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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.119781" ID-Pensoft-Pub="1313-2970-1198-17" ID-Pensoft-UUID="9A0A0A5CD22451C7ADAE5C8460C568DC" ID-ZooBank="BDC5B2CD1BB442AE8672E57CC0FBBF6F" ModsDocID="1313-2970-1198-17" checkinTime="1713962616471" checkinUser="pensoft" docAuthor="Dyachkov, Yurii V. &amp; Bonato, Lucio" docDate="2024" docId="B9897045E6415D7E9857F8FC4CDA1A0A" docLanguage="en" docName="ZooKeys 1198: 17-54" docOrigin="ZooKeys 1198" docPubDate="2024-04-23" docSource="http://dx.doi.org/10.3897/zookeys.1198.119781" docTitle="Escaryus sibiricus Cook 1899" docType="treatment" docVersion="1" id="9A0A0A5CD22451C7ADAE5C8460C568DC" lastPageNumber="17" masterDocId="9A0A0A5CD22451C7ADAE5C8460C568DC" masterDocTitle="An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia" masterLastPageNumber="54" masterPageNumber="17" pageNumber="17" updateTime="1713962616471" updateUser="pensoft">
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<mods:title id="C3D897D9FBBB8C5E7655F83DE7E90C15">An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia</mods:title>
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<mods:namePart id="A603E70BF7CF5DC9683349EFB198C638">Dyachkov, Yurii V.</mods:namePart>
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<mods:affiliation id="E93395CC2211A75135DE6BA08C146078">Altai State University, Lenin Avenue, 61, 656049, Barnaul, Russia &amp; Tomsk State University, Lenin Avenue, 36, 634050, Tomsk, Russia &amp; Western Caspian University, Istiglaliyyat Street, 31, Baku, Azerbaijan</mods:affiliation>
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<mods:namePart id="31E3724B0A3A46D99DD9203A06E16200">Bonato, Lucio</mods:namePart>
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<mods:affiliation id="33CEFDBC2F46A6A3FB1CBC2F10D15690">Dipartimento di Biologia, Universita di Padova, via U. Bassi 58 b, 35131 Padova, Italy</mods:affiliation>
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<treatment id="B9897045E6415D7E9857F8FC4CDA1A0A" LSID="urn:lsid:plazi:treatment:B9897045E6415D7E9857F8FC4CDA1A0A" httpUri="http://treatment.plazi.org/id/B9897045E6415D7E9857F8FC4CDA1A0A" lastPageNumber="17" pageId="0" pageNumber="17">
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<paragraph id="68B5BC625DEDDAB9F68A9FDEF09C6B4C" pageId="0" pageNumber="17">
37.
<taxonomicName id="9CFECC5C2A9AD12C68FAE61D19D98A63" ID-CoL="6GT9F" LSID="B9897045-E641-5D7E-9857-F8FC4CDA1A0A" authority="Cook, 1899" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">Escaryus sibiricus Cook, 1899</taxonomicName>
</paragraph>
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<subSubSection id="B0705314A266013481F935B735261850" pageId="0" pageNumber="17" type="reference_group">
<paragraph id="F4732396B79D389C46552749348B8235" pageId="0" pageNumber="17">
<taxonomicName id="02736837514A2CF3A1C70ECC76D33735" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">Escaryus sibiricus</taxonomicName>
<bibRefCitation id="5D006D400B5A5FC9159F4D2585BCF3E8" author="Cook, OF" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="17" pagination="303 - 312" refId="B26" refString="Cook, OF, 1899. The Geophiloidea of Florida Keys. Proceedings of the Entomological Society of Washington 4: 303 - 312" title="The Geophiloidea of Florida Keys." volume="4" year="1899">Cook 1899</bibRefCitation>
: 304.
</paragraph>
<paragraph id="8381755D566117645DBA6910D448D6CF" pageId="0" pageNumber="17">
<taxonomicName id="ED24D1E3D4B343D4FE2C598F76CF100B" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">Escaryus sibiricus</taxonomicName>
-
<bibRefCitation id="EE64196334E6D797DE1F7B0DE8C2BE2D" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems 1904</bibRefCitation>
: 122;
<bibRefCitation id="AAF48B0C2FB48110473B8B5DD1DD47EC" author="Attems, C" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="17" pagination="291 - 305" refId="B6" refString="Attems, C, 1927. Neue Chilopoden. Zoologischer Anzeiger 72: 291 - 305" title="Neue Chilopoden." volume="72" year="1927">1927</bibRefCitation>
: 301;
<bibRefCitation id="A83B54424FC5078E4B6C4EB271B95DCF" DOI="https://doi.org/10.1515/9783111430638" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="17" pagination="1 - 388" refId="B7" refString="Attems, C, 1929. Myriapoda. 1. Geophilomorpha. Das Tierreich 52: 1 - 388, DOI: https://doi.org/10.1515/9783111430638" title="Myriapoda. 1. Geophilomorpha." url="https://doi.org/10.1515/9783111430638" volume="52" year="1929">1929</bibRefCitation>
: 95;
<bibRefCitation id="D6CF5F8F2A1FD0E25DD40D992342C896" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
: 135;
<bibRefCitation id="D4714D62668379A042341AB96B9B4B9B" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
: 94, 113;
<bibRefCitation id="6280A86DA762F6FA0CE923F5142EA8B6" author="Ganin, GN" journalOrPublisher="Dalnauka, Vladivostok-Khabarovsk" pageId="0" pageNumber="17" refId="B48" refString="Ganin, GN, 1997. Soil Invertebrates of the Ussuri Taiga. Dalnauka, Vladivostok-Khabarovsk" title="Soil Invertebrates of the Ussuri Taiga." year="1997">Ganin 1997</bibRefCitation>
: 141;
<bibRefCitation id="342224F7AF202243AA7D16E21F27CDB5" DOI="https://doi.org/10.11646/zootaxa.4977.1.1" author="Thofern, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="17" pagination="1 - 103" refId="B101" refString="Thofern, D, Duperre, N, Harms, D, 2021. An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg. Zootaxa 4977 (1): 1 - 103, DOI: https://doi.org/10.11646/zootaxa.4977.1.1" title="An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg." url="https://doi.org/10.11646/zootaxa.4977.1.1" volume="4977" year="2021">Thofern et al. 2021</bibRefCitation>
: 30.
</paragraph>
</subSubSection>
<subSubSection id="8390975599349C02AABEA4B61A7CC677" pageId="0" pageNumber="17" type="type locality">
<paragraph id="A134EFE392C000C5C2B5F84DFEC69EF5" pageId="0" pageNumber="17">Type locality.</paragraph>
<paragraph id="9A7D35197BC2BE2DD8063EF758A1B8DE" pageId="0" pageNumber="17">
Russia: Maritime krai:
<normalizedToken id="463D6DB7A595873AA59E95454879614A" originalValue="“Vladivostock”">&quot;Vladivostock&quot;</normalizedToken>
(
<bibRefCitation id="299226B43EF172E46F723B14F45A3882" author="Cook, OF" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="17" pagination="303 - 312" refId="B26" refString="Cook, OF, 1899. The Geophiloidea of Florida Keys. Proceedings of the Entomological Society of Washington 4: 303 - 312" title="The Geophiloidea of Florida Keys." volume="4" year="1899">Cook 1899</bibRefCitation>
) = Vladivostok city,
<geoCoordinate id="8E6C04C69D43DEB7259AE71E00A36E52" degrees="43" direction="north" minutes="7" orientation="latitude" precision="925" value="43.116665">43°7'N</geoCoordinate>
,
<geoCoordinate id="D984E8B68E0BA92BB169302EB427A86D" degrees="131" direction="east" minutes="54" orientation="longitude" precision="925" value="131.9">131°54'E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="B45C09805402E1F040694B0B096A0940" pageId="0" pageNumber="17" type="type series">
<paragraph id="44C35149C8BD6FADAE143AA4623A168F" pageId="0" pageNumber="17">Type series.</paragraph>
<paragraph id="9610F9AA1D753268DF989CA7A69B93F2" pageId="0" pageNumber="17">
<emphasis id="7F89109F1E597F266D9B993304F6C0FB" bold="true" italics="true" pageId="0" pageNumber="17">Syntypes</emphasis>
: 12 specimens, both sexes, including 10 specimens deposited in ZMH (
<bibRefCitation id="487AC235D555FC97D41423DD64F8D873" DOI="https://doi.org/10.11646/zootaxa.4977.1.1" author="Thofern, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="17" pagination="1 - 103" refId="B101" refString="Thofern, D, Duperre, N, Harms, D, 2021. An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg. Zootaxa 4977 (1): 1 - 103, DOI: https://doi.org/10.11646/zootaxa.4977.1.1" title="An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg." url="https://doi.org/10.11646/zootaxa.4977.1.1" volume="4977" year="2021">Thofern et al. 2021</bibRefCitation>
) and 2 specimens deposited in NHMW (
<bibRefCitation id="3C8649EF149FE8F61E75F004BFB25220" author="Ilie, V" journalOrPublisher="Verlag des Naturhistorischen Museum in Vienna" pageId="0" pageNumber="17" pagination="1 - 75" refId="B55" refString="Ilie, V, Schiller, E, Stagl, V, 2009. Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien. Verlag des Naturhistorischen Museum in Vienna 22 (4): 1 - 75" title="Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien." volume="22" year="2009">Ilie et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8CDAE4CF057F0D937F83939A35AB456B" pageId="0" pageNumber="17" type="diagnosis">
<paragraph id="B7B6819FFE4203E530E51BC3EE5ED0D2" pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph id="69411AFE87E346D6BE504B2A43B0BF8B" pageId="0" pageNumber="17">
An
<taxonomicName id="96E3B7C0E576CE950042AEFC5429D7A7" authorityName="Cook &amp; Collins" authorityYear="1891" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="77A4729D9B8AAAD48CD12FE3FCF93B99" italics="true" pageId="0" pageNumber="17">Escaryus</emphasis>
</taxonomicName>
species with body length reaching 6.5 cm; labral arc relatively shallow, with middle denticles obtuse and lateral ones long and pointed; first maxillae with two pairs of lappets; forcipular trochanteroprefemur with a small distal bulge, other forcipular articles without denticles; 49-51 leg-bearing segments; metasternite of the ultimate leg-bearing segment rectangular, ~ 2
<normalizedToken id="E078F671F24AEFBABEB45B74F41AEBBE" originalValue="×">x</normalizedToken>
as long as wide; coxal pores of different size, on both ventral and lateral sides, including a pair of much larger ventral pores on each coxopleuron; anal pores absent.
</paragraph>
</subSubSection>
<subSubSection id="0B5AFE8E7F4EFAD5AD2082B8C2E9B9FF" pageId="0" pageNumber="17" type="distribution">
<paragraph id="995259F81CF9B65F74BFD988437CBA6C" pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph id="4FD8B7C47D7574C83615FD6C3C6F41C8" pageId="0" pageNumber="17">
Far East: Maritime krai (
<bibRefCitation id="4A4D3B02299833941603AF5999A11FEE" author="Cook, OF" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="17" pagination="303 - 312" refId="B26" refString="Cook, OF, 1899. The Geophiloidea of Florida Keys. Proceedings of the Entomological Society of Washington 4: 303 - 312" title="The Geophiloidea of Florida Keys." volume="4" year="1899">Cook 1899</bibRefCitation>
;
<bibRefCitation id="54D3FA7751482A2C76BA855E30B436F0" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems 1904</bibRefCitation>
) and Amur oblast (
<bibRefCitation id="4715069102D0F2F782A87209A90F1640" author="Ganin, GN" journalOrPublisher="Dalnauka, Vladivostok-Khabarovsk" pageId="0" pageNumber="17" refId="B48" refString="Ganin, GN, 1997. Soil Invertebrates of the Ussuri Taiga. Dalnauka, Vladivostok-Khabarovsk" title="Soil Invertebrates of the Ussuri Taiga." year="1997">Ganin 1997</bibRefCitation>
). Outside Asian Russia: no records.
</paragraph>
</subSubSection>
<subSubSection id="C72A4CA7D87695B1BADD4484DBD5CF2C" pageId="0" pageNumber="17" type="remarks">
<paragraph id="D5373BF3660FF33D0A38A9EA973113FC" pageId="0" pageNumber="17">Remarks.</paragraph>
<paragraph id="2BB17AD96D84275AFDF8069C0187CB4A" pageId="0" pageNumber="17">
<bibRefCitation id="537C6D610689D0F38B57D49112672163" author="Titova, LP" editor="Aleinikova, MM" journalOrPublisher="Materials of the 3 th All-Union Conference, Kazan. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="165 - 166" refId="B102" refString="Titova, LP, 1969. Geophilids of the USSR fauna and news in the distribution of the fam. Mecistocephalidae. In: Aleinikova, MM, Ed., Problems of soil zoology. Materials of the 3th All-Union Conference, Kazan. Nauka Publisher, Moscow: 165 - 166" title="Geophilids of the USSR fauna and news in the distribution of the fam. Mecistocephalidae." volumeTitle="Problems of soil zoology." year="1969">Titova (1969)</bibRefCitation>
reported this species from Western Siberia (Kemerovo oblast) but later (
<bibRefCitation id="463BC81372F5D5F1396111EAB865C9C2" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Titova 1973</bibRefCitation>
) she regarded the same record as
<taxonomicName id="9116574025F52271D624EB9DE215A794" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="9C8DEC844E8423D49D2DB1464C1E92F8" italics="true" pageId="0" pageNumber="17">E. koreanus</emphasis>
</taxonomicName>
, even though she also suggested that the latter species can be a junior synonym of
<taxonomicName id="D973788A5C0206C893C22D3DBE2B5EC7" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">
<emphasis id="8682B18150B08E3A22FD872050D48DDE" italics="true" pageId="0" pageNumber="17">E. sibiricus</emphasis>
</taxonomicName>
. The anal pores were indicated as absent by
<bibRefCitation id="D0B942DCA8F5E484E01E1E41CBC2875C" author="Cook, OF" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="17" pagination="303 - 312" refId="B26" refString="Cook, OF, 1899. The Geophiloidea of Florida Keys. Proceedings of the Entomological Society of Washington 4: 303 - 312" title="The Geophiloidea of Florida Keys." volume="4" year="1899">Cook (1899)</bibRefCitation>
and
<bibRefCitation id="41A917C652C719B14A59E3083C897E1C" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems (1904)</bibRefCitation>
but illustrated as present in specimens identified as
<taxonomicName id="7A4CBA2F67025FC8AC020BC3D23C3A76" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">
<emphasis id="0F46AE19D3E59804DE54BA6AB7A843ED" italics="true" pageId="0" pageNumber="17">E. sibiricus</emphasis>
</taxonomicName>
by
<bibRefCitation id="65F4D6BEDEB6E86534D439247E8779D9" DOI="https://doi.org/10.11646/zootaxa.4977.1.1" author="Thofern, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="17" pagination="1 - 103" refId="B101" refString="Thofern, D, Duperre, N, Harms, D, 2021. An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg. Zootaxa 4977 (1): 1 - 103, DOI: https://doi.org/10.11646/zootaxa.4977.1.1" title="An annotated type catalogue of the centipedes (Myriapoda: Chilopoda) held in the Zoological Museum Hamburg." url="https://doi.org/10.11646/zootaxa.4977.1.1" volume="4977" year="2021">Thofern et al. (2021)</bibRefCitation>
, and they are known to be present in
<taxonomicName id="6834E4DA878620C4661E4C7C66BBFDD0" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="FFA7DB0FFAC3FCB5C8C2E189E7028AFA" italics="true" pageId="0" pageNumber="17">E. koreanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>