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<document id="4084900200CB6F8B0C2B764D420BD915" ID-DOI="10.1111/zoj.12362" ID-ISSN="0024-4082" ID-Zenodo-Dep="10109299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235092894" checkinUser="marcus" docAuthor="Dellapé, Pablo M., Melo, María C. &amp; Henry, Thomas J." docDate="2016" docId="69250127E9695002FEAC346EFB3FE82A" docLanguage="en" docName="zoj12362.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (1)" docSource="http://dx.doi.org/10.1111/zoj.12362" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus penai Dellapé &amp; Melo &amp; Henry 2016, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="103" masterDocId="951C795FE9235048FF9E3541FFCCEE5A" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="103" updateTime="1700019240479" updateUser="ExternalLinkService">
<mods:mods id="843E6376AF4B3EC0B0A2EF21785378DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="68E27D5CABA1492D8018CE8DC33D9C4F">
<mods:title id="B371D9419C1910F42EC1E517727786CE">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="69063DA01C6A3F250131556525EA9496">Dellapé, Pablo M.</mods:namePart>
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<mods:namePart id="355DB6B88DD7F753160108A2EBD8CCF0">Melo, María C.</mods:namePart>
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<mods:namePart id="C593FB21234823DB2B3831D52509DB0E">Henry, Thomas J.</mods:namePart>
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<mods:title id="DF4BAD4ED97870547F590AEA683EC84A">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="C2135E4C0EA5D005A29C875820E781C0">2016</mods:date>
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<mods:number id="2D6ED623068696BA684B1EEF38DDB1F2">2016-04-17</mods:number>
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<mods:detail id="286F5B2975D1C7F324CABB831C327CCC" type="volume">
<mods:number id="583ECF225A0F0AFB6545BE1D3CCA7FFF">177</mods:number>
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<mods:number id="95F1B4971A8889960339394FFEA92C12">1</mods:number>
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<mods:url id="F5C63657F2DB2531FD92EC760E98D166">http://dx.doi.org/10.1111/zoj.12362</mods:url>
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<mods:classification id="EA565DF91296D5EAD5E7B6C8C6947D63">journal article</mods:classification>
<mods:identifier id="E90220C6A639C8B10B32D644C69A882A" type="DOI">10.1111/zoj.12362</mods:identifier>
<mods:identifier id="36A7CD0813FBC19D76580EE613C47F52" type="ISSN">0024-4082</mods:identifier>
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<treatment id="69250127E9695002FEAC346EFB3FE82A" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A" httpUri="http://treatment.plazi.org/id/69250127E9695002FEAC346EFB3FE82A" lastPageNumber="103" pageId="74" pageNumber="103">
<subSubSection id="A996E3BAE9695002FEAC346EFD98EF1E" box="[306,596,303,324]" pageId="74" pageNumber="103" type="nomenclature">
<paragraph id="E133B031E9695002FEAC346EFD98EF1E" blockId="74.[306,596,303,324]" box="[306,596,303,324]" pageId="74" pageNumber="103">
<heading id="BA7B075DE9695002FEAC346EFD98EF1E" box="[306,596,303,324]" centered="true" fontSize="9" level="2" pageId="74" pageNumber="103" reason="4">
<taxonomicName id="268CCBB2E9695002FEAC346EFE3DEF19" ID-CoL="3KXMR" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[306,497,303,324]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="103" phylum="Arthropoda" rank="species" species="penai" status="sp. nov.">HERAEUS PENAI</taxonomicName>
<emphasis id="D3F86C23E9695002FE663473FD98EF1E" bold="true" box="[504,596,303,324]" pageId="74" pageNumber="103">
<taxonomicNameLabel id="C8CBD158E9695002FE663473FD98EF1E" box="[504,596,303,324]" pageId="74" pageNumber="103" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A996E3BAE9695002FF603414FB3FE82A" pageId="74" pageNumber="103" type="description">
<paragraph id="E133B031E9695002FF603414FD45EF31" blockId="74.[254,649,341,363]" box="[254,649,341,363]" pageId="74" pageNumber="103">
(
<figureCitation id="79B7ACB4E9695002FE983414FEADEF30" box="[262,353,341,362]" captionStart="Figure 29" captionStartId="53.[164,241,1163,1182]" captionTargetBox="[168,1440,199,1129]" captionTargetId="figure-224@53.[164,1444,195,1133]" captionTargetPageId="53" captionText="Figure 29. Distributional map of Heraeus apicalis sp. nov., Heraeus bahiensis sp. nov., Heraeus bolivianus sp. nov., and Heraeus penai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128637" httpUri="https://zenodo.org/record/10128637/files/figure.png" pageId="74" pageNumber="103">FIGS 29</figureCitation>
,
<figureCitation id="79B7ACB4E9695002FEF03414FE6EEF30" box="[366,418,341,362]" captionStart="Figure 30" captionStartId="55.[164,241,1815,1834]" captionTargetBox="[165,1444,193,1786]" captionTargetId="figure-0@55.[164,1444,193,1786]" captionTargetPageId="55" captionText="Figure 30. Aedeagus. A, Heraeus bahiensis sp. nov.; B, Heraeus bolivianus sp. nov.; C, Heraeus brevirostris sp. nov.; D, Heraeus concolor Slater &amp; Baranowski, 1994; E, Heraeus ecuatorianus sp. nov.; F, Heraeus mesoamericanus sp. nov.; G, Heraeus mexicanus sp. nov.; H, Heraeus penai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128639" httpUri="https://zenodo.org/record/10128639/files/figure.png" pageId="74" pageNumber="103">30H</figureCitation>
,
<figureCitation id="79B7ACB4E9695002FE313414FE2DEF31" box="[431,481,341,363]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="74" pageNumber="103">36G</figureCitation>
,
<figureCitation id="79B7ACB4E9695002FE703414FDECEF30" box="[494,544,341,362]" captionStart="Figure 39" captionStartId="70.[144,220,1059,1078]" captionTargetBox="[305,1261,196,1028]" captionTargetId="figure-296@70.[303,1263,194,1030]" captionTargetPageId="70" captionText="Figure 39. Legs: A, Heraeus nicaraguensis sp. nov.; B, Heraeus pacificus Barber, 1925; C, Heraeus pallidinervis sp. nov.; D, Heraeus penai sp. nov.; E, Heraeus plebejus Stål, 1874; F, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128659" httpUri="https://zenodo.org/record/10128659/files/figure.png" pageId="74" pageNumber="103">39D</figureCitation>
,
<figureCitation id="79B7ACB4E9695002FDB33414FD4DEF31" box="[557,641,341,363]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="74" pageNumber="103">40M–P</figureCitation>
)
</paragraph>
<paragraph id="E133B031E9695002FF0E343DFECCEFCB" blockId="74.[144,759,380,708]" box="[144,256,380,401]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FF0E343DFECCEFCB" box="[144,256,380,401]" italics="true" pageId="74" pageNumber="103">Diagnosis</emphasis>
</paragraph>
<paragraph id="E133B031E9695002FF0E34DBFEECEC56" blockId="74.[144,759,380,708]" pageId="74" pageNumber="103">Labium almost extending to metacoxae. Distiflagellomere with a narrow pale band sub-basally. Membrane brown, with veins paler. Metafemur with a narrow subapical brown band.</paragraph>
<paragraph id="E133B031E9695002FF363754FDABEC9E" blockId="74.[144,759,380,708]" pageId="74" pageNumber="103">
In addition to
<taxonomicName id="268CCBB2E9695002FE1E3754FDF2EC70" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[384,574,533,554]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="103" phylum="Arthropoda" rank="species" species="penai" status="sp. nov.">
<emphasis id="D3F86C23E9695002FE1E3754FDF2EC70" box="[384,574,533,554]" italics="true" pageId="74" pageNumber="103">Heraeus penai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E9695002FDD63754FD65EC70" box="[584,681,533,554]" pageId="74" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
, only
<taxonomicName id="268CCBB2E9695002FF0E3775FE85EC12" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[144,329,563,585]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="103" phylum="Arthropoda" rank="species" species="pallidinervis" status="sp. nov.">
<emphasis id="D3F86C23E9695002FF0E3775FE85EC12" box="[144,329,563,585]" italics="true" pageId="74" pageNumber="103">H. pallidinervis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E9695002FECE3775FE64EC13" box="[336,424,564,585]" pageId="74" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="268CCBB2E9695002FE753775FDF7EC12" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[491,571,563,585]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="74" pageNumber="103" phylum="Arthropoda" rank="species" species="loja" status="sp. nov.">
<emphasis id="D3F86C23E9695002FE753775FDF7EC12" box="[491,571,563,585]" italics="true" pageId="74" pageNumber="103">H. loja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E9695002FDDC3775FD55EC13" box="[578,665,564,585]" pageId="74" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
have a narrow pale band on distiflagellomere, occupying about one-quarter of segment length, and a long labium extending to metacoxae. The male genitalic characters, however, will differentiate these species.
</paragraph>
<paragraph id="E133B031E9695002FF0E37ADFE40ED5B" blockId="74.[144,759,748,1902]" box="[144,396,748,769]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FF0E37ADFE40ED5B" box="[144,396,748,769]" italics="true" pageId="74" pageNumber="103">
Description (
<figureCitation id="79B7ACB4E9695002FEBC37ADFE48ED5B" box="[290,388,748,769]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="74" pageNumber="103">Fig. 36G</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="E133B031E9695002FF0E364BFE91ED7A" blockId="74.[144,759,748,1902]" box="[144,349,778,800]" pageId="74" pageNumber="103">Total length 5.61.</paragraph>
<paragraph id="E133B031E9695002FF363668FE43EB53" blockId="74.[144,759,748,1902]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FF363668FF26ED64" box="[168,234,809,830]" italics="true" pageId="74" pageNumber="103">Head:</emphasis>
Brown, shiny, coriaceous, with abundant short recumbent and sparse long erect setae. Head length 1.22, width 0.84. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.43, interocellar width 0.26. Labium pale brown with erect setae, almost extending to metacoxae. Labial segment lengths: I 0.83, II 0.96, III 0.80, and IV 0.32. Antennae pale brown, except apex of scapus, pedicel, and basiflagellomere; distiflagellomere darker, with a narrow pale band subbasally; all segments with abundant short semi-erect and sparse longer erect setae. Antennal lengths: scape 0.46, pedicel 0.96, basiflagellomere 0.86, and distiflagellomere 0.98. Length of pale band on distiflagellomere 0.24.
</paragraph>
<paragraph id="E133B031E9695002FF363053FA42EF6C" blockId="74.[144,759,748,1902]" lastBlockId="74.[806,1422,197,677]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FF363053FF32EB7D" box="[168,254,1298,1319]" italics="true" pageId="74" pageNumber="103">Thorax:</emphasis>
Anterior pronotal lobe brown, collar and posterior lobe paler, with irregular paler areas, humeral angles with a whitish spot; collar delimited posteriorly by a punctuate sulcus; punctate, punctures larger on posterior lobe; anterior and posterior lobes with short recumbent and erect setae, longer erect setae on anterior lobe. Collar length 0.07, anterior lobe length 0.53, posterior lobe length 0.50; anterior lobe width 0.90, posterior lobe width 1.39. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, with a darker longitudinal median stripe, pruinose, punctate, with short recumbent and long erect setae, as for anterior pronotal lobe. Hemelytra brown, costal margin pale on proximal three-quarters, apical half of corium darker, with a subapical whitish spot and a small pale rounded spot internally (
<figureCitation id="79B7ACB4E9695002FEC033BDFE06E948" box="[350,458,1788,1810]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="74" pageNumber="103">Fig. 36G</figureCitation>
); setae short and semierect; membrane brown with pale areas, veins paler. Legs: Coxae, protrochanter, profemur, except distally, and a subapical narrow band on metafemur brown, re- mainder of legs pale brown; mesofemur with a narrow band subapically and apex of tibiae darker (
<figureCitation id="79B7ACB4E9695002FA8035A2FA4DEEA3" box="[1310,1409,227,249]" captionStart="Figure 39" captionStartId="70.[144,220,1059,1078]" captionTargetBox="[305,1261,196,1028]" captionTargetId="figure-296@70.[303,1263,194,1030]" captionTargetPageId="70" captionText="Figure 39. Legs: A, Heraeus nicaraguensis sp. nov.; B, Heraeus pacificus Barber, 1925; C, Heraeus pallidinervis sp. nov.; D, Heraeus penai sp. nov.; E, Heraeus plebejus Stål, 1874; F, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128659" httpUri="https://zenodo.org/record/10128659/files/figure.png" pageId="74" pageNumber="103">Fig. 39D</figureCitation>
); setae abundant and semi-erect, longest on profemur. Profemur with spines restricted to apical two-thirds.
</paragraph>
<paragraph id="E133B031E9695002FCA1347EFC4CECFF" blockId="74.[806,1422,197,677]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FCA1347EFC7CEF0E" box="[831,944,319,340]" italics="true" pageId="74" pageNumber="103">Abdomen:</emphasis>
Brown, with abundant short recumbent setae. Male genitalia: Pygophore (
<figureCitation id="79B7ACB4E9695002FB05341FFAD2EF29" box="[1179,1310,350,372]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="74" pageNumber="103">Fig. 40M, N</figureCitation>
) rounded, anterior margin of dorsal aperture slightly rounded, inner projections subquadrangular; declivent posteriorly in lateral view. Outer projection of parameres not developed (
<figureCitation id="79B7ACB4E9695002FC3B3499FBEBEFB7" box="[933,1063,472,494]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="74" pageNumber="103">Fig. 40O, P</figureCitation>
). Aedeagus (
<figureCitation id="79B7ACB4E9695002FB293499FAD0EFB4" box="[1207,1308,472,494]" captionStart="Figure 30" captionStartId="55.[164,241,1815,1834]" captionTargetBox="[165,1444,193,1786]" captionTargetId="figure-0@55.[164,1444,193,1786]" captionTargetPageId="55" captionText="Figure 30. Aedeagus. A, Heraeus bahiensis sp. nov.; B, Heraeus bolivianus sp. nov.; C, Heraeus brevirostris sp. nov.; D, Heraeus concolor Slater &amp; Baranowski, 1994; E, Heraeus ecuatorianus sp. nov.; F, Heraeus mesoamericanus sp. nov.; G, Heraeus mexicanus sp. nov.; H, Heraeus penai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128639" httpUri="https://zenodo.org/record/10128639/files/figure.png" pageId="74" pageNumber="103">Fig. 30H</figureCitation>
): conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with two pairs of lobes, the anterior lobe short, unsclerotized, and with many spines, and the posterior lobe larger and sclerotized, with a few spines distally;
<emphasis id="D3F86C23E9695002FBB73733FADEECDD" box="[1065,1298,626,647]" italics="true" pageId="74" pageNumber="103">processus gonopori</emphasis>
long and slender.
</paragraph>
<paragraph id="E133B031E9695002FCB8378EFC7DECBE" blockId="74.[806,1309,719,771]" box="[806,945,719,740]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FCB8378EFC7DECBE" box="[806,945,719,740]" italics="true" pageId="74" pageNumber="103">Distribution</emphasis>
</paragraph>
<paragraph id="E133B031E9695002FCB837ACFAD1ED59" blockId="74.[806,1309,719,771]" box="[806,1309,749,771]" pageId="74" pageNumber="103">
<collectingCountry id="999BF0A1E9695002FCB837ACFC50ED59" box="[806,924,749,771]" name="Argentina" pageId="74" pageNumber="103">Argentina</collectingCountry>
,
<collectingCountry id="999BF0A1E9695002FC3437ACFC31ED59" box="[938,1021,749,771]" name="Bolivia" pageId="74" pageNumber="103">Bolivia</collectingCountry>
, and
<collectingCountry id="999BF0A1E9695002FBDE37AFFB62ED59" box="[1088,1198,750,771]" name="Paraguay" pageId="74" pageNumber="103">Paraguay</collectingCountry>
(
<figureCitation id="79B7ACB4E9695002FB5E37ACFADDED59" box="[1216,1297,749,771]" captionStart="Figure 29" captionStartId="53.[164,241,1163,1182]" captionTargetBox="[168,1440,199,1129]" captionTargetId="figure-224@53.[164,1444,195,1133]" captionTargetPageId="53" captionText="Figure 29. Distributional map of Heraeus apicalis sp. nov., Heraeus bahiensis sp. nov., Heraeus bolivianus sp. nov., and Heraeus penai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128637" httpUri="https://zenodo.org/record/10128637/files/figure.png" pageId="74" pageNumber="103">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="E133B031E9695002FCB8366DFC51ED1B" blockId="74.[806,1421,812,896]" box="[806,925,812,833]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FCB8366DFC51ED1B" box="[806,925,812,833]" italics="true" pageId="74" pageNumber="103">Etymology</emphasis>
</paragraph>
<paragraph id="E133B031E9695002FCB8360AFAA5ED25" blockId="74.[806,1421,812,896]" pageId="74" pageNumber="103">
This species is named after the late Luiz E. Peña, who collected the
<typeStatus id="3E370E93E9695002FC5F362BFBEEEDDA" box="[961,1058,874,896]" pageId="74" pageNumber="103" type="holotype">holotype</typeStatus>
and some of the
<typeStatus id="3E370E93E9695002FB6F362BFAA9ED25" box="[1265,1381,874,895]" pageId="74" pageNumber="103" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="E133B031E9695002FCB836EBFC0FEDE4" blockId="74.[806,1421,937,1020]" box="[806,963,937,959]" pageId="74" pageNumber="103">
<emphasis id="D3F86C23E9695002FCB836EBFC0FEDE4" box="[806,963,937,959]" italics="true" pageId="74" pageNumber="103">Type material</emphasis>
</paragraph>
<paragraph id="E133B031E9695002FCB83686FAAEEDA6" blockId="74.[806,1421,937,1020]" pageId="74" pageNumber="103">
<materialsCitation id="51E4BA6CE9695002FCB83686FAAEEDA6" collectingDate="1984-12-17" collectionCode="USNM" collectorName="V. Grande &amp; L. E. Pena" country="Bolivia" elevation="1900" location="Mataral" pageId="74" pageNumber="103" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="D3F86C23E9695002FCB83686FC41ED86" box="[806,909,967,988]" italics="true" pageId="74" pageNumber="103">
<typeStatus id="3E370E93E9695002FCB83686FC4BED86" box="[806,903,967,988]" pageId="74" pageNumber="103" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E9695002FC0A3686FC60ED87" box="[916,940,967,989]" pageId="74" pageNumber="103" type="male">♂</specimenCount>
,
<emphasis id="D3F86C23E9695002FC253686FBE1ED87" bold="true" box="[955,1069,967,989]" pageId="74" pageNumber="103">
<collectingCountry id="999BF0A1E9695002FC253686FBE1ED87" box="[955,1069,967,989]" name="Bolivia" pageId="74" pageNumber="103">BOLIVIA</collectingCountry>
</emphasis>
,
<location id="E453E6EAE9695002FBA53686FB59ED87" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FBA53686FB59ED87" box="[1083,1173,967,989]" country="Bolivia" name="Mataral" pageId="74" pageNumber="103">Mataral</location>
(N),
<collectorName id="4C79D5E7E9695002FB533689FAF3ED87" box="[1229,1343,967,989]" pageId="74" pageNumber="103">V. Grande</collectorName>
,
<elevation id="6AA15702E9695002FAD43689FC7BEDA6" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="74" pageNumber="103" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<quantity id="26741DD4E9695002FAD43689FCB6EDA6" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="74" pageNumber="103" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800– 2000 m</quantity>
a.s.l.
</elevation>
, 15/
<date id="953296F1E9695002FC7636A7FBBFEDA6" box="[1000,1139,998,1020]" pageId="74" pageNumber="103" value="1984-12-17">
<collectingDate id="85766F19E9695002FC7636A7FBBFEDA6" box="[1000,1139,998,1020]" pageId="74" pageNumber="103" value="1984-12-17">17-XII-1984</collectingDate>
</date>
,
<collectorName id="4C79D5E7E9695002FB1F36A6FB3DEDA6" box="[1153,1265,999,1020]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
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).
</materialsCitation>
</paragraph>
<paragraph id="E133B031E9695002FCB83154FB3FE82A" blockId="74.[806,1422,1044,1648]" pageId="74" pageNumber="103">
<materialsCitation id="51E4BA6CE9695002FCB83154FA45EA70" box="[806,1417,1044,1066]" collectingDate="1984-12-17" collectionCode="USNM" collectorName="V. Grande &amp; L. E. Pena" country="Bolivia" elevation="1900" location="Mataral" pageId="74" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<emphasis id="D3F86C23E9695002FCB83154FC56EA70" box="[806,922,1045,1066]" italics="true" pageId="74" pageNumber="103">
<typeStatus id="3E370E93E9695002FCB83154FC58EA70" box="[806,916,1045,1066]" pageId="74" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E9695002FC3F3155FC0AEA70" box="[929,966,1044,1066]" pageId="74" pageNumber="103" type="male">2♂</specimenCount>
,
<specimenCount id="F78A7BB8E9695002FC4B3155FC3AEA70" box="[981,1014,1044,1066]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
, same data as for holotype (
<collectionCode id="879D28F4E9695002FAAF3155FA4EEA70" box="[1329,1410,1044,1066]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FCA13172FBDEEA3D" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" location="Chuquisaca" municipality="Monteagudo" pageId="74" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FCA13172FCAAEA12" box="[831,870,1074,1096]" pageId="74" pageNumber="103" type="male">2♂</specimenCount>
,
<specimenCount id="F78A7BB8E9695002FCE63172FC57EA12" box="[888,923,1074,1096]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<location id="E453E6EAE9695002FC303172FBF5EA13" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FC303172FBF5EA13" box="[942,1081,1075,1097]" country="Bolivia" municipality="Monteagudo" name="Chuquisaca" pageId="74" pageNumber="103">Chuquisaca</location>
,
<collectingMunicipality id="01572A4BE9695002FBD73172FB17EA13" box="[1097,1243,1075,1097]" pageId="74" pageNumber="103">Monteagudo</collectingMunicipality>
, 24-XII-[19]84,
<collectorName id="4C79D5E7E9695002FCA13113FC67EA3D" box="[831,939,1106,1127]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FC243110FBC7EA3D" box="[954,1035,1105,1127]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FB833110FC46EADC" collectionCode="MLP" collectorName="Pena" country="Bolivia" pageId="74" pageNumber="103" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FB833110FB8EEA3D" box="[1053,1090,1105,1127]" pageId="74" pageNumber="103" type="male">1♂</specimenCount>
,
<specimenCount id="F78A7BB8E9695002FBCF3110FBBEEA3D" box="[1105,1138,1105,1127]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
, 24-XII-[19]84, L.E.
<collectorName id="4C79D5E7E9695002FACB3113FA41EA3D" box="[1365,1421,1106,1127]" pageId="74" pageNumber="103">Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FCD83131FC4EEADC" box="[838,898,1136,1158]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="74" pageNumber="103">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FC093131FB1EEAFF" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" elevation="1600" location="Muyupampa" pageId="74" pageNumber="103" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FC093131FC72EADF" box="[919,958,1135,1157]" pageId="74" pageNumber="103" type="male">4♂</specimenCount>
,
<specimenCount id="F78A7BB8E9695002FC503131FC3CEADF" box="[974,1008,1135,1157]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
, (E)
<location id="E453E6EAE9695002FBB53131FB72EADF" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FBB53131FB72EADF" box="[1067,1214,1136,1157]" country="Bolivia" name="Muyupampa" pageId="74" pageNumber="103">Muyupampa</location>
,
<elevation id="6AA15702E9695002FB553131FA90EADC" box="[1227,1372,1136,1158]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="103" unit="m" value="1600.0">
<quantity id="26741DD4E9695002FB553131FAD3EADF" box="[1227,1311,1136,1158]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="103" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
, 21/ 25-XII-[19]84,
<collectorName id="4C79D5E7E9695002FC4131CEFB87EAFE" box="[991,1099,1167,1188]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
G. (
<collectionCode id="879D28F4E9695002FBE431CEFB07EAFF" box="[1146,1227,1167,1189]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<emphasis id="D3F86C23E9695002FB4331CEFAE6EAFE" box="[1245,1322,1167,1188]" italics="true" pageId="74" pageNumber="103">La Paz</emphasis>
:
<materialsCitation id="51E4BA6CE9695002FAA731CEFC69EAB8" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" elevation="1200" location="Rio Coroico" pageId="74" pageNumber="103" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FAA731CEFA96EAFE" box="[1337,1370,1166,1188]" pageId="74" pageNumber="103" type="female">3♀</specimenCount>
,
<location id="E453E6EAE9695002FAF631CFFC55EA99" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FAF631CFFC55EA99" country="Bolivia" name="Rio Coroico" pageId="74" pageNumber="103">Rio Coroico</location>
,
<elevation id="6AA15702E9695002FC3731ECFBF2EA99" box="[937,1086,1197,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="74" pageNumber="103" unit="m" value="1200.0">
<quantity id="26741DD4E9695002FC3731ECFC32EA98" box="[937,1022,1197,1218]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="74" pageNumber="103" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
, 24/26-XI-[19]84,
<collectorName id="4C79D5E7E9695002FA8431ECFA41EA98" box="[1306,1421,1197,1218]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FCD9318DFC51EAB8" box="[839,925,1228,1250]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FC2B318DFBB4EB5A" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" location="Tres Esteros" municipality="Guanay" pageId="74" pageNumber="103" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FC2B318DFC15EABB" box="[949,985,1227,1249]" pageId="74" pageNumber="103" type="female">7♀</specimenCount>
,
<location id="E453E6EAE9695002FC72318DFB45EABB" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FC72318DFB45EABB" box="[1004,1161,1228,1249]" country="Bolivia" municipality="Guanay" name="Tres Esteros" pageId="74" pageNumber="103">Tres Esteros</location>
,
<collectingMunicipality id="01572A4BE9695002FB04318DFB3BEABB" box="[1178,1271,1228,1249]" pageId="74" pageNumber="103">Guanay</collectingMunicipality>
, 19/25-VIII- [19]89,
<collectorName id="4C79D5E7E9695002FC0731AAFBC6EB5A" box="[921,1034,1259,1280]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FB8331ABFBBCEB5A" box="[1053,1136,1258,1280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FB1931ABFAF0EB45" collectingDate="1984-12-01" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" elevation="1700" location="Yungas" municipality="Coripata" pageId="74" pageNumber="103" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FB1931ABFB66EB5A" box="[1159,1194,1258,1280]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<location id="E453E6EAE9695002FB2531AAFADEEB5A" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FB2531AAFADEEB5A" box="[1211,1298,1259,1280]" country="Bolivia" municipality="Coripata" name="Yungas" pageId="74" pageNumber="103">Yungas</location>
,
<collectingMunicipality id="01572A4BE9695002FABC31ABFA45EB5A" box="[1314,1417,1258,1280]" pageId="74" pageNumber="103">Coripata</collectingMunicipality>
,
<elevation id="6AA15702E9695002FCA13048FC02EB45" box="[831,974,1289,1311]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="74" pageNumber="103" unit="m" value="1700.0">
<quantity id="26741DD4E9695002FCA13048FC5EEB44" box="[831,914,1289,1310]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="74" pageNumber="103" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
,
<date id="953296F1E9695002FC423048FB9BEB44" box="[988,1111,1289,1310]" pageId="74" pageNumber="103" value="1984-12-01">
<collectingDate id="85766F19E9695002FC423048FB9BEB44" box="[988,1111,1289,1310]" pageId="74" pageNumber="103" value="1984-12-01">1-XII-1984</collectingDate>
</date>
,
<collectorName id="4C79D5E7E9695002FBFA3048FB1FEB44" box="[1124,1235,1289,1310]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FB7C3048FAF8EB45" box="[1250,1332,1289,1311]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FAD63048FA45EB67" collectionCode="USNM" collectorName="N La Paz &amp; L. E. Pena" country="Bolivia" location="Mapiri" pageId="74" pageNumber="103" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FAD63048FAA6EB44" box="[1352,1386,1288,1310]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<collectorName id="4C79D5E7E9695002FAE73048FC42EB67" pageId="74" pageNumber="103">N La Paz</collectorName>
,
<location id="E453E6EAE9695002FC073066FC2AEB67" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FC073066FC2AEB67" box="[921,998,1319,1341]" country="Bolivia" name="Mapiri" pageId="74" pageNumber="103">Mapiri</location>
, 10/16-VIII-[19]89,
<collectorName id="4C79D5E7E9695002FB273069FAE8EB67" box="[1209,1316,1320,1341]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FAAC3066FA4EEB67" box="[1330,1410,1319,1341]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FCA13007FB51EB21" collectingDate="1984-12-26" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" elevation="1400" location="Yungas" municipality="Pte. Mururata" pageId="74" pageNumber="103" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FCA13007FCADEB01" box="[831,865,1349,1371]" pageId="74" pageNumber="103" type="female">2♀</specimenCount>
,
<location id="E453E6EAE9695002FCF13007FC09EB01" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FCF13007FC09EB01" box="[879,965,1350,1371]" country="Bolivia" municipality="Pte. Mururata" name="Yungas" pageId="74" pageNumber="103">Yungas</location>
,
<collectingMunicipality id="01572A4BE9695002FC4C3007FBB7EB01" box="[978,1147,1350,1371]" pageId="74" pageNumber="103">Pte. Mururata</collectingMunicipality>
,
<elevation id="6AA15702E9695002FB193007FA91EB06" box="[1159,1373,1350,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="74" pageNumber="103" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">
<quantity id="26741DD4E9695002FB193007FAD3EB01" box="[1159,1311,1350,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="74" pageNumber="103" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">1200–1600 m</quantity>
a.s.l.
</elevation>
, 24/
<date id="953296F1E9695002FCA13025FC0EEB20" box="[831,962,1380,1402]" pageId="74" pageNumber="103" value="1984-12-26">
<collectingDate id="85766F19E9695002FCA13025FC0EEB20" box="[831,962,1380,1402]" pageId="74" pageNumber="103" value="1984-12-26">26-XII-1984</collectingDate>
</date>
,
<collectorName id="4C79D5E7E9695002FC533024FBF5EB20" box="[973,1081,1381,1402]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FBD83024FB5AEB21" box="[1094,1174,1381,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<emphasis id="D3F86C23E9695002FB363024FAEAEB20" box="[1192,1318,1381,1402]" italics="true" pageId="74" pageNumber="103">Santa Cruz</emphasis>
:
<materialsCitation id="51E4BA6CE9695002FAAD3024FA98EBC3" collectionCode="USNM" collectorName="L. E. Pena" country="Bolivia" location="Pto. Camacho" pageId="74" pageNumber="103" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FAAD3024FA98EB20" box="[1331,1364,1380,1402]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<location id="E453E6EAE9695002FAFC3024FC6AEBC3" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FAFC3024FC6AEBC3" country="Bolivia" name="Pto. Camacho" pageId="74" pageNumber="103">Pto. Camacho</location>
(S), 20-XII-[19]84,
<collectorName id="4C79D5E7E9695002FBE330C5FB27EBC3" box="[1149,1259,1412,1433]" pageId="74" pageNumber="103">L.E. Peña</collectorName>
(
<collectionCode id="879D28F4E9695002FB6530C2FA81EBC3" box="[1275,1357,1411,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<emphasis id="D3F86C23E9695002FAC130C2FC09EBED" bold="true" pageId="74" pageNumber="103">AR- GENTINA:</emphasis>
<emphasis id="D3F86C23E9695002FC5030E3FBC0EBED" box="[974,1036,1442,1463]" italics="true" pageId="74" pageNumber="103">Salta</emphasis>
:
<materialsCitation id="51E4BA6CE9695002FB8230E3FB42EB8C" collectionCode="IFML" collectorName="R. Golbach" country="Algeria" location="Abra Grande" pageId="74" pageNumber="103" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FB8230E3FB8FEBED" box="[1052,1091,1441,1463]" pageId="74" pageNumber="103" type="male">1♂</specimenCount>
,
<collectingCountry id="999BF0A1E9695002FBCC30E3FB43EBE2" box="[1106,1167,1442,1464]" name="Algeria" pageId="74" pageNumber="103">Orán</collectingCountry>
,
<location id="E453E6EAE9695002FB0230E3FAF9EBE2" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FB0230E3FAF9EBE2" box="[1180,1333,1442,1464]" country="Algeria" name="Abra Grande" pageId="74" pageNumber="103">Abra Grande</location>
, 10-I/1- III-[19]67,
<collectorName id="4C79D5E7E9695002FC273080FBFEEB8C" box="[953,1074,1472,1494]" pageId="74" pageNumber="103">R. Golbach</collectorName>
(
<collectionCode id="879D28F4E9695002FBDC3081FB4BEB8C" box="[1090,1159,1472,1494]" pageId="74" pageNumber="103">IFML</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FB043081FB67EBAF" collectingDate="1971-07-12" collectionCode="IFML" collectorName="Porter-Stange" country="Algeria" location="Oran" pageId="74" pageNumber="103" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FB043081FB70EB8C" box="[1178,1212,1472,1494]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<collectingCountry id="999BF0A1E9695002FB553081FACBEB8C" box="[1227,1287,1472,1494]" name="Algeria" pageId="74" pageNumber="103">Orán</collectingCountry>
, 5 R.N.,
<date id="953296F1E9695002FAF43081FC69EBAE" pageId="74" pageNumber="103" value="1971-07-12">
<collectingDate id="85766F19E9695002FAF43081FC69EBAE" pageId="74" pageNumber="103" value="1971-07-12">12- VII-1971</collectingDate>
</date>
,
<collectorName id="4C79D5E7E9695002FC2F309EFB83EBAE" box="[945,1103,1503,1524]" pageId="74" pageNumber="103">Porter-Stange</collectorName>
(
<collectionCode id="879D28F4E9695002FBC0309EFB6FEBAF" box="[1118,1187,1503,1525]" pageId="74" pageNumber="103">IFML</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FB28309EFB35E84E" collectionCode="IADIZA" collectorName="Roig" country="Algeria" location="Oran" pageId="74" pageNumber="103" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FB28309EFB14EBAE" box="[1206,1240,1502,1524]" pageId="74" pageNumber="103" type="female">1♀</specimenCount>
,
<quantity id="26741DD4E9695002FB79309EFAE1EBAF" box="[1255,1325,1503,1525]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="74" pageNumber="103" unit="km" value="15.0">15 km</quantity>
S
<collectingCountry id="999BF0A1E9695002FAD2309EFA45EBAF" box="[1356,1417,1503,1525]" name="Algeria" pageId="74" pageNumber="103">Orán</collectingCountry>
, 13-VI-[19]95,
<collectingCountry id="999BF0A1E9695002FC4130BCFBEAE849" box="[991,1062,1533,1555]" name="Indonesia" pageId="74" pageNumber="103">Flores</collectingCountry>
&amp;
<collectorName id="4C79D5E7E9695002FBD130BCFB4EE849" box="[1103,1154,1533,1555]" pageId="74" pageNumber="103">Roig</collectorName>
(
<collectionCode id="879D28F4E9695002FB0D30BFFB3EE84E" box="[1171,1266,1534,1556]" pageId="74" pageNumber="103">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE9695002FA9930BFFA4AE849" box="[1287,1414,1534,1555]" country="Algeria" location="Catamarca" pageId="74" pageNumber="103" specimenCount="1" typeStatus="paratype">
<emphasis id="D3F86C23E9695002FA9930BFFA4AE849" box="[1287,1414,1534,1555]" italics="true" pageId="74" pageNumber="103">
<location id="E453E6EAE9695002FA9930BFFA4AE849" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FA9930BFFA4AE849" box="[1287,1414,1534,1555]" country="Algeria" name="Catamarca" pageId="74" pageNumber="103">Catamarca</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="51E4BA6CE9695002FCA1335DFAF0E80B" collectionCode="IADIZA" collectorName="El Arenal &amp; S. Roig" country="Algeria" elevation="1106" latitude="-29.080265" longLatPrecision="1" longitude="-65.51442" pageId="74" pageNumber="103" specimenCount="6" specimenCount-female="2" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="F78A7BB8E9695002FCA1335DFCAAE868" box="[831,870,1564,1586]" pageId="74" pageNumber="103" type="male">4♂</specimenCount>
,
<specimenCount id="F78A7BB8E9695002FCE4335DFC51E868" box="[890,925,1564,1586]" pageId="74" pageNumber="103" type="female">2♀</specimenCount>
,
<collectorName id="4C79D5E7E9695002FC2F335DFBE7E868" box="[945,1067,1564,1586]" pageId="74" pageNumber="103">El Arenal</collectorName>
,
<geoCoordinate id="84B8D6F6E9695002FBA2335DFB1AE86B" box="[1084,1238,1564,1586]" degrees="29" direction="south" minutes="4" orientation="latitude" pageId="74" pageNumber="103" precision="1" seconds="48.95" value="-29.080265">29°4′48.95″S</geoCoordinate>
,
<geoCoordinate id="84B8D6F6E9695002FB76335DFA45E868" box="[1256,1417,1564,1586]" degrees="65" direction="west" minutes="30" orientation="longitude" pageId="74" pageNumber="103" precision="1" seconds="51.9" value="-65.51442">65°30′51.9″W</geoCoordinate>
,
<elevation id="6AA15702E9695002FCA1337AFC1EE80B" box="[831,978,1595,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.1059999999999999" pageId="74" pageNumber="103" unit="m" value="1106.0">
<quantity id="26741DD4E9695002FCA1337AFC5EE80A" box="[831,914,1595,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.1059999999999999" pageId="74" pageNumber="103" unit="m" value="1106.0">1106 m</quantity>
a.s.l.
</elevation>
, XII-[20]03,
<collectorName id="4C79D5E7E9695002FBF0337AFB0CE80B" box="[1134,1216,1595,1617]" pageId="74" pageNumber="103">S. Roig</collectorName>
(
<collectionCode id="879D28F4E9695002FB4A337AFAFAE80B" box="[1236,1334,1595,1617]" pageId="74" pageNumber="103">IADIZA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="51E4BA6CE9695002FAD2337AFB23E82A" collectingDate="1993-11" collectionCode="USNM" country="Paraguay" location="Capiata" pageId="74" pageNumber="103" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="D3F86C23E9695002FAD2337AFCBAE835" italics="true" pageId="74" pageNumber="103">
<collectingCountry id="999BF0A1E9695002FAD2337AFCBAE835" name="Paraguay" pageId="74" pageNumber="103">Paraguay</collectingCountry>
</emphasis>
:
<specimenCount id="F78A7BB8E9695002FC18331BFC61E835" box="[902,941,1625,1647]" pageId="74" pageNumber="103" type="male">1♂</specimenCount>
,
<location id="E453E6EAE9695002FC233318FBD6E835" LSID="urn:lsid:plazi:treatment:69250127E9695002FEAC346EFB3FE82A:E453E6EAE9695002FC233318FBD6E835" box="[957,1050,1625,1647]" country="Paraguay" name="Capiata" pageId="74" pageNumber="103">Capiata</location>
,
<date id="953296F1E9695002FBB6331BFB4EE835" box="[1064,1154,1626,1647]" pageId="74" pageNumber="103" value="1993-11">
<collectingDate id="85766F19E9695002FBB6331BFB4EE835" box="[1064,1154,1626,1647]" pageId="74" pageNumber="103" value="1993-11">XI-1993</collectingDate>
</date>
(
<collectionCode id="879D28F4E9695002FB0A331BFB24E82A" box="[1172,1256,1626,1648]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="103" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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