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<document ID-DOI="http://doi.org/10.5281/zenodo.5980869" ID-GBIF-Dataset="641cd26f-54db-4069-9513-6ad899ed6b4e" ID-GBIF-Taxon="140561241" ID-ISSN="1175-5326" ID-Zenodo-Dep="5980869" ID-ZooBank="2F102698-EE0A-4FC9-AFCF-178222993E9B" checkinTime="1517396089111" checkinUser="plazi" docAuthor="Hsu, Yu-Feng" docDate="2018" docId="9766E10DD758FF84C489C378638EF81F" docLanguage="en" docName="zootaxa.4377.1.10.pdf" docOrigin="Zootaxa 1" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2449cff7" docStyleName="zootaxa.2013.journal_article" docTitle="Chrysozephyrus laoshanensis Hsu 2018, sp. nov." docType="treatment" docVersion="5" lastPageId="5" lastPageNumber="148" masterDocId="6B5F9975D759FF81C41EC76C6722FFA5" masterDocTitle="A new species of Chrysozephyrus associated with Lithocarpus corneus (Lepidoptera: Lycaenidae: Theclini)" masterLastPageNumber="150" masterPageNumber="143" pageId="1" pageNumber="144" updateTime="1644126660039" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Chrysozephyrus associated with Lithocarpus corneus (Lepidoptera: Lycaenidae: Theclini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hsu, Yu-Feng</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-01-31</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>143</mods:start>
<mods:end>150</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">https://doi.org/10.11646/zootaxa.4377.1.10</mods:identifier>
<mods:identifier type="GBIF-Dataset">641cd26f-54db-4069-9513-6ad899ed6b4e</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">1163194</mods:identifier>
<mods:identifier type="ZooBank">2F102698-EE0A-4FC9-AFCF-178222993E9B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5980869" ID-GBIF-Taxon="140561241" ID-Zenodo-Dep="5980869" LSID="urn:lsid:plazi:treatment:9766E10DD758FF84C489C378638EF81F" httpUri="http://treatment.plazi.org/id/9766E10DD758FF84C489C378638EF81F" lastPageId="5" lastPageNumber="148" pageId="1" pageNumber="144">
<subSubSection pageId="1" pageNumber="144" type="nomenclature">
<paragraph blockId="1.[151,674,1044,1104]" box="[151,674,1044,1071]" pageId="1" pageNumber="144">
<heading bold="true" box="[151,674,1044,1071]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
<emphasis bold="true" box="[151,674,1044,1071]" pageId="1" pageNumber="144">
<taxonomicName authority="Hsu" authorityName="Hsu" authorityYear="2018" box="[151,568,1044,1071]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis bold="true" box="[151,508,1044,1070]" italics="true" pageId="1" pageNumber="144">Chrysozephyrus laoshanensis</emphasis>
Hsu
</taxonomicName>
,
<taxonomicNameLabel box="[581,674,1045,1071]" pageId="1" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[151,674,1044,1104]" box="[151,510,1079,1104]" pageId="1" pageNumber="144">
(
<figureCitation box="[159,266,1079,1104]" captionStart="FIGURES 18" captionStartId="3.[151,264,1941,1964]" captionTargetBox="[162,1410,195,1915]" captionTargetId="figure@3.[150,1436,183,1931]" captionTargetPageId="3" captionText="FIGURES 18. Adults of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 1, upperside of holotype male, 2, underside of holotype male, 3, upperside of paratype female, 4, underside of paratype female, 5, upperside of male C. watsoni, 7, underside of male C. watsoni, 7, upperside of female C. watsoni, 8. underside of female C.watsoni. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1163196/files/figure.png" pageId="1" pageNumber="144">Figs. 14</figureCitation>
,
<figureCitation box="[278,377,1080,1104]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="1" pageNumber="144">9, 11, 13</figureCitation>
, 15, 1722)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="144" type="materials_examined">
<paragraph blockId="1.[151,1437,1143,2029]" pageId="1" pageNumber="144">
<materialsCitation ID-GBIF-Occurrence="1806155422" collectingDate="2016-04-02" collectingDateMax="2017-04-27" collectingDateMin="2016-04-02" collectionCode="IOZ" country="China" county="Type" elevation="1300" location="Tianlin Xian" municipality="Baise Shi" pageId="1" pageNumber="144" specimenCode="HSU 16" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis bold="true" box="[151,465,1143,1168]" pageId="1" pageNumber="144">
<collectingCounty box="[151,210,1143,1168]" pageId="1" pageNumber="144">
<typeStatus box="[151,210,1143,1168]" pageId="1" pageNumber="144">Type</typeStatus>
</collectingCounty>
materials.
<typeStatus box="[349,460,1143,1168]" pageId="1" pageNumber="144">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[476,500,1144,1167]" pageId="1" pageNumber="144" type="male"></specimenCount>
:
<collectingCountry box="[515,608,1144,1168]" name="China" pageId="1" pageNumber="144">CHINA</collectingCountry>
:
<collectingRegion box="[621,718,1143,1168]" country="China" name="Guangxi" pageId="1" pageNumber="144">Guangxi</collectingRegion>
Prov.,
<collectingMunicipality box="[804,917,1143,1168]" pageId="1" pageNumber="144">Baise Shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:9766E10DD758FF84C489C378638EF81F:1A1006C0D758FF80C7BDC31B6315FB35" box="[931,1079,1143,1168]" country="China" county="Type" municipality="Baise Shi" name="Tianlin Xian" pageId="1" pageNumber="144" stateProvince="Guangxi">Tianlin Xian</location>
,
<location LSID="urn:lsid:plazi:treatment:9766E10DD758FF84C489C378638EF81F:1A1006C0D758FF80C058C31B622AFB35" box="[1094,1288,1143,1168]" country="China" county="Type" municipality="Baise Shi" name="Cenwanglaoshan" pageId="1" pageNumber="144" stateProvince="Guangxi">Cenwanglaoshan</location>
,
<elevation box="[1303,1393,1144,1168]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="144" unit="m" value="1300.0">
<quantity box="[1303,1393,1144,1168]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="144" unit="m" value="1300.0">1300 m</quantity>
</elevation>
,
<collectingDate pageId="1" pageNumber="144" value="2016-04-02" valueMax="2016-04-03" valueMin="2016-04-02">
<date pageId="1" pageNumber="144" value="2016-04-02" valueMax="2016-04-03" valueMin="2016-04-02">2 3.IV.2016</date>
</collectingDate>
, reared from
<taxonomicName box="[413,645,1177,1201]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="species" species="corneus">
<emphasis box="[413,645,1177,1201]" italics="true" pageId="1" pageNumber="144">Lithocarpus corneus</emphasis>
</taxonomicName>
(
<specimenCode box="[659,752,1177,1202]" collectionCode="HSU" country="0" httpUri="http://grbio.org/cool/h9vj-e322" name="Humboldt State University" pageId="1" pageNumber="144">HSU 16</specimenCode>
D8), emgd.
<collectingDate box="[885,1016,1178,1202]" pageId="1" pageNumber="144" value="2017-04-27">
<date box="[885,1016,1178,1202]" pageId="1" pageNumber="144" value="2017-04-27">27. IV.2017</date>
</collectingDate>
(
<collectionCode box="[1031,1083,1178,1202]" pageId="1" pageNumber="144">IOZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[151,1437,1143,2029]" pageId="1" pageNumber="144">
<materialsCitation ID-GBIF-Occurrence="1806155424" collectionCode="HSU" pageId="1" pageNumber="144" specimenCode="HSU 16" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<typeStatus box="[199,319,1210,1235]" pageId="1" pageNumber="144">
<emphasis bold="true" box="[199,319,1210,1235]" pageId="1" pageNumber="144">Paratypes</emphasis>
</typeStatus>
.
<specimenCount box="[333,398,1210,1235]" pageId="1" pageNumber="144" type="male">5♂♂</specimenCount>
<specimenCount box="[398,447,1210,1235]" pageId="1" pageNumber="144" type="female">3♀♀</specimenCount>
, same locality and date as for
<typeStatus box="[792,893,1210,1235]" pageId="1" pageNumber="144">holotype</typeStatus>
, reared from
<taxonomicName box="[1044,1277,1211,1235]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="species" species="corneus">
<emphasis box="[1044,1277,1211,1235]" italics="true" pageId="1" pageNumber="144">Lithocarpus corneus</emphasis>
</taxonomicName>
(
<specimenCode box="[1291,1386,1210,1235]" collectionCode="HSU" country="0" httpUri="http://grbio.org/cool/h9vj-e322" name="Humboldt State University" pageId="1" pageNumber="144">HSU 16</specimenCode>
D8), emgd
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="1806155425" collectingDate="2016-04-23" collectingDateMax="2016-04-27" collectingDateMin="2016-04-23" collectionCode="IOZ, NTNU" country="China" county="Type" elevation="1300" location="Tianlin Xian" municipality="Baise Shi" pageId="1" pageNumber="144" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Guangxi" typeStatus="paratype">
<collectingDate box="[230,396,1246,1271]" pageId="1" pageNumber="144" value="2016-04-23" valueMax="2016-04-27" valueMin="2016-04-23">
<date box="[230,396,1246,1271]" pageId="1" pageNumber="144" value="2016-04-23" valueMax="2016-04-27" valueMin="2016-04-23">2327.IV.2016</date>
</collectingDate>
(3 dissected, genitalia YFH 1122, 1158, 1159) (
<collectionCode box="[954,1004,1247,1271]" pageId="1" pageNumber="144">IOZ</collectionCode>
,
<collectionCode box="[1019,1102,1247,1271]" country="China" httpUri="http://grbio.org/cool/6zr8-87q6" name="National Taiwan Normal University" pageId="1" pageNumber="144">NTNU</collectionCode>
);
<specimenCount box="[1123,1188,1247,1271]" pageId="1" pageNumber="144" type="male">2♂♂</specimenCount>
<specimenCount box="[1187,1238,1247,1271]" pageId="1" pageNumber="144" type="female">4♀♀</specimenCount>
, same locality as holotype
</materialsCitation>
,
<date box="[268,421,1282,1307]" pageId="1" pageNumber="144" value="2017-04-06" valueMax="2017-04-07" valueMin="2017-04-06">67. IV.2017</date>
, reared from
<taxonomicName box="[586,822,1283,1307]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="species" species="corneus">
<emphasis box="[586,822,1283,1307]" italics="true" pageId="1" pageNumber="144">Lithocarpus corneus</emphasis>
</taxonomicName>
(HSU 17D2), emgd.
<materialsCitation ID-GBIF-Occurrence="1806155423" collectingDate="2017-04-30" collectingDateMax="2017-05-01" collectingDateMin="2017-04-30" collectionCode="NTNU" country="Vietnam" elevation="1250" location="Ha Giang" pageId="1" pageNumber="144" specimenCount="1" specimenCount-female="1" stateProvince="Ha Giang" typeStatus="paratype">
<collectingDate box="[1083,1266,1283,1307]" pageId="1" pageNumber="144" value="2017-04-30" valueMax="2017-05-01" valueMin="2017-04-30">
<date box="[1083,1266,1283,1307]" pageId="1" pageNumber="144" value="2017-04-30" valueMax="2017-05-01" valueMin="2017-04-30">30.IV.1.V.2017</date>
</collectingDate>
(
<collectionCode box="[1286,1370,1283,1307]" country="China" httpUri="http://grbio.org/cool/6zr8-87q6" name="National Taiwan Normal University" pageId="1" pageNumber="144">NTNU</collectionCode>
).
<specimenCount box="[1393,1430,1283,1307]" pageId="1" pageNumber="144" type="female">1♀</specimenCount>
:
<collectingCountry box="[151,285,1319,1343]" name="Vietnam" pageId="1" pageNumber="144">VIETNAM</collectingCountry>
:
<collectingRegion box="[295,402,1318,1343]" country="Vietnam" name="Ha Giang" pageId="1" pageNumber="144">Ha Giang</collectingRegion>
Prov., Meo Vac,
<elevation box="[596,683,1318,1343]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="144" unit="m" value="1250.0">
<quantity box="[596,683,1318,1343]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="144" unit="m" value="1250.0">1250 m</quantity>
</elevation>
</materialsCitation>
,
<date box="[695,814,1319,1343]" pageId="1" pageNumber="144" value="2014-05-22">22.V.2014</date>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="145" pageId="1" pageNumber="144" type="description">
<paragraph blockId="1.[151,1437,1143,2029]" lastBlockId="2.[151,1437,151,1974]" lastPageId="2" lastPageNumber="145" pageId="1" pageNumber="144">
<emphasis bold="true" box="[199,337,1354,1379]" pageId="1" pageNumber="144">Description</emphasis>
. Male (
<figureCitation box="[424,534,1354,1379]" captionStart="FIGURES 18" captionStartId="3.[151,264,1941,1964]" captionTargetBox="[162,1410,195,1915]" captionTargetId="figure@3.[150,1436,183,1931]" captionTargetPageId="3" captionText="FIGURES 18. Adults of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 1, upperside of holotype male, 2, underside of holotype male, 3, upperside of paratype female, 4, underside of paratype female, 5, upperside of male C. watsoni, 7, underside of male C. watsoni, 7, upperside of female C. watsoni, 8. underside of female C.watsoni. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1163196/files/figure.png" pageId="1" pageNumber="144">Figs. 12</figureCitation>
): FL
<quantity box="[594,757,1354,1379]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" metricValueMax="1.9600000000000002" metricValueMin="1.6199999999999999" pageId="1" pageNumber="144" unit="mm" value="17.9" valueMax="19.6" valueMin="16.2">16.219.6 mm</quantity>
(mean, 18.2 ±
<quantity box="[926,1014,1355,1379]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="144" unit="mm" value="1.4">1.4 mm</quantity>
, n = 8); AL
<quantity box="[1153,1288,1355,1379]" metricMagnitude="-3" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.7" metricValueMin="7.2" pageId="1" pageNumber="144" unit="mm" value="7.949999999999999" valueMax="8.7" valueMin="7.2">7.28.7 mm</quantity>
(mean, 8.1 ±
<quantity box="[151,239,1390,1415]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="144" unit="mm" value="0.5">0.5 mm</quantity>
, n = 8). Head: Hairy, vertex, frons dark brown but sparsely bearing white hairs mesad; a narrow white rim surrounding eye; eye semi-oval, densely covered with long, buff setae; labial palpus porrect, with 3rd segment pointed downwards, covered with dark brown scaling mottled with white laterally; maxillary palpus reduced, inconspicuous; proboscis unscaled; antenna smooth scaled, naked at terminal end of nudum. Thorax: Dark brown dorsad, white ventrad; legs with tarsus of fore leg stubby; white, mottled with brown. Forewing: Termen, costa slightly concave, dorsum straight. Ground color of upperside dark brown, overlaid with large area of metallic green tinged with yellow scaling proximally, covering most of forewing, leaving merely a narrow dark brown margin. Fringe white, prominent. Ground color of underside buff gray. Discal spot forming brown bar edged with white. Distal band of central symmetry system represented as tilted white line edged with brown proximally, strongly dislocated inwards in cell Cu2. Submarginal band and &quot;g&quot;-element (per
<bibRefCitation author="Nijhout" box="[966,1117,1709,1734]" pageId="1" pageNumber="144" refString="Nijhout, H. F. (1991) The development and evolution of butterfly wing patterns. Smithsonian Institution Press, Washington and London, xvi + 297 pp." type="book" year="1991">Nijhout 1991</bibRefCitation>
) fused into prominent, dark brown band edged with white, attenuate towards apex. Fringe white, prominent. Hindwing: Contour of wing slightly produced at distal end of M1 and Cu1; Cu2 bearing long, tail-like projection. Metallic scaling on upperside less extensive than that of forewing, with dark brown areas including a patch along costa, narrow bands along termen and dorsum. Narrow, metallic blue line at distal ends of cells along termen. Ground color of underside buff. Discal spot forming brown bar edged with white. Distal band of central symmetry system forming prominent white line edged with dark brown proximally, forming straight line from dorsum to vein Cu1, running outwards in cell Cu1, dislocated proximally as tick-shaped or a straight line in cell Cu2, retracing into straight line in cell 2A, and becoming a slightly curved line in cell 1A. Submarginal band consisting of faint, broad, white band, a black, oval spot enclosed within orange circle in cell Cu1, and a tornal orange patch edged by black and metallic blue scaling anteriorly and posteriorly; &quot;g&quot;-element consisting of a string of arch-like, faint white markings. Fringe white, prominent. Abdomen: Dark brown dorsad, white ventrad.
<emphasis bold="true" box="[829,939,225,250]" pageId="2" pageNumber="145">Genitalia</emphasis>
(
<figureCitation box="[958,1133,225,250]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Figs. 9, 11, 13</figureCitation>
): Ring-shaped sclerites of 9+10 segments with posterior end nearly straight, a transverse notch present anteriorly; saccus flap-like, shallowly depressed; brachium abruptly bent over, forming elongate arm with distal end up-curved; elbow bearing a prominent, simple spine with pointed end; valva pincer-like, bearing two distal, flatten processes, with dorsal one (ampulla) longer, stout basally, twisted half-way, with distal portion thin, dagger-like, pointing meso-posteriad; ventral one (harpe) with distal end pointed, shorter, flattened, slightly up-curved. Phallus elongate, nearly straight with truncated caudal end; aedeagus approximately 1.77x phallobase in length, cornuti present, forming a cluster of small spinules at caudal end. Juxta broad, pincer-like with deep dorsal notch.
</paragraph>
<paragraph blockId="2.[151,1437,151,1974]" pageId="2" pageNumber="145">
<emphasis bold="true" box="[199,286,509,534]" pageId="2" pageNumber="145">Female</emphasis>
(
<figureCitation box="[301,411,509,534]" captionStart="FIGURES 18" captionStartId="3.[151,264,1941,1964]" captionTargetBox="[162,1410,195,1915]" captionTargetId="figure@3.[150,1436,183,1931]" captionTargetPageId="3" captionText="FIGURES 18. Adults of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 1, upperside of holotype male, 2, underside of holotype male, 3, upperside of paratype female, 4, underside of paratype female, 5, upperside of male C. watsoni, 7, underside of male C. watsoni, 7, upperside of female C. watsoni, 8. underside of female C.watsoni. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1163196/files/figure.png" pageId="2" pageNumber="145">Figs. 34</figureCitation>
). FL 16.0
<quantity box="[535,634,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="2" pageNumber="145" unit="mm" value="19.3">19.3 mm</quantity>
(mean, 18.2 ±
<quantity box="[804,892,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="145" unit="mm" value="1.1">1.1 mm</quantity>
, n = 8); AL
<quantity box="[1031,1166,509,534]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="8.7" metricValueMin="7.5" pageId="2" pageNumber="145" unit="mm" value="8.1" valueMax="8.7" valueMin="7.5">7.58.7 mm</quantity>
(mean, 8.3 ±
<quantity box="[1322,1410,510,534]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="145" unit="mm" value="0.4">0.4 mm</quantity>
, n = 8). Body, wing patterns of underside as described for male except scaling lacking gray tone. Fore leg with tarsus segmented. Wing upperside uniformly brown with prominent, proximal, metallic, pale blue patch on forewing. Metallic blue line at distal ends of cells along termen more prominent than that of male.
<emphasis bold="true" box="[1204,1314,617,642]" pageId="2" pageNumber="145">Genitalia</emphasis>
(Fig. 15): Apophyses posteriores elongate, slender, curved, abruptly enlarged, flattened basad. Sterigma with lamella antevaginallis heavily sclerotized, rench-like with prominent, trident, medial cleft; lamella postvaginalis forming moderately sclerotized wall with transverse wrinkles. Posterior half of ductus bursae tubular, weakly sclerotized; anterior half enlarged, pouch-like, membranous. Corpus bursae rounded, bearing a pair of small, discal, signa with invaginated blade. Papillae anales lobe-like, setose.
</paragraph>
<paragraph blockId="2.[151,1437,151,1974]" pageId="2" pageNumber="145">
<emphasis bold="true" box="[199,331,833,858]" pageId="2" pageNumber="145">Immatures</emphasis>
. Ova (
<figureCitation box="[413,498,833,858]" captionStart="FIGURES1722" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[150,1436,193,1588]" captionTargetId="figure@5.[150,1436,193,1588]" captionTargetPageId="5" captionText="FIGURES1722.Hostplant and immatures of Chrysozephyrus laoshanensis, sp. nov.: 17, host plant Lithocarpus corneus var. angustifolius at type locality, 18, ovum, 19, 3rd instar larva resting on underside of young leaf, 20, mature (4th instar larva) larva, 21, dorsal view of pupa, 22, lateral view of pupa." httpUri="https://zenodo.org/record/1163200/files/figure.png" pageId="2" pageNumber="145">Fig. 18</figureCitation>
) approximately 0.73 ±
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in diameter, 0.44 ±
<quantity box="[1120,1223,834,858]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="145" unit="mm" value="0.01">0.01 mm</quantity>
in height (n = 8), spherical but compressed, white, surface with fine sculpture forming framework with short stub-like processes. Larvae (
<figureCitation box="[245,380,905,930]" captionStart="FIGURES1722" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[150,1436,193,1588]" captionTargetId="figure@5.[150,1436,193,1588]" captionTargetPageId="5" captionText="FIGURES1722.Hostplant and immatures of Chrysozephyrus laoshanensis, sp. nov.: 17, host plant Lithocarpus corneus var. angustifolius at type locality, 18, ovum, 19, 3rd instar larva resting on underside of young leaf, 20, mature (4th instar larva) larva, 21, dorsal view of pupa, 22, lateral view of pupa." httpUri="https://zenodo.org/record/1163200/files/figure.png" pageId="2" pageNumber="145">Figs. 1920</figureCitation>
) with four instars. Head dark brown, glossy. Body slug-like, of typically lycaenid form, with A8 produced laterally in final instar; surface bearing transparent setae. Coloration of body olive-green in younger instars, becoming dark green to brown when full-grown. White chevrons edged by brown subdorsally. A dark, medial, longitudinal line dorsally, abruptly broaden cephalically. T1 shield diamond-shaped. Anal lobe somewhat square. Spiracles dark brown. Full-grown larva 17.0 mm in length (n = 5). Pupae (
<figureCitation box="[1071,1198,1049,1074]" captionStart="FIGURES1722" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[150,1436,193,1588]" captionTargetId="figure@5.[150,1436,193,1588]" captionTargetPageId="5" captionText="FIGURES1722.Hostplant and immatures of Chrysozephyrus laoshanensis, sp. nov.: 17, host plant Lithocarpus corneus var. angustifolius at type locality, 18, ovum, 19, 3rd instar larva resting on underside of young leaf, 20, mature (4th instar larva) larva, 21, dorsal view of pupa, 22, lateral view of pupa." httpUri="https://zenodo.org/record/1163200/files/figure.png" pageId="2" pageNumber="145">Fig. 2122</figureCitation>
): 11.55 ±
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in length, 4.48 ±
<quantity box="[313,412,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="145" unit="mm" value="0.2">0.20 mm</quantity>
in width of T2, 5.52 ±
<quantity box="[671,771,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="2" pageNumber="145" unit="mm" value="0.18">0.18 mm</quantity>
in width of A3, 4.90 ±
<quantity box="[1033,1133,1086,1110]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="145" unit="mm" value="0.23">0.23 mm</quantity>
in height of A3 (n = 5); of typical lycaenid form, brown tinged with yellow, mottled with dark brown dorsally, a dark brown medial line present dorsad on abdomen; wings with extensive dark brown markings; spiracles cream white.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="145" type="diagnosis">
<paragraph blockId="2.[151,1437,151,1974]" pageId="2" pageNumber="145">
<emphasis bold="true" box="[199,314,1193,1218]" pageId="2" pageNumber="145">Diagnosis</emphasis>
. The general morphology of the genitalia (
<figureCitation box="[802,961,1193,1218]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Figs 9, 11, 13</figureCitation>
, 15) in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[1050,1229,1193,1217]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[1050,1229,1193,1217]" italics="true" pageId="2" pageNumber="145">C. laoshanensis</emphasis>
</taxonomicName>
conforms with all congeners, including a straight phallus, cornuti as minute spinules, ductus bursae tubular, sclerotized caudally and pouch-like, membranous cephalically (
<bibRefCitation author="Shirozu" box="[595,924,1265,1290]" pageId="2" pageNumber="145" refString="Shirozu T. &amp; Yamamoto, H. (1956) A generic revision and the phylogeny of the tribe Theclini (Lepidoptera; Lycaenidae). Sieboldia, 1, 329 - 421." type="journal article" year="1956">Shirozu &amp; Yamamoto, 1956</bibRefCitation>
). However,
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[1064,1245,1265,1289]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[1064,1245,1265,1289]" italics="true" pageId="2" pageNumber="145">C. laoshanensis</emphasis>
</taxonomicName>
possesses a few peculiar features not seen in other
<taxonomicName authorityName="Shirozu and Yamamoto" authorityYear="1956" box="[549,730,1301,1325]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis box="[549,730,1301,1325]" italics="true" pageId="2" pageNumber="145">Chrysozephyrus</emphasis>
</taxonomicName>
species, including pincer-shaped valva and elbow position of the brachium forming a simple spine. The valvae of all other
<taxonomicName authorityName="Shirozu and Yamamoto" authorityYear="1956" box="[847,1028,1337,1361]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis box="[847,1028,1337,1361]" italics="true" pageId="2" pageNumber="145">Chrysozephyrus</emphasis>
</taxonomicName>
species have the harpe only weakly represented, thus not pincer-like in shape; elbow position of brachium is serrate, bearing small spines in other
<taxonomicName authorityName="Shirozu and Yamamoto" authorityYear="1956" box="[151,332,1409,1433]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis box="[151,332,1409,1433]" italics="true" pageId="2" pageNumber="145">Chrysozephyrus</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph blockId="2.[151,1437,151,1974]" pageId="2" pageNumber="145">
In terms of wing patterns,
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[492,669,1446,1470]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[492,669,1446,1470]" italics="true" pageId="2" pageNumber="145">C. laoshanensis</emphasis>
</taxonomicName>
is most similar to the sympatric,
<taxonomicName box="[1038,1115,1446,1469]" class="Magnoliopsida" family="Ericaceae" genus="Lyonia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="145" phylum="Tracheophyta" rank="genus">
<emphasis box="[1038,1115,1446,1469]" italics="true" pageId="2" pageNumber="145">Lyonia</emphasis>
</taxonomicName>
-associated
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1245,1364,1446,1469]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1245,1364,1446,1469]" italics="true" pageId="2" pageNumber="145">C. watsoni</emphasis>
</taxonomicName>
(
<figureCitation captionStart="FIGURES 18" captionStartId="3.[151,264,1941,1964]" captionTargetBox="[162,1410,195,1915]" captionTargetId="figure@3.[150,1436,183,1931]" captionTargetPageId="3" captionText="FIGURES 18. Adults of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 1, upperside of holotype male, 2, underside of holotype male, 3, upperside of paratype female, 4, underside of paratype female, 5, upperside of male C. watsoni, 7, underside of male C. watsoni, 7, upperside of female C. watsoni, 8. underside of female C.watsoni. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1163196/files/figure.png" pageId="2" pageNumber="145">Figs. 58</figureCitation>
). They may be separated using the following diagnostic characteristics: The distal band of central symmetry system on the hindwing underside forms a tick-shaped or straight white line in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[1063,1208,1518,1542]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[1063,1208,1518,1542]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
, whereas it forms a broken line in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[319,406,1554,1577]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[319,406,1554,1577]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
. The distal band of central symmetry system on the forewing of the male is prominent in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[151,296,1590,1614]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[151,296,1590,1614]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
, whereas it is obsolete or barely recognizable in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[895,982,1590,1613]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[895,982,1590,1613]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
. Dark brown margin on the forewing upperside of the male is narrow, nearly equal in width throughout the length in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[1044,1189,1625,1649]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[1044,1189,1625,1649]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
, whereas it is thicker, broaden anteriorly, forming a prominent dark area at the apex in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[874,961,1662,1685]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[874,961,1662,1685]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
. In
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[1005,1150,1662,1686]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[1005,1150,1662,1686]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
, the metallic blue scaling on the wing upperside of the female is only found on forewing and pale blue in color; no orange spot is present on the forewing. By contrast, in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[487,574,1734,1757]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[487,574,1734,1757]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
, it is blue or purple, sometimes also found on the hindwing, and additional orange spots sometimes are found on the forewing. The brachium possesses a prominent spine at elbow in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[151,296,1806,1830]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[151,296,1806,1830]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
(
<figureCitation box="[312,381,1805,1830]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 9</figureCitation>
), but a conical process with serrate end in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[868,955,1806,1829]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[868,955,1806,1829]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
(
<figureCitation box="[971,1055,1805,1830]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 10</figureCitation>
). The aedeagus is longer than the phallobase in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[308,453,1841,1865]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[308,453,1841,1865]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
(
<figureCitation box="[469,552,1841,1866]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 11</figureCitation>
), but nearly equal in length in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[901,988,1842,1865]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[901,988,1842,1865]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
(
<figureCitation box="[1005,1089,1841,1866]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 12</figureCitation>
). The valva is pincer-like with two elongate processes with acute distal end in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[693,838,1878,1902]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[693,838,1878,1902]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
(
<figureCitation box="[854,938,1877,1902]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 13</figureCitation>
), but lobe-like with only a digitate, inwardpointing process in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[378,465,1914,1937]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[378,465,1914,1937]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
(
<figureCitation box="[483,569,1913,1938]" captionStart="FIGURES 914" captionStartId="4.[151,264,1880,1903]" captionTargetBox="[262,1320,210,1830]" captionTargetId="figure@4.[245,1356,193,1859]" captionTargetPageId="4" captionText="FIGURES 914. Male genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 9. caudal view of tegumen of C. laoshanensis, sp. nov. with brachia attached, 10, caudal view of tegumen of C. watsoni with brachia attached, 11, lateral view of phallus of C. laoshanensis, sp. nov., 12, lateral view of phallus of C. watsoni, 13. Caudal view of valvae of C. laoshanensis, sp. nov., 14, Caudal view of valvae of C. watsoni. FIGURES 1516. Female genitalia of Chrysozephyrus laoshanensis, sp. nov. and sympatric C. watsoni: 15, Chrysozephyrus laoshanensis, sp. nov., 16, C. watsoni." httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="2" pageNumber="145">Fig. 14</figureCitation>
). The lamella antevaginallis broad, rench-like with a trident medial cleft in
<taxonomicName authorityName="Hsu" authorityYear="2018" box="[151,296,1950,1974]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="laoshanensis">
<emphasis box="[151,296,1950,1974]" italics="true" pageId="2" pageNumber="145">laoshanensis</emphasis>
</taxonomicName>
(Fig. 15), whereas it is narrow, bump-like with shallow medial cleft in
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1094,1181,1950,1973]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1094,1181,1950,1973]" italics="true" pageId="2" pageNumber="145">watsoni</emphasis>
</taxonomicName>
(Fig. 16).
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/1163196/files/figure.png" pageId="3" pageNumber="146" targetBox="[162,1410,195,1915]" targetPageId="3">
<paragraph blockId="3.[151,1436,1941,2025]" pageId="3" pageNumber="146">
<emphasis bold="true" box="[151,315,1941,1964]" pageId="3" pageNumber="146">FIGURES 18.</emphasis>
Adults of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[424,725,1942,1964]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[424,725,1942,1964]" italics="true" pageId="3" pageNumber="146">Chrysozephyrus laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[738,819,1942,1963]" pageId="3" pageNumber="146" rank="species">
<emphasis bold="true" box="[738,819,1942,1963]" pageId="3" pageNumber="146">sp. nov.</emphasis>
</taxonomicNameLabel>
and sympatric
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[977,1086,1943,1964]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[977,1086,1943,1964]" italics="true" pageId="3" pageNumber="146">C. watsoni</emphasis>
</taxonomicName>
: 1, upperside of holotype male, 2, underside of holotype male, 3, upperside of paratype female, 4, underside of paratype female, 5, upperside of male
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1321,1430,1973,1995]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1321,1430,1973,1995]" italics="true" pageId="3" pageNumber="146">C. watsoni</emphasis>
</taxonomicName>
, 7, underside of male
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[364,472,2004,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[364,472,2004,2025]" italics="true" pageId="3" pageNumber="146">C. watsoni</emphasis>
</taxonomicName>
, 7, upperside of female
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[717,826,2004,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[717,826,2004,2025]" italics="true" pageId="3" pageNumber="146">C. watsoni</emphasis>
</taxonomicName>
, 8. underside of female
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1071,1179,2004,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1071,1179,2004,2025]" italics="true" pageId="3" pageNumber="146">C. watsoni</emphasis>
</taxonomicName>
. Scale bar = 1 cm.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/1163198/files/figure.png" pageId="4" pageNumber="147" targetBox="[262,1320,210,1830]" targetPageId="4">
<paragraph blockId="4.[151,1436,1880,2025]" pageId="4" pageNumber="147">
<emphasis bold="true" box="[151,332,1880,1903]" pageId="4" pageNumber="147">FIGURES 914.</emphasis>
Male genitalia of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[537,842,1881,1903]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[537,842,1881,1903]" italics="true" pageId="4" pageNumber="147">Chrysozephyrus laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[859,944,1881,1902]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[859,944,1881,1902]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
and sympatric
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1115,1229,1882,1903]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1115,1229,1882,1903]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
: 9. caudal view of tegumen of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[274,436,1911,1933]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[274,436,1911,1933]" italics="true" pageId="4" pageNumber="147">C. laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[450,531,1912,1933]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[450,531,1912,1933]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
with brachia attached, 10, caudal view of tegumen of
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1093,1203,1912,1934]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1093,1203,1912,1934]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
with brachia attached, 11, lateral view of phallus of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[450,611,1942,1964]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[450,611,1942,1964]" italics="true" pageId="4" pageNumber="147">C. laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[624,705,1943,1964]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[624,705,1943,1964]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
, 12, lateral view of phallus of
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1019,1128,1943,1964]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1019,1128,1943,1964]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
, 13. Caudal view of valvae of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[151,314,1973,1995]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[151,314,1973,1995]" italics="true" pageId="4" pageNumber="147">C. laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[328,411,1973,1994]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[328,411,1973,1994]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
, 14, Caudal view of valvae of
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[739,850,1974,1995]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[739,850,1974,1995]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
.
<emphasis bold="true" box="[865,1056,1972,1995]" pageId="4" pageNumber="147">FIGURES 1516.</emphasis>
Female genitalia of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis italics="true" pageId="4" pageNumber="147">Chrysozephyrus laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[294,375,2004,2025]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[294,375,2004,2025]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
and sympatric
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[531,639,2004,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[531,639,2004,2025]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
: 15,
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[690,990,2003,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[690,990,2003,2025]" italics="true" pageId="4" pageNumber="147">Chrysozephyrus laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1003,1083,2004,2025]" pageId="4" pageNumber="147" rank="species">
<emphasis bold="true" box="[1003,1083,2004,2025]" pageId="4" pageNumber="147">sp. nov.</emphasis>
</taxonomicNameLabel>
, 16,
<taxonomicName baseAuthorityName="Evans" baseAuthorityYear="1927" box="[1134,1242,2004,2025]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="watsoni">
<emphasis box="[1134,1242,2004,2025]" italics="true" pageId="4" pageNumber="147">C. watsoni</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/1163200/files/figure.png" pageId="5" pageNumber="148" targetBox="[150,1436,193,1588]" targetPageId="5">
<paragraph blockId="5.[151,1437,1609,1693]" pageId="5" pageNumber="148">
<emphasis bold="true" box="[151,340,1609,1632]" pageId="5" pageNumber="148">FIGURES 1722.</emphasis>
Hostplant and immatures of
<taxonomicName authority="Hsu, 2018" authorityName="Hsu" authorityYear="2018" box="[633,934,1609,1631]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="laoshanensis" status="sp. nov.">
<emphasis box="[633,934,1609,1631]" italics="true" pageId="5" pageNumber="148">Chrysozephyrus laoshanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[946,1026,1610,1631]" pageId="5" pageNumber="148" rank="species">
<emphasis bold="true" box="[946,1026,1610,1631]" pageId="5" pageNumber="148">sp. nov.</emphasis>
</taxonomicNameLabel>
: 17, host plant
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="148" phylum="Tracheophyta" rank="variety" species="corneus" variety="angustifolius">
<emphasis box="[1184,1392,1609,1631]" italics="true" pageId="5" pageNumber="148">Lithocarpus corneus</emphasis>
var.
<emphasis box="[151,282,1640,1662]" italics="true" pageId="5" pageNumber="148">angustifolius</emphasis>
</taxonomicName>
at type locality, 18, ovum, 19, 3rd instar larva resting on underside of young leaf, 20, mature (4th instar larva) larva, 21, dorsal view of pupa, 22, lateral view of pupa.
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="148" type="biology_ecology">
<paragraph blockId="5.[151,1436,1737,1978]" box="[199,987,1737,1762]" pageId="5" pageNumber="148">
<emphasis bold="true" box="[199,324,1737,1762]" pageId="5" pageNumber="148">Host plant</emphasis>
.
<taxonomicName box="[338,769,1738,1762]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="148" phylum="Tracheophyta" rank="variety" species="corneus" variety="angustifolius">
<emphasis box="[338,569,1738,1762]" italics="true" pageId="5" pageNumber="148">Lithocarpus corneus</emphasis>
var.
<emphasis box="[624,769,1738,1762]" italics="true" pageId="5" pageNumber="148">angustifolius</emphasis>
</taxonomicName>
(
<figureCitation box="[784,865,1737,1762]" captionStart="FIGURES1722" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[150,1436,193,1588]" captionTargetId="figure@5.[150,1436,193,1588]" captionTargetPageId="5" captionText="FIGURES1722.Hostplant and immatures of Chrysozephyrus laoshanensis, sp. nov.: 17, host plant Lithocarpus corneus var. angustifolius at type locality, 18, ovum, 19, 3rd instar larva resting on underside of young leaf, 20, mature (4th instar larva) larva, 21, dorsal view of pupa, 22, lateral view of pupa." httpUri="https://zenodo.org/record/1163200/files/figure.png" pageId="5" pageNumber="148">Fig. 17</figureCitation>
) in nature.
</paragraph>
<paragraph blockId="5.[151,1436,1737,1978]" pageId="5" pageNumber="148">
<emphasis bold="true" box="[199,323,1773,1798]" pageId="5" pageNumber="148">Bionomics</emphasis>
. Univoltine, overwintering as an ovum, which is singly attached on the underside of twigs. Ovum hatch by late March in the
<typeStatus box="[454,502,1810,1834]" pageId="5" pageNumber="148">type</typeStatus>
locality. Larva fed on only soft foliage. Adults emerged by late April to early May under laboratory conditions. Eggs were present by early July, laid singly on small twigs of the host plant in shady positions approximately
<quantity box="[428,499,1882,1906]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="5" pageNumber="148" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">14 m</quantity>
above the ground.
</paragraph>
</subSubSection>
<subSubSection box="[199,1109,1917,1942]" pageId="5" pageNumber="148" type="distribution">
<paragraph blockId="5.[151,1436,1737,1978]" box="[199,1109,1917,1942]" pageId="5" pageNumber="148">
<emphasis bold="true" box="[199,352,1917,1942]" pageId="5" pageNumber="148">Distribution.</emphasis>
<collectingCountry box="[359,426,1917,1942]" name="China" pageId="5" pageNumber="148">China</collectingCountry>
(
<collectingRegion box="[441,540,1917,1942]" country="China" name="Guangxi" pageId="5" pageNumber="148">Guangxi</collectingRegion>
) and northern
<collectingCountry box="[704,799,1917,1942]" name="Vietnam" pageId="5" pageNumber="148">Vietnam</collectingCountry>
(
<collectingRegion box="[814,925,1917,1942]" country="Vietnam" name="Ha Giang" pageId="5" pageNumber="148">Ha Giang</collectingRegion>
) for the present.
</paragraph>
</subSubSection>
<subSubSection box="[199,1196,1953,1978]" pageId="5" pageNumber="148" type="etymology">
<paragraph blockId="5.[151,1436,1737,1978]" box="[199,1196,1953,1978]" pageId="5" pageNumber="148">
<emphasis bold="true" box="[199,326,1953,1978]" pageId="5" pageNumber="148">Etymology</emphasis>
. The specific name refers to the abbreviation of the name of the
<typeStatus box="[1051,1099,1954,1978]" pageId="5" pageNumber="148">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>