB1FC25C8D8" ID-DOI="http://doi.org/10.5281/zenodo.3458937" ID-Zenodo-Dep="3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="9" pageNumber="212" startId="9.[151,250,1136,1161]" targetBox="[151,1435,174,1117]" targetPageId="9">
|
||
<paragraph id="4D4936A7E037755DFF3FFBB1FC25C8D8" blockId="9.[151,1436,1136,1233]" pageId="9" pageNumber="212">
|
||
<emphasis id="7F82EAB5E037755DFF3FFBB1FE8AC880" bold="true" box="[151,275,1136,1161]" pageId="9" pageNumber="212">FIGURE 5.</emphasis>
|
||
Live colouration and habitat of
|
||
<taxonomicName id="8AF64D24E037755DFDF3FBB0FCCAC880" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[603,851,1137,1161]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFDF3FBB0FCCAC880" box="[603,851,1137,1161]" italics="true" pageId="9" pageNumber="212">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFCF1FBB0FC33C880" bold="true" box="[857,938,1137,1161]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFCF1FBB0FC33C880" box="[857,938,1137,1161]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, live crab
|
||
<emphasis id="7F82EAB5E037755DFB86FBB3FBF4C880" box="[1070,1133,1138,1161]" italics="true" pageId="9" pageNumber="212">in situ</emphasis>
|
||
at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D4936A7E037755DFF6FFAE2FDCACA1F" blockId="9.[151,1437,1315,1954]" pageId="9" pageNumber="212">
|
||
<emphasis id="7F82EAB5E037755DFF6FFAE2FE08C937" bold="true" box="[199,401,1315,1342]" pageId="9" pageNumber="212">Ecological notes.</emphasis>
|
||
<taxonomicName id="8AF64D24E037755DFE32FAE5FD34C937" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[410,685,1316,1342]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFE32FAE5FD34C937" box="[410,685,1316,1342]" italics="true" pageId="9" pageNumber="212">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFD1FFAE5FC8BC937" bold="true" box="[695,786,1316,1342]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFD1FFAE5FC8BC937" box="[695,786,1316,1342]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occupies a wide range of habitats such as Shola forests, mountainous streams, natural springs, rice fields, and banana/betel nut/cassava plantations. These crabs live in shallow burrows (<
|
||
<quantity id="8A0E9B42E037755DFEE2FAAAFE16C98F" box="[330,399,1387,1414]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="212" unit="cm" value="15.0">15 cm</quantity>
|
||
deep) under small and moist boulders (<
|
||
<quantity id="8A0E9B42E037755DFCC9FAADFC3CC98F" box="[865,933,1388,1414]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="212" unit="cm" value="40.0">40 cm</quantity>
|
||
diameter) (
|
||
<figureCitation id="D5CD2A22E037755DFB8FFAAAFB39C98F" box="[1063,1184,1387,1414]" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="9" pageNumber="212">Fig. 5B, C</figureCitation>
|
||
). They are also known to inhabit muddy banks along the water channels of paddy fields and various plantations (
|
||
<figureCitation id="D5CD2A22E037755DFB3CFA4EFB77C9A3" box="[1172,1262,1423,1450]" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="9" pageNumber="212">Fig. 5D</figureCitation>
|
||
). The new species seems to be locally abundant in rather elevated areas of Wayanad district.
|
||
<taxonomicName id="8AF64D24E037755DFBA6FA75FAB9C9C7" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[1038,1312,1460,1486]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFBA6FA75FAB9C9C7" box="[1038,1312,1460,1486]" italics="true" pageId="9" pageNumber="212">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFA80FA75FA1BC9C7" bold="true" box="[1320,1410,1460,1486]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFA80FA75FA1BC9C7" box="[1320,1410,1460,1486]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is syntopic with
|
||
<taxonomicName id="8AF64D24E037755DFE9BFA19FD9BC9FB" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[307,514,1496,1522]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E037755DFE9BFA19FD9BC9FB" box="[307,514,1496,1522]" italics="true" pageId="9" pageNumber="212">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AF64D24E037755DFD9EFA19FCA2C9FB" authority="(Roux, 1931)" authorityName="Roux" authorityYear="1931" baseAuthorityName="Roux" baseAuthorityYear="1931" box="[566,827,1496,1522]" class="Malacostraca" family="Gecarcinucidae" genus="Vela" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="carli">
|
||
<emphasis id="7F82EAB5E037755DFD9EFA19FD38C9FB" box="[566,673,1496,1522]" italics="true" pageId="9" pageNumber="212">Vela carli</emphasis>
|
||
(Roux, 1931)
|
||
</taxonomicName>
|
||
at Kambamala, Kappikalam, Puthiyidam, and Varayal, where soft substrates dominate (
|
||
<figureCitation id="D5CD2A22E037755DFDA9FA3AFDDBCA1F" box="[513,578,1531,1558]" captionStart="FIGURE 1" captionStartId="1.[151,250,1897,1922]" captionTargetBox="[176,1411,166,1877]" captionTargetId="figure@1.[176,1411,166,1877]" captionTargetPageId="1" captionText="FIGURE 1. Map showing distribution of Arcithelphusa tumpikkai sp. nov. and A. cochleariformis. Collection/observation sites (1–17): 1, Makkimala (type locality); 2, Kambamala; 3, Puthiyidam; 4, Kappikalam; 5, Varayal; 6, Thirunelly; 7, Bavali; 8, 9, Ondayangadi; 10, Pakkom; 11, Mananthavady; 12, Chettapalam; 13, Ellumannam; 14, Panamaram; 15, Padinjarathara; 16, Kambalakkad; 17, Muthanga." figureDoi="http://doi.org/10.5281/zenodo.3458927" httpUri="https://zenodo.org/record/3458927/files/figure.png" pageId="9" pageNumber="212">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E037755DFF6FF9DEFDA3CB50" blockId="9.[151,1437,1315,1954]" pageId="9" pageNumber="212">
|
||
The ecological requirement of
|
||
<taxonomicName id="8AF64D24E037755DFD8EF9E1FD2CCA33" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[550,693,1568,1594]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFD8EF9E1FD2CCA33" box="[550,693,1568,1594]" italics="true" pageId="9" pageNumber="212">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFD15F9E1FC81CA33" bold="true" box="[701,792,1568,1594]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFD15F9E1FC81CA33" box="[701,792,1568,1594]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, however, is different from
|
||
<taxonomicName id="8AF64D24E037755DFBF1F9E1FAB3CA33" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[1113,1322,1568,1594]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E037755DFBF1F9E1FAB3CA33" box="[1113,1322,1568,1594]" italics="true" pageId="9" pageNumber="212">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
. The new species generally prefers to stay near water bodies or moist places (
|
||
<figureCitation id="D5CD2A22E037755DFC0CF982FBB9CA57" box="[932,1056,1603,1630]" captionStart="FIGURE 5" captionStartId="9.[151,250,1136,1161]" captionTargetBox="[151,1435,174,1117]" captionTargetId="figure@9.[151,1435,174,1117]" captionTargetPageId="9" captionText="FIGURE 5. Live colouration and habitat of Arcithelphusa tumpikkai sp. nov. A, live crab in situ at Makkimala (type locality); B, small boulders (indicated by red arrow) at Makkimala; C, shallow burrow under small and moist boulder at Makkimala; D, burrows (indicated by red arrow) along water channels of rice fields at Varayal." figureDoi="http://doi.org/10.5281/zenodo.3458937" httpUri="https://zenodo.org/record/3458937/files/figure.png" pageId="9" pageNumber="212">Fig. 5C, D</figureCitation>
|
||
) whereas
|
||
<taxonomicName id="8AF64D24E037755DFB3FF985FAF3CA57" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[1175,1386,1604,1630]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E037755DFB3FF985FAF3CA57" box="[1175,1386,1604,1630]" italics="true" pageId="9" pageNumber="212">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
is a more terrestrial crab that can be found adjacent to or away from water bodies (see
|
||
<bibRefCitation id="29674B56E037755DFB91F9A6FAC7CA8B" author="Pati, S. K. & Sudha Devi, A. R." box="[1081,1374,1639,1666]" pageId="9" pageNumber="212" pagination="35" refId="ref8106" refString="Pati, S. K. & Sudha Devi, A. R. (2015 a) Description of a new genus and new species of freshwater crab (Brachyura: Gecarcinucidae) from the Western Ghats, Kerala India. Zoological Studies, 54, 35. https: // doi. org / 10.1186 / s 40555 - 015 - 0112 - 0" type="journal article" year="2015">Pati & Sudha Devi 2015a</bibRefCitation>
|
||
). The difference in the habitat preferences between both the species could be related to the condition of the flagellum on their third maxilliped exopod, which is relatively well-developed in
|
||
<taxonomicName id="8AF64D24E037755DFC30F971FBBFCAC3" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[920,1062,1712,1738]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFC30F971FBBFCAC3" box="[920,1062,1712,1738]" italics="true" pageId="9" pageNumber="212">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFB86F971FB11CAC3" bold="true" box="[1070,1160,1712,1738]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFB86F971FB11CAC3" box="[1070,1160,1712,1738]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and altogether absent in
|
||
<taxonomicName id="8AF64D24E037755DFF3FF915FEF0CAE7" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[151,361,1748,1774]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E037755DFF3FF915FEF0CAE7" box="[151,361,1748,1774]" italics="true" pageId="9" pageNumber="212">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="8AF64D24E037755DFED0F915FE7FCAE7" baseAuthorityName="Roux" baseAuthorityYear="1931" box="[376,486,1748,1774]" class="Malacostraca" family="Gecarcinucidae" genus="Vela" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="carli">
|
||
<emphasis id="7F82EAB5E037755DFED0F915FE7FCAE7" box="[376,486,1748,1774]" italics="true" pageId="9" pageNumber="212">Vela carli</emphasis>
|
||
</taxonomicName>
|
||
is a larger species with a well-developed flagellum on the third maxilliped exopod, and its burrows are deeper and larger than that of the
|
||
<taxonomicName id="8AF64D24E037755DFCACF939FC39CB1B" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[772,928,1784,1810]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E037755DFCACF939FC39CB1B" box="[772,928,1784,1810]" italics="true" pageId="9" pageNumber="212">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
species. The burrows of
|
||
<taxonomicName id="8AF64D24E037755DFB6CF939FA8ECB1B" baseAuthorityName="Roux" baseAuthorityYear="1931" box="[1220,1303,1784,1810]" class="Malacostraca" family="Gecarcinucidae" genus="Vela" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="carli">
|
||
<emphasis id="7F82EAB5E037755DFB6CF939FA8ECB1B" box="[1220,1303,1784,1810]" italics="true" pageId="9" pageNumber="212">V. carli</emphasis>
|
||
</taxonomicName>
|
||
are usually water filled, which can be seen along rice field embankments, margins of natural springs, and water channels of banana/betel nut/coconut plantations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE037755DFF6FF8A2FDF4CBAB" pageId="9" pageNumber="212" type="distribution">
|
||
<paragraph id="4D4936A7E037755DFF6FF8A2FDF4CBAB" blockId="9.[151,1437,1315,1954]" pageId="9" pageNumber="212">
|
||
<emphasis id="7F82EAB5E037755DFF6FF8A2FEF8CB77" bold="true" box="[199,353,1891,1918]" pageId="9" pageNumber="212">Distribution.</emphasis>
|
||
<taxonomicName id="8AF64D24E037755DFECEF8A5FDEFCB77" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[358,630,1892,1918]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="212" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E037755DFECEF8A5FDEFCB77" box="[358,630,1892,1918]" italics="true" pageId="9" pageNumber="212">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E037755DFDD4F8A5FD4CCB77" bold="true" box="[636,725,1892,1918]" pageId="9" pageNumber="212">
|
||
<taxonomicNameLabel id="64B157CEE037755DFDD4F8A5FD4CCB77" box="[636,725,1892,1918]" pageId="9" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is endemic to the Western Ghats and currently known only from Wayanad district of
|
||
<collectingRegion id="8F32F845E037755DFED1F846FE5FCBA8" box="[377,454,1927,1953]" country="India" name="Kerala" pageId="9" pageNumber="212">Kerala</collectingRegion>
|
||
,
|
||
<collectingCountry id="35E17637E037755DFE79F846FD92CBA8" box="[465,523,1927,1953]" name="India" pageId="9" pageNumber="212">India</collectingCountry>
|
||
(
|
||
<figureCitation id="D5CD2A22E037755DFDB2F846FDC5CBAB" box="[538,604,1927,1954]" captionStart="FIGURE 1" captionStartId="1.[151,250,1897,1922]" captionTargetBox="[176,1411,166,1877]" captionTargetId="figure@1.[176,1411,166,1877]" captionTargetPageId="1" captionText="FIGURE 1. Map showing distribution of Arcithelphusa tumpikkai sp. nov. and A. cochleariformis. Collection/observation sites (1–17): 1, Makkimala (type locality); 2, Kambamala; 3, Puthiyidam; 4, Kappikalam; 5, Varayal; 6, Thirunelly; 7, Bavali; 8, 9, Ondayangadi; 10, Pakkom; 11, Mananthavady; 12, Chettapalam; 13, Ellumannam; 14, Panamaram; 15, Padinjarathara; 16, Kambalakkad; 17, Muthanga." figureDoi="http://doi.org/10.5281/zenodo.3458927" httpUri="https://zenodo.org/record/3458927/files/figure.png" pageId="9" pageNumber="212">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |