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<mods:title id="9CE8D84129884C86395E9A9B865807FC">The genus Dedalopterus Sabatinelli &amp; Pontuale, 1998 (Scarabaeidae: Melolonthinae: Leucopholini) in Laos and Vietnam, with description of a new species</mods:title>
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<mods:namePart id="53AF118B892EDA06941F9B771E8DAA58">Pham, Phu Van</mods:namePart>
<mods:affiliation id="7E538D1761BE6BD5CA03274BA8E361D3">Institute of Ecology and Biological resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam. &amp; Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn; Germany.</mods:affiliation>
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<mods:affiliation id="24E45943B6037B04FF9FD6E021D927BF">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn; Germany.</mods:affiliation>
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Key to the species of 
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<emphasis id="2DB2620DFFDCFFDDFE1BFC9EFDEA3D3F" bold="true" box="[413,574,828,855]" italics="true" pageId="10" pageNumber="151">Dedalopterus</emphasis>
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(males only)
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Based on the key of 
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, modified and updated
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<keyLead id="A4371D2AFFDCFFDDFF11FC3DFA483DDC" box="[151,1436,927,949]" pageId="10" pageNumber="151">1 Antennal club less than twice as long as remaining antennomeres................................................ 2</keyLead>
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<keyLead id="A4371D2AFFDCFFDDFF11FC1FFA483DBB" box="[151,1436,957,979]" pageId="10" pageNumber="151">- Antennal club twice or more than twice as long as remaining antennomeres....................................... 6</keyLead>
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<paragraph id="1F79BE1FFFDCFFDDFF11FC78FA483D87" blockId="10.[151,1437,927,1595]" box="[151,1436,986,1008]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FC78FA483D87" box="[151,1436,986,1008]" pageId="10" pageNumber="151">2 Parameres of unequal length............................................................................ 3</keyLead>
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<paragraph id="1F79BE1FFFDCFFDDFF11FC5AFA483A64" blockId="10.[151,1437,927,1595]" box="[151,1436,1015,1037]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FC5AFA483A64" box="[151,1436,1015,1037]" pageId="10" pageNumber="151">- Parameres of equal length.............................................................................. 4</keyLead>
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<keyLead id="A4371D2AFFDCFFDDFF11FBB7FA483A42" box="[151,1436,1045,1067]" pageId="10" pageNumber="151">
3 Protibia bidentate, tegument of elytra light-brown to yellow, parameres short............... 
<taxonomicName id="D8C6C59CFFDCFFDDFB0EFBB7FA483A42" ID-CoL="3XLRZ" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1160,1436,1045,1067]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="2DB2620DFFDCFFDDFB0EFBB7FAD93A42" box="[1160,1293,1045,1066]" italics="true" pageId="10" pageNumber="151">D. fujianensis</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFBB7FA413A42" author="Zhang, Y. W." box="[1306,1429,1045,1067]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
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- Protibia tridentate (basal tooth small), tegument of elytra dark brown, parameres longer.................................................................................................... 
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<emphasis id="2DB2620DFFDCFFDDFBBEFBF2FB553A0C" box="[1080,1153,1103,1125]" italics="true" pageId="10" pageNumber="151">D. itohi</emphasis>
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4 Parameres curved ventrally by 40 
<superScript id="E8B31357FFDCFFDDFDA2FBCEFDFF3A10" attach="left" box="[548,555,1132,1144]" fontSize="5" pageId="10" pageNumber="151">o</superScript>
at distal third (lateral view).......................... 
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<emphasis id="2DB2620DFFDCFFDDFBEFFBCFFB6F3AEA" box="[1129,1211,1133,1154]" italics="true" pageId="10" pageNumber="151">D. fencli</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFB44FBCFFA483AEA" author="Zidek, J. &amp; Krajcik, M." box="[1218,1436,1133,1155]" pageId="10" pageNumber="151" pagination="1 - 14" refId="ref6749" refString="Zidek, J. &amp; Krajcik, M. (2007) Remarks on the genus Dedalopterus (Melolonthinae: Leucopholini), with descriptions of two new species. Animma. x, 21, 1 - 14." type="journal article" year="2007">Zídek &amp; Krajčík, 2007</bibRefCitation>
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<paragraph id="1F79BE1FFFDCFFDDFF11FB28FA483AC8" blockId="10.[151,1437,927,1595]" box="[151,1436,1162,1184]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB28FA483AC8" box="[151,1436,1162,1184]" pageId="10" pageNumber="151">- Parameres entirely straight (lateral view)................................................................... 5</keyLead>
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<paragraph id="1F79BE1FFFDCFFDDFF11FB05FA483AD4" blockId="10.[151,1437,927,1595]" box="[151,1436,1191,1213]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB05FA483AD4" box="[151,1436,1191,1213]" pageId="10" pageNumber="151">
5 Right paramere rather straight and larger.......................................... 
<taxonomicName id="D8C6C59CFFDCFFDDFBEBFB0AFA483AD4" ID-CoL="3XLS4" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1133,1436,1191,1213]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="melanodiscus">
<emphasis id="2DB2620DFFDCFFDDFBEBFB0AFAD93AD4" box="[1133,1293,1191,1213]" italics="true" pageId="10" pageNumber="151">D. melanodiscus</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFB05FA413AD4" author="Zhang, Y. W." box="[1306,1429,1191,1213]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
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<keyLead id="A4371D2AFFDCFFDDFF11FB67FA483AB2" box="[151,1436,1221,1243]" pageId="10" pageNumber="151">
- Right paramere curved moderately dorsally at apex and slender.......................... 
<taxonomicName id="D8C6C59CFFDCFFDDFBFBFB67FA483AB2" ID-CoL="3XLS2" authority="(Zhang, 2002)" baseAuthorityName="Zhang" baseAuthorityYear="2002" box="[1149,1436,1221,1243]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="2DB2620DFFDCFFDDFBFBFB67FAD93AB2" box="[1149,1293,1221,1242]" italics="true" pageId="10" pageNumber="151">D. hainanensis</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFB67FA423AB2" author="Zhang, Y. W." box="[1306,1430,1221,1243]" pageId="10" pageNumber="151" pagination="327 - 336" refId="ref6696" refString="Zhang, Y. W. (2002) Coleoptera: Scarabaeoidea. In: Huang, F. S. (Ed.), Forest Insects of Hainan. Science Press, Beijing, pp. 327 - 336. [in Chinese, with English description of new species]" type="book chapter" year="2002">Zhang, 2002</bibRefCitation>
)
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<paragraph id="1F79BE1FFFDCFFDDFF11FB40FA483A9F" blockId="10.[151,1437,927,1595]" box="[151,1436,1250,1272]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FB40FA483A9F" box="[151,1436,1250,1272]" pageId="10" pageNumber="151">6 Pygidium apically rounded.............................................................................. 7</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FAA2FA483B7C" blockId="10.[151,1437,927,1595]" box="[151,1436,1279,1301]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FAA2FA483B7C" box="[151,1436,1279,1301]" pageId="10" pageNumber="151">- Pygidium apically triangular............................................................................. 8</keyLead>
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</keyStep>
<keyStep id="A432A6BAFFDCFFDDFF11FABFFA483BE2" pageId="10" pageNumber="151">
<paragraph id="1F79BE1FFFDCFFDDFF11FABFFA483B27" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FABFFA483B27" pageId="10" pageNumber="151">
7 Parameres rather straight and long, right paramere width almost equal to the left one (see 
<figureCitation id="87FDA29AFFDCFFDDFBEAFABFFB7D3B5A" box="[1132,1193,1309,1331]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[323,1265,358,1905]" captionTargetId="figure-83@4.[310,1278,345,1905]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Dedalopterus lexuanhuei Pham, Ahrens and Sabatinelli, new species (holotype,A–C); D. malyszi Bunalski, 2001 (D–F). A, D, parameres in left lateral view; B, E, parameres in dorsal view. C, F, parameres in right lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611473" httpUri="https://zenodo.org/record/10611473/files/figure.png" pageId="10" pageNumber="151">Fig. 3</figureCitation>
; 
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).......................................................................... 
<taxonomicName id="D8C6C59CFFDCFFDDFB27FA99FA483B27" ID-CoL="6C8Y5" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[1185,1436,1338,1360]" class="Insecta" family="Scarabaeidae" genus="Cyphochilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="2DB2620DFFDCFFDDFB27FA99FADA3B38" box="[1185,1294,1339,1360]" italics="true" pageId="10" pageNumber="151">D. signatus</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFA98FA413B27" author="Moser, J." box="[1306,1429,1338,1359]" pageId="10" pageNumber="151" pagination="353 - 367" refId="ref6425" refString="Moser, J. (1908) Neue Melolonthiden und Cetoniden. Annales de la Societe Entomologique de Belgique, 52 (12), 353 - 367. https: // doi. org / 10.5962 / bhl. part. 5014" type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
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<paragraph id="1F79BE1FFFDCFFDDFF11FAFAFA483BE2" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FAFAFA483BE2" pageId="10" pageNumber="151">
- Parameres rather curved and short, right paramere narrower than left one (see 
<figureCitation id="87FDA29AFFDCFFDDFC56FAF5FBDE3B04" box="[976,1034,1367,1389]" captionStart="FIGURE 2" captionStartId="3.[151,250,1444,1468]" captionTargetBox="[370,1260,238,1348]" captionTargetId="figure-20@3.[328,1260,181,1420]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in dorsal view. A–B, D. lexuanhuei Pham, Ahrens and Sabatinelli, new species. A, ♂ holotype; B, ♀ paratype; C–D, D. malyszi Bunalski, 2001 (Vietnam: Sa Pa; C, ♂; D, ♀). Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10611471" httpUri="https://zenodo.org/record/10611471/files/figure.png" pageId="10" pageNumber="151">Fig. 2</figureCitation>
, 
<bibRefCitation id="7B57C3EEFFDCFFDDFB92FAF5FAFE3B04" author="Sabatinelli, G. &amp; Pontuale, G." box="[1044,1322,1367,1389]" pageId="10" pageNumber="151" pagination="60 - 76" refId="ref6468" refString="Sabatinelli, G. &amp; Pontuale, G. (1998) Description of the genus Dedalopterus and notes on genus Malaisius Arrow and Cyphochilus Waterhouse (Col. Scarabaeoidea, Melolonthinae). Lambillionea, 97 (1), 60 - 76." type="journal article" year="1998">Sabatinelli &amp; Pontuale, 1998</bibRefCitation>
).................................................................................... 
<taxonomicName id="D8C6C59CFFDCFFDDFBF8FAD7FA483BE2" ID-CoL="3XLS3" authority="(Zhang, 1990)" baseAuthorityName="Zhang" baseAuthorityYear="1990" box="[1150,1436,1397,1419]" class="Insecta" family="Scarabaeidae" genus="Malaisius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="2DB2620DFFDCFFDDFBF8FAD7FAD93BE2" box="[1150,1293,1397,1418]" italics="true" pageId="10" pageNumber="151">D. intermedius</emphasis>
(
<bibRefCitation id="7B57C3EEFFDCFFDDFA9CFAD7FA413BE2" author="Zhang, Y. W." box="[1306,1429,1397,1419]" pageId="10" pageNumber="151" pagination="188 - 195" refId="ref6656" refString="Zhang, Y. W. (1990) Notes on the genus Malaisius Arrow with four new species from China (Col. Melolonthinae). Acta Zootaxonomica Sinica, 15 (2), 188 - 195." type="journal article" year="1990">Zhang, 1990</bibRefCitation>
)
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<keyLead id="A4371D2AFFDCFFDDFF11FA30FA483BCF" box="[151,1436,1426,1448]" pageId="10" pageNumber="151">8 Frontal depression deep; apical segment of maxillary palp with a large callus on its outer surface...................... 9</keyLead>
</paragraph>
<paragraph id="1F79BE1FFFDCFFDDFF11FA12FA483B8A" blockId="10.[151,1437,927,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11FA12FA483B8A" pageId="10" pageNumber="151">
- Frontal depression superficial, apical segment of maxillary palp with a minor callus on its outer surface................................................................................................. 
<taxonomicName id="D8C6C59CFFDCFFDDFB26FA6FFA483B8A" ID-CoL="6CGVD" authority="Bunalski, 2001" authorityName="Bunalski" authorityYear="2001" box="[1184,1436,1485,1507]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="malyszi">
<emphasis id="2DB2620DFFDCFFDDFB26FA6FFAD13B8A" box="[1184,1285,1485,1506]" italics="true" pageId="10" pageNumber="151">D. malyszi</emphasis>
<bibRefCitation id="7B57C3EEFFDCFFDDFA8DFA6FFA483B8A" author="Bunalski, M." box="[1291,1436,1485,1507]" pageId="10" pageNumber="151" pagination="247 - 251" refId="ref6144" refString="Bunalski, M. (2001) A new species of the genus Dedalopterus (Col. Melolonthidae) from Vietnam. Polish Journal of Entomology, 70, 247 - 251." type="journal article" year="2001">Bunalski, 2001</bibRefCitation>
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<keyLead id="A4371D2AFFDCFFDDFF11FA48FA493875" pageId="10" pageNumber="151">
9 Parameres unequal in length, right paramere shorter and slender than the left one............................................................................................ 
<taxonomicName id="D8C6C59CFFDCFFDDFC0EF9AAFAC93875" authority="Pham, Ahrens&amp; Sabatinelli" authorityName="Pham, Ahrens &amp; Sabatinelli" authorityYear="2024" box="[904,1309,1543,1565]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="lexuanhuei" status="new species">
<emphasis id="2DB2620DFFDCFFDDFC0EF9AAFBDA3874" box="[904,1038,1543,1565]" italics="true" pageId="10" pageNumber="151">D. lexuanhuei</emphasis>
Pham, Ahrens&amp; Sabatinelli
</taxonomicName>
, 
<emphasis id="2DB2620DFFDCFFDDFAA0F9A5FA493875" bold="true" box="[1318,1437,1543,1565]" pageId="10" pageNumber="151">
<taxonomicNameLabel id="3681DF76FFDCFFDDFAA0F9A5FA493875" box="[1318,1437,1543,1565]" pageId="10" pageNumber="151" rank="species">new species</taxonomicNameLabel>
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<paragraph id="1F79BE1FFFDCFFDDFF11F987FA483852" blockId="10.[151,1437,927,1595]" box="[151,1436,1573,1595]" pageId="10" pageNumber="151">
<keyLead id="A4371D2AFFDCFFDDFF11F987FA483852" box="[151,1436,1573,1595]" pageId="10" pageNumber="151">
- Parameres equal in length............................................. 
<taxonomicName id="D8C6C59CFFDCFFDDFB87F987FA483852" ID-CoL="6CGVF" authority="Sabatinelli &amp; Pontuale, 1998" authorityName="Sabatinelli &amp; Pontuale" authorityYear="1998" box="[1025,1436,1573,1595]" class="Insecta" family="Scarabaeidae" genus="Dedalopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="2DB2620DFFDCFFDDFB87F987FB553852" box="[1025,1153,1573,1594]" italics="true" pageId="10" pageNumber="151">D. pulchellus</emphasis>
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