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<mods:title id="82C584A44BE031924A4393C4C04618CF">Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes</mods:title>
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<taxonomicName id="4C7E4D5CFFABFF9BFF1BCC84C8EDE33E" ID-CoL="76XJQ" authority="Bernhauer, 1909" authorityName="Bernhauer" authorityYear="1909" box="[151,645,499,526]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
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<emphasis id="B90AEACDFFABFF9BFF1BCC84CBD8E33D" bold="true" box="[151,432,500,526]" italics="true" pageId="7" pageNumber="38">Philonthus bonariensis</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFE3BCC83C8EDE33E" author="Bernhauer" box="[439,645,499,525]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1909) Zur Staphylinidenfauna von Sudamerika (6. Beitrag.). Bolletino della Societa Entomologica Italiana, 40 (1908), 225 - 251." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
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(
<figureCitation id="13452A5AFFABFF9BFF13CF66CB79E303" box="[159,273,534,560]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="7" pageNumber="38">Figs. 79</figureCitation>
;
<figureCitation id="13452A5AFFABFF9BFE91CF66CB04E303" box="[285,364,534,560]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="7" pageNumber="38">
Figs.
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15 in Chani-Posse 2004)
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<taxonomicName id="4C7E4D5CFFABFF9BFF1BCF14C8EAE348" ID-CoL="76XJQ" authority="Bernhauer, 1909: 245" authorityName="Bernhauer" authorityPageNumber="245" authorityYear="1909" box="[151,642,612,635]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFF1BCF14CBE6E348" box="[151,398,612,635]" italics="true" pageId="7" pageNumber="38">Philonthus bonariensis</emphasis>
<treatmentCitation id="0ADF10CEFFABFF9BFE1ACF14C8EAE348" author="Bernhauer" box="[406,642,612,635]" page="245" pageId="7" pageNumber="38" year="1909">
<bibRefCitation id="EFEF4B2EFFABFF9BFE1ACF14C824E348" author="Bernhauer" box="[406,588,612,635]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1909) Zur Staphylinidenfauna von Sudamerika (6. Beitrag.). Bolletino della Societa Entomologica Italiana, 40 (1908), 225 - 251." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
: 245
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;
<treatmentCitation id="0ADF10CEFFABFF9BFD03CF14C994E348" author="Bernhauer" box="[655,1020,612,635]" page="330" pageId="7" pageNumber="38" year="1914">
<bibRefCitation id="EFEF4B2EFFABFF9BFD03CF14C9A0E348" author="Bernhauer" box="[655,968,612,635]" pageId="7" pageNumber="41" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 5 (57). W. Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
: 330
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(catalog);
<treatmentCitation id="0ADF10CEFFABFF9BFBFECF14CF46E348" author="Bruch" box="[1138,1326,612,635]" page="444" pageId="7" pageNumber="38" year="1915">
<bibRefCitation id="EFEF4B2EFFABFF9BFBFECF14CE92E348" author="Bruch" box="[1138,1274,612,635]" pageId="7" pageNumber="41" refString="Bruch, C. (1915) Catalogo Sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235 - 302." type="journal article" year="1915">Bruch, 1915</bibRefCitation>
: 444
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(catalog);
<treatmentCitation id="0ADF10CEFFABFF9BFF4BCFF4CBAFE3A8" author="Blackwelder" box="[199,455,644,667]" page="132" pageId="7" pageNumber="38" year="1944">
<bibRefCitation id="EFEF4B2EFFABFF9BFF4BCFF4CBFCE3A8" author="Blackwelder" box="[199,404,644,667]" pageId="7" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 132
</treatmentCitation>
(list);
<treatmentCitation id="0ADF10CEFFABFF9BFD9DCFF4C95AE3A8" author="Coiffait" box="[529,818,644,667]" page="355" pageId="7" pageNumber="38" year="1968">
<bibRefCitation id="EFEF4B2EFFABFF9BFD9DCFF4C896E3A8" author="Coiffait" box="[529,766,644,667]" pageId="7" pageNumber="41" refString="Coiffait, H. &amp; Saiz, F. (1968) Les Staphylinidae (sensu lato) du Chili. In: Delamare Deboutteville, C. &amp; Rapoport, E. (Eds.), Biologie de l'Amerique Australe. Etudes sur la faune du sol, 4. Centre National de la Recherche Scientifique, Paris, pp. 339 - 468." type="journal article" year="1968">Coiffait &amp; Sáiz, 1968</bibRefCitation>
: 355
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(misidentified as
<taxonomicName id="4C7E4D5CFFABFF9BFC7ACFF4CEBFE3A8" ID-CoL="76ZGR" box="[1014,1239,644,667]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="hepaticus">
<emphasis id="B90AEACDFFABFF9BFC7ACFF4CEBFE3A8" box="[1014,1239,644,667]" italics="true" pageId="7" pageNumber="38">Philonthus hepaticus</emphasis>
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);
<treatmentCitation id="0ADF10CEFFABFF9BFB7CCFF4CFF4E3A8" author="Frank" box="[1264,1436,644,667]" page="38" pageId="7" pageNumber="38" year="1982">
<bibRefCitation id="EFEF4B2EFFABFF9BFB7CCFF4CF1EE3A8" author="Frank" box="[1264,1398,644,667]" pageId="7" pageNumber="42" refString="Frank, J. H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agr. Exp. Stations, Technical Bulletin, 824, i - vii + 1 - 118 pp." type="journal article" year="1982">Frank, 1982</bibRefCitation>
: 38
</treatmentCitation>
(list);
<treatmentCitation id="0ADF10CEFFABFF9BFE8BCFD4CB8BE388" author="Herman" box="[263,483,676,699]" page="2762" pageId="7" pageNumber="38" year="2001">
<bibRefCitation id="EFEF4B2EFFABFF9BFE8BCFD4CBCBE388" author="Herman" box="[263,419,676,699]" pageId="7" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2762
</treatmentCitation>
(catalog); Chani-Posse, 2004: 222 (characters), 230 (list).
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</paragraph>
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<taxonomicName id="4C7E4D5CFFABFF9BFF1BCFB4C8CDE3E8" ID-CoL="76XKJ" authority="Bernhauer, 1916: 36" authorityName="Bernhauer" authorityPageNumber="36" authorityYear="1916" box="[151,677,708,731]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFF1BCFB4CBC6E3E8" box="[151,430,708,731]" italics="true" pageId="7" pageNumber="38">Philonthus catamarcanus</emphasis>
<treatmentCitation id="0ADF10CEFFABFF9BFE37CFB4C8CDE3E8" author="Bernhauer" box="[443,677,708,731]" page="36" pageId="7" pageNumber="38" year="1916">
<bibRefCitation id="EFEF4B2EFFABFF9BFE37CFB4C81EE3E8" author="Bernhauer" box="[443,630,708,731]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer, 1916</bibRefCitation>
: 36
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</taxonomicName>
;
<treatmentCitation id="0ADF10CEFFABFF9BFD3BCFB4C9AFE3E8" author="Scheerpeltz" box="[695,967,708,731]" page="1335" pageId="7" pageNumber="38" year="1933">
<bibRefCitation id="EFEF4B2EFFABFF9BFD3BCFB4C9E9E3E8" author="Scheerpeltz" box="[695,897,708,731]" pageId="7" pageNumber="42" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1335
</treatmentCitation>
(catalog);
<treatmentCitation id="0ADF10CEFFABFF9BFBC4CFB4CF3CE3E8" author="Blackwelder" box="[1096,1364,708,731]" page="132" pageId="7" pageNumber="38" year="1944">
<bibRefCitation id="EFEF4B2EFFABFF9BFBC4CFB4CF73E3E8" author="Blackwelder" box="[1096,1307,708,731]" pageId="7" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 132
</treatmentCitation>
(list);
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<bibRefCitation id="EFEF4B2EFFABFF9BFF4BCF94CB0BE3C8" author="Herman" box="[199,355,740,763]" pageId="7" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2772
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(catalog); Chani-Posse, 2004: 230 (list),
<emphasis id="B90AEACDFFABFF9BFCD6CF93C9D5E3C8" bold="true" box="[858,957,739,763]" pageId="7" pageNumber="38">syn. nov.</emphasis>
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<taxonomicName id="4C7E4D5CFFABFF9BFF1BCE74C833E228" ID-CoL="4GC58" authority="Bernhauer, 1921: 169" authorityName="Bernhauer" authorityPageNumber="169" authorityYear="1921" box="[151,603,772,795]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFF1BCE74CB09E228" box="[151,353,772,795]" italics="true" pageId="7" pageNumber="38">Philonthus weiseri</emphasis>
<treatmentCitation id="0ADF10CEFFABFF9BFEE7CE74C833E228" author="Bernhauer" box="[363,603,772,795]" page="169" pageId="7" pageNumber="38" year="1921">
<bibRefCitation id="EFEF4B2EFFABFF9BFEE7CE74C84BE228" author="Bernhauer" box="[363,547,772,795]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
: 169
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;
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: 1368
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(catalog);
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: 135
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(list);
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<bibRefCitation id="EFEF4B2EFFABFF9BFAB3CE74CB68E208" author="Herman" pageId="7" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3003
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(catalog); Chani-Posse, 2004: 232 (list),
<emphasis id="B90AEACDFFABFF9BFD7BCE53C932E208" bold="true" box="[759,858,803,827]" pageId="7" pageNumber="38">syn. nov.</emphasis>
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<typeStatus id="54C5887DFFABFF9BFF1BCE1ACABCE2B7" box="[151,212,874,900]" pageId="7" pageNumber="38">Type</typeStatus>
material.
<typeStatus id="54C5887DFFABFF9BFED9CE1ACBB9E2B7" box="[341,465,874,900]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
</emphasis>
of
<taxonomicName id="4C7E4D5CFFABFF9BFE76CE1AC962E2B0" box="[506,778,874,899]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFE76CE1AC962E2B0" box="[506,778,874,899]" italics="true" pageId="7" pageNumber="38">Philonthus bonariensis</emphasis>
</taxonomicName>
(here designated;
<figureCitation id="13452A5AFFABFF9BFC68CE1ACE5CE2B7" box="[996,1076,874,900]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="7" pageNumber="38">Figs. 7</figureCitation>
ab), 3, with labels: “Buenos Aires /
<date id="FFC0101FFFABFF9BFF78CEE1CB21E298" box="[244,329,913,939]" pageId="7" pageNumber="38" value="1998-12">XII.98</date>
.
<date id="FFC0101FFFABFF9BFED9CEE1CBF2E298" box="[341,410,913,939]" pageId="7" pageNumber="38" value="1999-03">III.99</date>
/ Silvestri”, “
<taxonomicName id="4C7E4D5CFFABFF9BFDC4CEE1C966E298" authority="Brh" authorityName="Brh" box="[584,782,913,939]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFDC4CEE1C8B0E299" box="[584,728,913,938]" italics="true" pageId="7" pageNumber="38">bonariensis</emphasis>
Brh
</taxonomicName>
n. sp / det. Bernhauer”, “
<taxonomicName id="4C7E4D5CFFABFF9BFBC1CEE1CF4AE298" authority="Brnh" authorityName="Brnh" box="[1101,1314,913,939]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFBC1CEE1CEB5E299" box="[1101,1245,913,938]" italics="true" pageId="7" pageNumber="38">bonariensis</emphasis>
Brnh
</taxonomicName>
/
<typeStatus id="54C5887DFFABFF9BFAB1CEE1CFE5E298" box="[1341,1421,913,939]" pageId="7" pageNumber="38">Typus</typeStatus>
”, “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFABFF9BFD68CEC8C937E2E1" box="[740,863,952,978]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFABFF9BFCEBCEC8CF25E2E1" authority="Bernhauer, 1909" authorityName="Bernhauer" authorityYear="1909" box="[871,1357,952,978]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFCEBCEC8CE16E2E2" box="[871,1150,952,977]" italics="true" pageId="7" pageNumber="38">Philonthus bonariensis</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFB0ACEC8CF25E2E1" author="Bernhauer" box="[1158,1357,952,978]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1909) Zur Staphylinidenfauna von Sudamerika (6. Beitrag.). Bolletino della Societa Entomologica Italiana, 40 (1908), 225 - 251." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFABFF9BFE7FCEAEC827E2CB" LSID="urn:lsid:biocol.org:col:34795" box="[499,591,990,1016]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFABFF9BFF4AC975CE3AE5D3" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
<emphasis id="B90AEACDFFABFF9BFF4AC975CB2AE52C" bold="true" box="[198,322,1029,1055]" pageId="7" pageNumber="38">
<typeStatus id="54C5887DFFABFF9BFF4AC975CB2AE52C" box="[198,322,1029,1055]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
</emphasis>
of
<taxonomicName id="4C7E4D5CFFABFF9BFEFFC975C83BE52D" box="[371,595,1029,1054]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFEFFC975C83BE52D" box="[371,595,1029,1054]" italics="true" pageId="7" pageNumber="38">Philonthus weiseri</emphasis>
</taxonomicName>
(
<figureCitation id="13452A5AFFABFF9BFDEBC975C8D6E52C" box="[615,702,1029,1055]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="7" pageNumber="38">Figs. 8</figureCitation>
ab), 3, with labels: “Rep.
<collectingCountry id="F369764FFFABFF9BFBA8C975CEF0E52C" box="[1060,1176,1029,1055]" name="Argentina" pageId="7" pageNumber="38">Argentina</collectingCountry>
/ Jujuy / C. Bruch”, “Cueva Iturbe /
<quantity id="4C869B3AFFABFF9BFED8C95CCBC6E575" box="[340,430,1068,1094]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="38" unit="m" value="3700.0">3700 m</quantity>
<date id="FFC0101FFFABFF9BFE3AC95CCB95E575" box="[438,509,1068,1094]" pageId="7" pageNumber="38">10.XI.</date>
919 / lg. Weiser”, “
<taxonomicName id="4C7E4D5CFFABFF9BFD68C95CC9E0E575" authority="Bernh" authorityName="Bernh" box="[740,904,1068,1094]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFD68C95CC951E576" box="[740,825,1068,1093]" italics="true" pageId="7" pageNumber="38">weiseri</emphasis>
Bernh
</taxonomicName>
/
<typeStatus id="54C5887DFFABFF9BFC2DC95CC986E575" box="[929,1006,1068,1094]" pageId="7" pageNumber="38">Typus</typeStatus>
”, “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFABFF9BFEB0C922CBDFE55F" box="[316,439,1106,1132]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFABFF9BFE4CC923C91AE55F" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" box="[448,882,1106,1132]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFE4CC923C8CAE55F" box="[448,674,1107,1132]" italics="true" pageId="7" pageNumber="38">Philonthus weiseri</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFD26C922C91AE55F" author="Bernhauer" box="[682,882,1106,1132]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFABFF9BFABEC922CFF8E55F" LSID="urn:lsid:biocol.org:col:34795" box="[1330,1424,1106,1132]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38">FMNH</collectionCode>
).
<emphasis id="B90AEACDFFABFF9BFF1BC908CB31E5A1" bold="true" box="[151,345,1144,1170]" pageId="7" pageNumber="38">
<typeStatus id="54C5887DFFABFF9BFF1BC908CB3CE5A1" box="[151,340,1144,1170]" pageId="7" pageNumber="38" type="paralectotype">Paralectotypes</typeStatus>
:
</emphasis>
3, Ƥ, with labels: “Rep.
<collectingCountry id="F369764FFFABFF9BFD04C909C895E5A0" box="[648,765,1145,1171]" name="Argentina" pageId="7" pageNumber="38">Argentina</collectingCountry>
/ Jujuy / C. Bruch”, “Cueva Iturbe /
<quantity id="4C869B3AFFABFF9BFB3FC909CF64E5A0" box="[1203,1292,1145,1171]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="38" unit="m" value="3700.0">3700 m</quantity>
<date id="FFC0101FFFABFF9BFA99C909CF34E5A0" box="[1301,1372,1145,1171]" pageId="7" pageNumber="38">10.XI.</date>
919 / lg. Weiser”, “
<taxonomicName id="4C7E4D5CFFABFF9BFECFC9D0CB99E589" authority="Bernh" authorityName="Bernh" box="[323,497,1184,1210]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFECFC9D0CBF5E58A" box="[323,413,1184,1209]" italics="true" pageId="7" pageNumber="38">weiseri</emphasis>
Bernh
</taxonomicName>
/ Coypus”, “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFABFF9BFB63C9D0CFF5E589" box="[1263,1437,1184,1210]" pageId="7" pageNumber="38" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFABFF9BFF1BC9B7C854E5D3" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" box="[151,572,1222,1248]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="weiseri">
<emphasis id="B90AEACDFFABFF9BFF1BC9B7CB19E5D3" box="[151,369,1223,1248]" italics="true" pageId="7" pageNumber="38">Philonthus weiseri</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFEF5C9B6C854E5D3" author="Bernhauer" box="[377,572,1222,1248]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFABFF9BFC66C9B6CE2EE5D3" LSID="urn:lsid:biocol.org:col:34795" box="[1002,1094,1222,1248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="38">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFABFF9BFF4AC99DC918E4FB" pageId="7" pageNumber="38" type="description">
<paragraph id="8BC136DFFFABFF9BFF4AC99DC918E4FB" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
<emphasis id="B90AEACDFFABFF9BFF4AC99DCB2AE434" bold="true" box="[198,322,1261,1287]" pageId="7" pageNumber="38">
<typeStatus id="54C5887DFFABFF9BFF4AC99DCB2AE434" box="[198,322,1261,1287]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
</emphasis>
of
<taxonomicName id="4C7E4D5CFFABFF9BFEE0C99DC8F0E435" box="[364,664,1261,1286]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFEE0C99DC8F0E435" box="[364,664,1261,1286]" italics="true" pageId="7" pageNumber="38">Philonthus catamarcanus</emphasis>
</taxonomicName>
(here designated;
<figureCitation id="13452A5AFFABFF9BFCF8C99DC9AEE434" box="[884,966,1261,1287]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="7" pageNumber="38">Figs. 9</figureCitation>
ab), Ƥ, with labels: “Rep.
<collectingCountry id="F369764FFFABFF9BFA9AC99DCFE2E434" box="[1302,1418,1261,1287]" name="Argentina" pageId="7" pageNumber="38">Argentina</collectingCountry>
/ Prov. Catamarca /
<date id="FFC0101FFFABFF9BFEF9C864CB92E41D" box="[373,506,1300,1326]" pageId="7" pageNumber="38" value="1912-11-19">19.11.1912</date>
/ C. Bruch”, “
<taxonomicName id="4C7E4D5CFFABFF9BFD25C864C9C1E41D" authority="Bernh." authorityName="Bernh." box="[681,937,1300,1326]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFD25C864C938E41E" box="[681,848,1300,1325]" italics="true" pageId="7" pageNumber="38">catamarcanus</emphasis>
Bernh.
</taxonomicName>
/
<typeStatus id="54C5887DFFABFF9BFC4EC864CE78E41D" box="[962,1040,1300,1326]" pageId="7" pageNumber="38">Typus</typeStatus>
”, “Chicago NHMus. / Bernhauer Collection”, “
<typeStatus id="54C5887DFFABFF9BFECEC84ACBA9E467" box="[322,449,1338,1364]" pageId="7" pageNumber="38" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFABFF9BFE46C84BC9B7E467" authority="Bernhauer, 1916" authorityName="Bernhauer" authorityYear="1916" box="[458,991,1338,1364]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFE46C84BC960E467" box="[458,776,1339,1364]" italics="true" pageId="7" pageNumber="38">Philonthus catamarcanus</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFC9DC84AC9B7E467" author="Bernhauer" box="[785,991,1338,1364]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer, 1916</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (FMNH).
<emphasis id="B90AEACDFFABFF9BFE9BC810CBB4E449" bold="true" box="[279,476,1376,1402]" pageId="7" pageNumber="38">
<typeStatus id="54C5887DFFABFF9BFE9BC810CBBEE449" box="[279,470,1376,1402]" pageId="7" pageNumber="38" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
Ƥ, with labels: “San Luis / 190”, “C. Bruch”, “
<taxonomicName id="4C7E4D5CFFABFF9BFBCFC811CE9CE449" box="[1091,1268,1377,1402]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFBCFC811CE9CE449" box="[1091,1268,1377,1402]" italics="true" pageId="7" pageNumber="38">catamarcanus</emphasis>
</taxonomicName>
det. Bernh. / Cotypus”, “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFABFF9BFC14C8F8CE23E491" box="[920,1099,1416,1442]" pageId="7" pageNumber="38" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFABFF9BFBD9C8F8CB31E4FB" authority="Bernhauer, 1916" authorityName="Bernhauer" authorityYear="1916" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFBD9C8F8CFF5E492" box="[1109,1437,1416,1441]" italics="true" pageId="7" pageNumber="38">Philonthus catamarcanus</emphasis>
<bibRefCitation id="EFEF4B2EFFABFF9BFF1BC8DECB31E4FB" author="Bernhauer" box="[151,345,1454,1480]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer, 1916</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (FMNH).
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFABFF94FF4AC8A5C9DCE181" lastPageId="8" lastPageNumber="39" pageId="7" pageNumber="38" type="diagnosis">
<paragraph id="8BC136DFFFABFF9BFF4AC8A5CFFBE750" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
<emphasis id="B90AEACDFFABFF9BFF4AC8A5CB2EE4DC" bold="true" box="[198,326,1493,1519]" pageId="7" pageNumber="38">Diagnosis.</emphasis>
<taxonomicName id="4C7E4D5CFFABFF9BFEC2C8A5C837E4DD" box="[334,607,1493,1518]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFEC2C8A5C837E4DD" box="[334,607,1493,1518]" italics="true" pageId="7" pageNumber="38">Philonthus bonariensis</emphasis>
</taxonomicName>
may be identified by the antennal segments 4 and 5 elongate, the last segment of labial palpi 2 times as long as the second, the abdominal tergum 10 with 5 to 6 apical setae in both sexes, the median lobe with a small tooth on the face adjacent to paramere, and the paramere elongate with 6 peg setae forming two short lateral rows at apex. For further details and illustrations see Chani-Posse (2004).
</paragraph>
<paragraph id="8BC136DFFFABFF9BFF4ACB1FCF2CE7CD" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
<emphasis id="B90AEACDFFABFF9BFF4ACB1FCB7CE7BA" bold="true" box="[198,276,1647,1673]" pageId="7" pageNumber="38">Notes.</emphasis>
I designate the
<typeStatus id="54C5887DFFABFF9BFE68CB00C838E7B9" box="[484,592,1648,1674]" pageId="7" pageNumber="38" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C7E4D5CFFABFF9BFD0ECB00C946E7BA" box="[642,814,1648,1673]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFABFF9BFD0ECB00C946E7BA" box="[642,814,1648,1673]" italics="true" pageId="7" pageNumber="38">P. bonariensis</emphasis>
</taxonomicName>
rather than assume that the examined
<typeStatus id="54C5887DFFABFF9BFA94CB00CF22E7B9" box="[1304,1354,1648,1674]" pageId="7" pageNumber="38">type</typeStatus>
is the
<typeStatus id="54C5887DFFABFF9BFF1BCBE6CB60E783" box="[151,264,1686,1712]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
, since in the original description
<bibRefCitation id="EFEF4B2EFFABFF9BFD23CBE6C9E4E783" author="Bernhauer" box="[687,908,1686,1712]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1909) Zur Staphylinidenfauna von Sudamerika (6. Beitrag.). Bolletino della Societa Entomologica Italiana, 40 (1908), 225 - 251." type="journal article" year="1909">Bernhauer (1909)</bibRefCitation>
gave no information about the number of specimens examined. Therefore, it is possible that this name was based on more than one specimen (
<bibRefCitation id="EFEF4B2EFFABFF9BFAD9CBCDCB80E7CD" author="ICZN" pageId="7" pageNumber="42" refString="ICZN (1999) International Code of Zoological Nomenclatur e. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999, Recommendation 73F</bibRefCitation>
). The
<typeStatus id="54C5887DFFABFF9BFDBDCB94C8F4E7CD" box="[561,668,1764,1790]" pageId="7" pageNumber="38" type="lectotype">lectotype</typeStatus>
agrees with the original description by
<bibRefCitation id="EFEF4B2EFFABFF9BFBE1CB94CF28E7CD" author="Bernhauer" box="[1133,1344,1764,1790]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1909) Zur Staphylinidenfauna von Sudamerika (6. Beitrag.). Bolletino della Societa Entomologica Italiana, 40 (1908), 225 - 251." type="journal article" year="1909">Bernhauer (1909)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC136DFFFABFF9BFF4ACA7AC9E8E641" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
In the original description of
<emphasis id="B90AEACDFFABFF9BFD97CA7BC8E0E617" box="[539,648,1803,1828]" italics="true" pageId="7" pageNumber="38">P. w e i s e r i</emphasis>
there is no indication about how many specimens were included in the
<typeStatus id="54C5887DFFABFF9BFF4ACA41CA94E678" box="[198,252,1841,1867]" pageId="7" pageNumber="38">type</typeStatus>
series. All three specimens listed above agree with the description by
<bibRefCitation id="EFEF4B2EFFABFF9BFBF1CA41CF37E678" author="Bernhauer" box="[1149,1375,1841,1867]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer (1921)</bibRefCitation>
. The specimen in better conditions is hereby designated as
<typeStatus id="54C5887DFFABFF9BFC80CA28C913E641" box="[780,891,1880,1906]" pageId="7" pageNumber="38" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC136DFFFABFF9BFF4ACA0EC80DE6D5" blockId="7.[151,1437,874,2022]" pageId="7" pageNumber="38">
In the original description of
<taxonomicName id="4C7E4D5CFFABFF9BFDA2CA0FC89BE6AB" box="[558,755,1919,1944]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFABFF9BFDA2CA0FC89BE6AB" box="[558,755,1919,1944]" italics="true" pageId="7" pageNumber="38">P. catamarcanus</emphasis>
</taxonomicName>
there is no indication about how many specimens were examined. Both specimens listed above agree with the description by
<bibRefCitation id="EFEF4B2EFFABFF9BFB8BCAD5CE8CE68C" author="Bernhauer" box="[1031,1252,1957,1983]" pageId="7" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer (1916)</bibRefCitation>
who gave two different
<typeStatus id="54C5887DFFABFF9BFE8FCABCCB5DE6D5" box="[259,309,1996,2022]" pageId="7" pageNumber="38">type</typeStatus>
localities for this species.
</paragraph>
<paragraph id="8BC136DFFFA4FF94FF1BCDE8C9DCE181" blockId="8.[151,1436,152,410]" box="[151,948,152,178]" pageId="8" pageNumber="39">
The specimen in better conditions is hereby designated as
<typeStatus id="54C5887DFFA4FF94FCCDCDE8C9D8E181" box="[833,944,152,178]" pageId="8" pageNumber="39" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFA4FF94FF4ACDCEC9D2E0A9" pageId="8" pageNumber="39" type="discussion">
<paragraph id="8BC136DFFFA4FF94FF4ACDCEC9D2E0A9" blockId="8.[151,1436,152,410]" pageId="8" pageNumber="39">
<emphasis id="B90AEACDFFA4FF94FF4ACDCECB3DE1EB" bold="true" box="[198,341,190,216]" pageId="8" pageNumber="39">Comments.</emphasis>
The
<typeStatus id="54C5887DFFA4FF94FE1CCDCECBA6E1EB" box="[400,462,190,216]" pageId="8" pageNumber="39">types</typeStatus>
of
<taxonomicName id="4C7E4D5CFFA4FF94FE79CDCFC8F3E1EB" box="[501,667,191,216]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFA4FF94FE79CDCFC8F3E1EB" box="[501,667,191,216]" italics="true" pageId="8" pageNumber="39">P. bonariensis</emphasis>
</taxonomicName>
,
<emphasis id="B90AEACDFFA4FF94FD25CDCFC97EE1EB" box="[681,790,191,216]" italics="true" pageId="8" pageNumber="39">P. w e i s e r i</emphasis>
and
<taxonomicName id="4C7E4D5CFFA4FF94FCDECDCFCE7CE1EB" box="[850,1044,191,216]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA4FF94FCDECDCFCE7CE1EB" box="[850,1044,191,216]" italics="true" pageId="8" pageNumber="39">P. catamarcanus</emphasis>
</taxonomicName>
(
<figureCitation id="13452A5AFFA4FF94FBA8CDCECE1DE1EB" box="[1060,1141,190,216]" captionStart="FIGURES 1 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a b) Philonthus quadraticeps (lectotype with labels *). (6 a b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a b) Philonthus bonariensis (lectotype with labels *). (8 a b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="8" pageNumber="39">Figs. 7</figureCitation>
ab, 8 ab, 9 ab) are all similar in external characters, and identical in characters of the male and female genitalia. All these names are here considered to be synonyms. The body size and the elytral color pattern vary in examined specimens but this variation is continuous. The elytral color pattern may range from piceous to piceous-brunneous with suture and apical margin of each elytron paler, as stated by Chani-Posse (2004) in her redescription of
<taxonomicName id="4C7E4D5CFFA4FF94FA0ACC29CB48E0AA" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B90AEACDFFA4FF94FA0ACC29CB48E0AA" italics="true" pageId="8" pageNumber="39">P. bonariensis</emphasis>
</taxonomicName>
, to one-colored elytra, brunneous to testaceo-brunneous.
</paragraph>
</subSubSection>
</treatment>
</document>