134 lines
18 KiB
XML
134 lines
18 KiB
XML
<document id="E1EF9CD6C89594A1AE709692EB35D377" ENCODING="UTF-8" ID-CLB-Dataset="52945" ID-GBIF-Dataset="b185fadf-4822-445b-a074-a56fdb2f718a" ModsDocAuthor="Borowiec, M. L." ModsDocID="21366" checkinTime="1244213982275" checkinUser="thomas" docAuthor="Borowiec, M. L." docDate="2007" docId="934972B786C31DD9FB7725AB7B082525" docLanguage="en" docName="21366_gg2.xml" docOrigin="Zootaxa 1642" docSource="http://www.antbase.org/ants/publications/21366/21366.pdf" docTitle="Tyrannomyrmex dux Borowiec, 2007, sp. n." docType="treatment" docVersion="14" lastPageNumber="68" masterDocId="D996C7CB8AB4EC51B8DC755216219072" masterDocTitle="A new species of Tyrannomyrmex (Hymenoptera: Formicidae: Myrmicinae) from India." masterLastPageNumber="65" masterPageNumber="65" pageNumber="66" updateTime="1732482678685" updateUser="ExternalLinkService">
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<mods:title id="1D9F3CF2EA8A1F3E8410468C1C6DD134">A new species of Tyrannomyrmex (Hymenoptera: Formicidae: Myrmicinae) from India.</mods:title>
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<mods:namePart id="100888ACC4B7D0261B2A501FD6AD878E">Borowiec, M. L.</mods:namePart>
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<mods:title id="90F3DDB0B8B831C5AB21603F84D873A6">Zootaxa</mods:title>
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<mods:date id="8F3BA9C5FB1103A72697D571A95E7F37">2007</mods:date>
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<mods:classification id="D02474E0AECCC8F647C6FC74FAAC320C">journal article</mods:classification>
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<mods:identifier id="14F8C593B50218530F35CCDF6C594565" type="HNS-Pub">21366</mods:identifier>
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<mods:identifier id="9604BFA25ECDC6D8928DDFEEA4BF13BF" type="ZooBank">D5F95884-1D25-402E-BAF9-B43A7C0A3FDB</mods:identifier>
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<treatment id="934972B786C31DD9FB7725AB7B082525" ID-DOI="http://doi.org/10.5281/zenodo.6243229" ID-GBIF-Taxon="100125840" ID-Zenodo-Dep="6243229" LSID="urn:lsid:plazi:treatment:934972B786C31DD9FB7725AB7B082525" httpUri="http://treatment.plazi.org/id/934972B786C31DD9FB7725AB7B082525" lastPageNumber="68" pageNumber="66" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Myrmicinae">
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<subSubSection id="B4406C4F271445C7DB256D492A998CD6" type="nomenclature">
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<paragraph id="7A7A5EB7A579021D8CC4D79B7E694D8F" pageNumber="66">
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<taxonomicName id="1800DBA6D13BC40122CC483324DADD09" ID-CoL="CN3KJ" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="66" phylum="Arthropoda" rank="species" species="dux">Tyrannomyrmex dux</taxonomicName>
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<taxonomicNameLabel id="C3BC91832B72A0519156CF96F9576CB1" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4781F221D19697DD8D599385E144C983" type="description">
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<paragraph id="64BC4ACB8504A120F3088D01FE4D29E8" pageNumber="66">(Figs. 1-4)</paragraph>
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<paragraph id="00C52687D062DC86C4571515571C729B" pageNumber="66">Worker measurements: HW 0.52, HL 0.71, EL 0.05, SL 0.56, PrW 0.49, WL 1.03, PL 0.39, PPL 0.21, PW 0.22, PPW 0.31, GL 0.77, TL 3.11, CI 73, SI 107.</paragraph>
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<paragraph id="49D313A38BE19C3110A479645772BB92" pageNumber="66">General appearance as in Fig. 1. Head longer than wide, sides and posterior outline slightly convex. Posterior head corners rounded. Mandibles triangular, masticatory margin edentate except two blunt apical teeth. Palp formula undeterminable. Clypeus convex in anterior part, posteriorly narrowly inserted between frontal lobes. Clypeus lacking carinae. Frontal lobes present and short but prominent, rounded and obscuring antennal sockets. Antennae 11-segmented with feebly defined three segmented club, apical segment being largest and equal in length to rest of funiculus, excluding pedicel. Scape slightly longer than head width, shallowly curved at base. Eyes small, composed of a few feebly defined ommatidia and situated slightly anterior to midlength of head (at 45% of HL).</paragraph>
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<paragraph id="45992588416C283F5682117DC8A41DE6" pageNumber="66">Mesosoma devoid of any kind of grooves or sutures, in side view forming a single broad and continuous convexity. In dorsal view it is widest in promesonotal area, slightly decreasing in width posteriorly. Anteroventral corner of pronotum rounded. Propodeal spiracle round and situated at distance about 3,5 diameters from propodeal margin. Propodeum armed with two very small, triangular, and acute teeth. Metapleural lobes broad, more or less rounded. Metapleural gland orifice not seen at 110X. Petiole without clearly differentiated node or peduncle, node lower than long, in dorsal view longer than wide. Postpetiole wider than long; postpetiolar sternite with transverse keel anteriorly, forming acute tooth in lateral view.</paragraph>
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<paragraph id="7A82CBCF31B6689C30AA8DF3237B13C4" pageNumber="66">Head and mesosoma heavily foveate. Foveae dense, irregular, usually less than 1/3 their diameter apart. Interspaces between foveae smooth and shining on head, sides of promesonotum and dorsal surface of mesosoma; those on katepisternum and sides of propodeum very feebly shagreened, appearing matt. Sculpture of petiole consisting of microreticulum in most of anterior part and irregular foveae on postero-dorsal surface. Sculpture of postpetiole microreticulate. Gaster covered with microreticulum fading before half length of first gastral tergite, remainder of gaster smooth.</paragraph>
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<paragraph id="A495D795FB2EDD525F0E7A335C7DF364" pageNumber="66">Body pilosity dense and abundant, composed of many erect hairs present on the head, meso- and metasoma; hairs on head and mesosoma arising from centers of foveae. Abundant hairs also present on body appendages, being present on antennal scape as well as funiculus; hairs present on all leg segments and on both dorsal and ventral surfaces of mandibles; those on ventral surface not differing visibly in structure from rest of pilosity. Body color reddish, appendages yellowish.</paragraph>
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</subSubSection>
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<subSubSection id="A6FAC81D23D38E5D7D112A288B8F4DF9" type="nomenclature">
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<paragraph id="2B21B7AC06F3DC06F5E38C98100A3BB6" pageNumber="66">Holotype deposited in the Department of Biodiversity and Evolutionary Taxonomy, Zoological Institute, University of Wroclaw, Przybyszewskiego 63/77, 51-148 Wroclaw, Poland.</paragraph>
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<paragraph id="9E9881B0E7C858C208049B4F1225DDC5" pageNumber="67">
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Holotype worker bears the following labels: white printed label
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<materialsCitation id="8075FB2B29F9158FE8E4620882D09F50" ID-GBIF-Occurrence="923903286" collectingDate="1999-06-30" collectorName="Z. Kejval, M. Tryzna" country="India" latitude="8.766666" location="Ponmudi hill resort, 30 km NE of Trivandrum, ca 1300-1500 m" longitude="77.1" stateProvince="Kerala state" typeStatus="Holotype">
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‘
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<collectingCountry id="2F6F5658F10709DF59ACD51CFAE9FE86">S-INDIA</collectingCountry>
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,
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<collectingRegion id="A99A0AA4A7927FFF3E2DF0F31ADAE6A7">Kerala state</collectingRegion>
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,
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<location id="EEC2D9DE2F8BAF0306ECDB3B4B9CE416" LSID="urn:lsid:plazi:treatment:934972B786C31DD9FB7725AB7B082525:EEC2D9DE2F8BAF0306ECDB3B4B9CE416" country="S-INDIA" latitude="8.766666" longitude="77.1" name="Ponmudi hill resort, 30 km NE of Trivandrum" stateProvince="Kerala state">Ponmudi hill resort, 30 km NE of Trivandrum</location>
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,
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<geoCoordinate id="F66FCBAFBC47EB3BFF59FEE6021ABB69" direction="east" orientation="longitude" value="77.1">77°06'E</geoCoordinate>
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<geoCoordinate id="6A8A385A97741625D1255BE3E5FA2C81" direction="north" orientation="latitude" value="8.766666">8°46'N</geoCoordinate>
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,
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<location id="97A83BB951F936E9CDBF83A57CCBEAA9" LSID="urn:lsid:plazi:treatment:934972B786C31DD9FB7725AB7B082525:97A83BB951F936E9CDBF83A57CCBEAA9" country="S-INDIA" latitude="8.766666" longitude="77.1" name="ca 1300-1500m" stateProvince="Kerala state">ca 1300-1500m</location>
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,
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<collectingDate id="43F8298521AD6458CEF7FB673A9612B0">
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28-
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<date id="5DD4DBDA6D046968BFBAFD3818D3A814" day="30" month="06" value="1999-06-30" year="1999">30.vi.1999</date>
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</collectingDate>
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,
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<collectorName id="B2F7F509C58CCD3F1E05D0C914C1E222">Z. Kejval & M. Tryzna</collectorName>
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leg.’
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</materialsCitation>
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and red, printed, edged in black, label ‘HOLOTYPUS
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<taxonomicName id="21596DA4D26127BB06E734FF36255226" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="67" phylum="Arthropoda" rank="species" species="dux">Tyrannomyrmex dux</taxonomicName>
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des. M. L. Borowiec 2007’.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2AB07D5EB75EB7D9DB7934AFF6A2B99" type="diagnosis">
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<paragraph id="537C80F666C80E7EDF66F5C268243755" pageNumber="67">Diagnosis.</paragraph>
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<paragraph id="16F99A6DD87C3A2532F8F900C2FE3ACD" lastPageNumber="68" pageNumber="67">
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<taxonomicName id="385E52CDC6D16BDC482032C63003DC48" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="67" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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is most easily distinguished from
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<taxonomicName id="CB01E83D93052F3114FEC6637C504D20" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182903" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex rex Fernández" order="Hymenoptera" pageNumber="67" phylum="Arthropoda" rank="species" species="rex">T. rex</taxonomicName>
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by the differences in pilosity, sculpture and the shape of the petiole.
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<taxonomicName id="4A037D1A9817DA7804061FDE90294308" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182903" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex rex Fernández" order="Hymenoptera" pageNumber="67" phylum="Arthropoda" rank="species" species="rex">T. rex</taxonomicName>
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is almost lacking pilosity on the mesosoma, while its whole surface is covered with erect hairs in
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<taxonomicName id="78A977ECC06A76F0A3060EE4CC740950" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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. Also the sculpturation is stronger in
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<taxonomicName id="4451F78F13583D6A6454BA7CF892B332" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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, especially on the mesosoma. While in
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<taxonomicName id="5916F4ACFB99D079431B267BA1A5B824" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182903" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex rex Fernández" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="rex">T. rex</taxonomicName>
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foveae on the mesosoma are small with most interspaces equal or wider than their diameter,
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<taxonomicName id="5A0FE971479B9D265BD343C20C318329" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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has them deeper, larger, irregular and almost touching each other. Also, the petiole in
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<taxonomicName id="EE36188E24B70B1FC9D5C43A1926BE26" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182903" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex rex Fernández" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="rex">T. rex</taxonomicName>
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is stout with a big node and short peduncle, while in
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<taxonomicName id="12EFCC71964E65765B1B2B97BF516246" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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the node is more slender, much lower than long (Fig. 4).
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</paragraph>
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</subSubSection>
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<subSubSection id="653C27D129C5A4DEEB1DD5E1FA26D64F" type="discussion">
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<paragraph id="52896074E86B846A75D5ECB84EFAA40D" pageNumber="68">Discussion.</paragraph>
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<paragraph id="62267D3AC83F89D433EB846D499D8DB9" pageNumber="68">
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Fernandez (2003) listed the following character combination as unique to
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<taxonomicName id="085FF06E1C98EA4750FD59C991510C5F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182902" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex Fernández" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="genus">Tyrannomyrmex</taxonomicName>
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:
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</paragraph>
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<paragraph id="B43CA75DC7BC74AD0D0FE9AD77338055" pageNumber="68">1) Mandibles with two teeth in the masticatory border, apical and smaller subapical.</paragraph>
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<paragraph id="08E1682D2E49EA7D5A22DB661EF99CA9" pageNumber="68">2) Inner ventral margin of masticatory border of mandibles with modified thick, cylindrical and transparent setae.</paragraph>
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<paragraph id="739596F82D95701746EBF6D1A320D69A" pageNumber="68">3) Clypeus not modified as central plate and devoid of carinae.</paragraph>
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<paragraph id="8F6CBB3095E4FD97C7AD1F39D577E217" pageNumber="68">4) Anterior clypeal margin with single median seta.</paragraph>
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<paragraph id="07955E3FE986F936AE3B0C007D0E8439" pageNumber="68">5) Palpal formula apparently 2,2.</paragraph>
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<paragraph id="E94E1C7561E0642BE21DB808329244CA" pageNumber="68">6) Antennae 11-segmented with ill-defined 3-segmented club.</paragraph>
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<paragraph id="967501317B4C431229D5C0E43BFCC7CC" pageNumber="68">7) Compound eyes small, reduced to a few ommatidia.</paragraph>
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<paragraph id="C9C7875088E92AD3164E2EC808B7D804" pageNumber="68">8) Frontal carinae and antennal scrobes absent.</paragraph>
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<paragraph id="5DBDD6E7114A1B0EE16A5E422F1D6B8A" pageNumber="68">9) Mesosoma without grooves.</paragraph>
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<paragraph id="C70C1938F5040F1550B804B20C5B2B5D" pageNumber="68">10) Propodeal spiracle round.</paragraph>
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<paragraph id="73CEFBECA02BEE56ED4A888A67BE46FA" pageNumber="68">11) Propodeal lobes large.</paragraph>
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<paragraph id="8048BC9AC7ABC55E21B22650BA5B9CF7" pageNumber="68">12) Sting large and robust.</paragraph>
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<paragraph id="725E5587738570510F239B65FC2681A2" pageNumber="68">
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<taxonomicName id="EC9FA4AE56F51FEE2AB44F921760ACAA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">Tyrannomyrmex dux</taxonomicName>
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agrees well with this diagnosis, but the setae on the venter of the mandibles (2) appear simple, not modified, and similar in structure to other setae of body. Though the sting is also extruded in
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<taxonomicName id="89FFA612B7776216F3C73650637616BD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229335" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex dux Borowiec" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="dux">T. dux</taxonomicName>
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, it is not as long as in
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<taxonomicName id="532E1A4863B596CF5499E0E0C83DD03D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182903" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex rex Fernández" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="species" species="rex">T. rex</taxonomicName>
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(12). Palp formula (5) could not be determined reliably. Thus at least characters 2) and 12) are not universal throughout
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<taxonomicName id="7E9B324D7B51FADB7DEB7A236A9119E9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182902" class="Insecta" family="Formicidae" genus="Tyrannomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Tyrannomyrmex Fernández" order="Hymenoptera" pageNumber="68" phylum="Arthropoda" rank="genus">Tyrannomyrmex</taxonomicName>
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. Formal placement of the genus in any tribe should be postponed until examination of other castes and more worker material is available allowing dissections.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |