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<document id="BB15C2DA76E928F61932E9C97B64613C" ID-CLB-Dataset="46282" ID-DOI="10.11646/zootaxa.3774.3.4" ID-GBIF-Dataset="811f2551-c599-4267-89c7-f3c71f8b8016" ID-ISSN="1175-5326" ID-Zenodo-Dep="230186" ID-ZooBank="1DEC9380-989D-4C21-AE36-8DCF03E0CBD2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393019391" checkinUser="plazi" docAuthor="Joharchi, Omid, Mašán, Peter &amp; Babaeian, Esmaeil" docDate="2014" docId="03F787F4FFEDB611FF28FB88FA3868C5" docLanguage="en" docName="zt03774p281.pdf" docOrigin="Zootaxa 3774 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eviphididae Joharchi, Mašán &amp; Babaeian, 2014, gen. nov." docType="key" docVersion="8" lastPageNumber="280" masterDocId="FFCEFF8CFFEEB614FFBFFF94FFA46917" masterDocTitle="A new genus and species of edaphic mite (Acari: Mesostigmata: Eviphididae) from Iran" masterLastPageNumber="281" masterPageNumber="275" pageNumber="278" updateTime="1698298440975" updateUser="ExternalLinkService">
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<mods:title id="692D34DFDE4D11C2A22DE5A31FD4D4D4">A new genus and species of edaphic mite (Acari: Mesostigmata: Eviphididae) from Iran</mods:title>
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<mods:namePart id="84B4D64FE7562F4B05EFE82104B0FC6F">Joharchi, Omid</mods:namePart>
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<mods:namePart id="F21EF0CC8021818A352FF9BD0C980455">Mašán, Peter</mods:namePart>
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<mods:title id="4380B612AE14A90B3EC1F7F853158DC4">Zootaxa</mods:title>
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<mods:date id="805DF19CE16DA9F79626EC3E9A5141F5">2014</mods:date>
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<treatment id="03F787F4FFEDB611FF28FB88FA3868C5" ID-DOI="http://doi.org/10.5281/zenodo.6124533" ID-GBIF-Taxon="119417263" ID-Zenodo-Dep="6124533" LSID="urn:lsid:plazi:treatment:03F787F4FFEDB611FF28FB88FA3868C5" httpUri="http://treatment.plazi.org/id/03F787F4FFEDB611FF28FB88FA3868C5" lastPageId="5" lastPageNumber="280" pageId="3" pageNumber="278">
<subSubSection id="C3446569FFEDB617FF28FB88FB996D88" pageId="3" pageNumber="278" type="nomenclature">
<paragraph id="8BE136E2FFEDB617FF28FB88FC256D21" blockId="3.[151,897,1052,1078]" box="[151,897,1052,1078]" pageId="3" pageNumber="278">
<heading id="D0A9818EFFEDB617FF28FB88FC256D21" bold="true" box="[151,897,1052,1078]" fontSize="11" level="1" pageId="3" pageNumber="278" reason="1">
<emphasis id="B92AEAF0FFEDB617FF28FB88FC256D21" bold="true" box="[151,897,1052,1078]" pageId="3" pageNumber="278">
Key to the world genera of the family
<taxonomicName id="4C5E4D61FFEDB617FDCFFB88FCA36D21" ID-CoL="6242M" ID-ENA="704094" box="[624,775,1052,1078]" class="Arachnida" family="Eviphididae" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="278" phylum="Arthropoda" rank="family">Eviphididae</taxonomicName>
(females)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28FBF6FB996D88" blockId="3.[151,1436,1122,1183]" pageId="3" pageNumber="278">
The key to the European genera of the family, presented by
<bibRefCitation id="EFCF4B13FFEDB617FCCCFBF7FB036D6C" author="Masan" box="[883,1191,1122,1147]" pageId="3" pageNumber="278" refString="Masan, P. &amp; Halliday, R. B. (2010) Review of the European genera of Eviphididae (Acari: Mesostigmata) and the species occurring in Slovakia. Zootaxa, 2585, 1 - 122." type="journal article" year="2010">Mašán &amp; Halliday (2010)</bibRefCitation>
, is modified here to accommodate non-European genera, including the new genus
<taxonomicName id="4C5E4D61FFEDB617FCF0FB12FC696D89" box="[847,973,1158,1182]" class="Arachnida" family="Eviphididae" genus="Pedoniphis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92AEAF0FFEDB617FCF0FB12FC696D89" box="[847,973,1158,1182]" italics="true" pageId="3" pageNumber="278">Pedoniphis</emphasis>
</taxonomicName>
<emphasis id="B92AEAF0FFEDB617FC6BFB12FB996D88" bold="true" box="[980,1085,1158,1183]" pageId="3" pageNumber="278">
<taxonomicNameLabel id="A219578BFFEDB617FC6BFB12FB996D88" box="[980,1085,1158,1183]" pageId="3" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C3446569FFEDB611FF28FB59FA3868C5" lastPageId="5" lastPageNumber="280" pageId="3" pageNumber="278" type="key">
<keyStep id="30AA2E47FFEDB617FF28FB59FA3F6C2C" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28FB59FA3F6C16" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28FB59FA3F6C16" pageId="3" pageNumber="278">
1. Palpgenu with five setae; dorsal shield with 11–17 pairs of setae; sternal shield absent, reduced or fragmented; at least one pair of sternal setae inserted on soft integument..............................................
<taxonomicName id="4C5E4D61FFEDB617FB16FB78FA3F6C16" authority="Halbert, 1920" authorityName="Halbert" authorityYear="1920" box="[1193,1435,1259,1281]" class="Arachnida" family="Eviphididae" genus="Thinoseius" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEDB617FB16FB78FAB56C16" box="[1193,1297,1260,1281]" italics="true" pageId="3" pageNumber="278">Thinoseius</emphasis>
Halbert, 1920
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28FA9DFA3F6C2C" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28FA9DFA3F6C2C" pageId="3" pageNumber="278">- Palpgenu with six setae; dorsal shield with 22–33 pairs of setae (if present); sternal shield entire, with three pairs of sternal setae............................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEDB617FF28FAD7FA3F6CFC" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28FAD7FA3F6C84" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28FAD7FA3F6C84" pageId="3" pageNumber="278">
2. Dorsal shield expanded laterally, covering wide lateral strips of ventral idiosoma, and capturing three pairs of ventral setae; opisthogastric soft integument with three pairs of setae; central projection of epistome trifurcate...............................................................................................
<taxonomicName id="4C5E4D61FFEDB617FB87FAEAFA3F6C84" authority="Skorupski &amp; Blaszak, 1997" authorityName="Skorupski &amp; Blaszak" authorityYear="1997" box="[1080,1435,1405,1427]" class="Arachnida" family="Eviphididae" genus="Rafaphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEDB617FB87FAEAFB296C84" box="[1080,1165,1406,1427]" italics="true" pageId="3" pageNumber="278">Rafaphis</emphasis>
Skorupski &amp; Błaszak, 1997
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28FA08FA3F6CFC" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28FA08FA3F6CFC" pageId="3" pageNumber="278">- Dorsal shield never expanded onto ventrolateral margins of idiosoma; all ventral setae inserted on soft or sclerotised integument; opisthogastric integument with more than three pairs of setae; central projection of epistome undivided or absent.................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEDB617FF28FA67FA3F6F8C" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28FA67FA386F54" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28FA67FA386F54" pageId="3" pageNumber="278">3. Dorsal, endopodal, peritrematal and anal shields weakly outlined, not observable and apparently absent due to very light and even sclerotisation of idiosomal integument; anterior margin of epistome with several large spines on each side of central projection; pulvillus of legs II–IV with elongated and pointed lateral lobes, projecting well beyond claws................ 4</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28F9D8FA3F6F8C" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F9D8FA3F6F8C" pageId="3" pageNumber="278">- Idiosomal integument well sclerotised, so dorsal and ventral shields clearly defined; anterior margin of epistome with small denticles or at most one spine on each side of central projection, or subtriangular and without central projection; pulvillus of legs II–IV with shorter and regularly rounded lateral lobes..................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEDB617FF28F937FA386E5C" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28F937FA386FE4" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F937FA386FE4" pageId="3" pageNumber="278">
4. Sternal setae st1 and st2 well separated and inserted in flat sternal integument; metasternal setae st4 on small oval platelets; genital setae st5 on epigynal shield; genu I and tibia I with three ventral setae, tibia II and genu II with one anterolateral seta..................................................................................
<taxonomicName id="4C5E4D61FFEDB617FB10F94AFA386FE4" authority="Karg, 1963" authorityName="Karg" authorityYear="1963" box="[1199,1436,1757,1779]" class="Arachnida" family="Eviphididae" genus="Crassicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEDB617FB10F94AFA8E6FE4" box="[1199,1322,1758,1779]" italics="true" pageId="3" pageNumber="278">Crassicheles</emphasis>
Karg, 1963
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28F968FA386E5C" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F968FA386E5C" pageId="3" pageNumber="278">
- Sternal setae st1 and st2 adjacent and inserted in massive protuberance between coxae II; metasternal setae st4 on soft integument; genital setae st5 outside epigynal shield; genu I and tibia I with two ventral setae, tibia II and genu II with two anterolateral setae.....................................................
<emphasis id="B92AEAF0FFEDB617FBB2F8A2FB0D6E5C" box="[1037,1193,1846,1867]" italics="true" pageId="3" pageNumber="278">Neocrassicheles</emphasis>
<bibRefCitation id="EFCF4B13FFEDB617FB0FF8A1FA386E5C" author="Masan" box="[1200,1436,1845,1867]" pageId="3" pageNumber="278" refString="Masan, P. &amp; Halliday, R. B. (2010) Review of the European genera of Eviphididae (Acari: Mesostigmata) and the species occurring in Slovakia. Zootaxa, 2585, 1 - 122." type="journal article" year="2010">Mašán &amp; Halliday, 2010</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEDB617FF28F8C7FA3F6ED6" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28F8C7FA3F6E91" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F8C7FA3F6E91" pageId="3" pageNumber="278">5. Peritrematal shields reduced and lacking post-stigmatic extensions; poststigmatic gland pores gp2 on soft cuticle; peritremes relatively wider; dorsal shield hypotrichous and never with standard set of 30 pairs of setae.......................... 6</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEDB617FF28F81AFA3F6ED6" blockId="3.[151,1436,1229,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F81AFA3F6ED6" pageId="3" pageNumber="278">- Peritrematal shields well developed, with poststigmatic extensions bearing poststigmatic gland pores gp2; peritremes relatively narrower; dorsal shield with standard set of 30 pairs of setae................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEDB610FF28F85CFA3F684A" lastPageId="4" lastPageNumber="279" pageId="3" pageNumber="278">
<paragraph id="8BE136E2FFEDB617FF28F85CFA386EC9" blockId="3.[151,1436,1229,2014]" box="[151,1436,1992,2014]" pageId="3" pageNumber="278">
<keyLead id="30AF95D7FFEDB617FF28F85CFA386EC9" box="[151,1436,1992,2014]" pageId="3" pageNumber="278">6. Epistome subtriangular, without lanceolate central projection on anterior margin; dorsal shield with 22–26 pairs of setae;</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF5EFF03FA3F684A" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
anterolateral margins of dorsal shield deeply incised to form band-shaped scutal elements bearing three pairs of podonotal setae (s2, r2 and r3); ventral setae JV3 and JV4 absent; exopodal platelets absent; seta av1 on tarsi II −IV modified, enlarged, thickened and tri-dentate at the tip; femora III, IV and genu IV with two ventral setae............................... 7
<keyLead id="30AF95D7FFEAB610FF28FF7BFA3F684A" pageId="4" pageNumber="279">- Epistome with lanceolate central projection; dorsal shield with 27–29 pairs of setae; anterolateral incisions of dorsal shield absent or only inconspicuously developed, with at most short laterodorsal scutal elements bearing one pair of podonotal setae (s2); ventral setae JV3 and JV4 present; exopodal platelets present; seta av1 on tarsi II −IV normal, setiform; femora III, IV and genu IV with one ventral seta............................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FEF1FA3868F8" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FEF1FA386880" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FEF1FA386880" pageId="4" pageNumber="279">
7. Dorsal shield with 26 pairs of setae (J1, J3, Z2 and Z5 present); small metasternal platelets present, each with seta st4 and lyrifissure
<date id="FFE01022FFEAB610FE98FE15FEEC6880" box="[295,328,385,407]" pageId="4" pageNumber="279">iv3</date>
; deutosternal groove with six rows of denticles....................
<taxonomicName id="4C5E4D61FFEAB610FBB6FE16FA386880" authority="Kazemi &amp; Moraza, 2008" authorityName="Kazemi &amp; Moraza" authorityYear="2008" box="[1033,1436,385,407]" class="Arachnida" family="Eviphididae" genus="Metacryptoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FBB6FE16FB036880" box="[1033,1191,386,407]" italics="true" pageId="4" pageNumber="279">Metacryptoseius</emphasis>
<bibRefCitation id="EFCF4B13FFEAB610FB12FE15FA386880" author="Kazemi" box="[1197,1436,385,407]" pageId="4" pageNumber="279" refString="Kazemi, S., Moraza, M. L., Kamali, K. &amp; Saboori, A. (2008) A new genus and three new species of Eviphididae (Acari: Mesostigmata) associated with scarab beetles in Iran. Zootaxa, 1852, 1 - 20." type="journal article" year="2008">Kazemi &amp; Moraza, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FE0BFA3868F8" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FE0BFA3868F8" pageId="4" pageNumber="279">
- Dorsal shield with 22−23 pairs of setae (J3, Z2 and Z5 absent; J1 sometimes absent); metasternal platelets absent, setae st4 on soft integument and lyrifissures
<date id="FFE01022FFEAB610FDA9FE28FD9168C5" box="[534,565,444,466]" pageId="4" pageNumber="279">iv3</date>
on posterior margin of sternal shield; deutosternal groove with eight rows of denticles.................................................................................
<taxonomicName id="4C5E4D61FFEAB610FB39FE4EFA3868F8" authority="Makarova, 1998" authorityName="Makarova" authorityYear="1998" box="[1158,1436,473,495]" class="Arachnida" family="Eviphididae" genus="Cryptoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB39FE4EFB5D68F8" box="[1158,1273,474,495]" italics="true" pageId="4" pageNumber="279">Cryptoseius</emphasis>
<bibRefCitation id="EFCF4B13FFEAB610FB40FE4DFA3868F8" author="Makarova" box="[1279,1436,473,495]" pageId="4" pageNumber="279" refString="Makarova, O. L. (1998) A new eviphidid mite genus (Parasitiformes: Mesostigmata: Eviphididae) associated with the dung beetle Scarabaeus transcaspius Stolfa (Coleoptera: Scarabaeidae) in Turkmenistan. Acarologia, 39, 115 - 122." type="journal article" year="1998">Makarova, 1998</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FE63FA386B88" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FE63FA386B5F" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FE63FA386B5F" pageId="4" pageNumber="279">
8. Dorsal shield with 29 pairs of setae (J2 absent); peritremes long, with anterior tip reaching dorsolateral surface and well beyond insertion of podonotal setae s2; podonotal shield completely covering anterior dorsal surface and bearing all podonotal setae; femur II with ventral seta
<date id="FFE01022FFEAB610FDA8FDA5FD8B6B50" box="[535,559,561,583]" pageId="4" pageNumber="279">v1</date>
modified into a robust spine................................
<emphasis id="B92AEAF0FFEAB610FB72FDA6FA386B5F" bold="true" box="[1229,1436,562,584]" pageId="4" pageNumber="279">
<taxonomicName id="4C5E4D61FFEAB610FB72FDA6FA986B5F" box="[1229,1340,562,584]" class="Arachnida" family="Eviphididae" genus="Pedoniphis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92AEAF0FFEAB610FB72FDA6FA986B5F" bold="true" box="[1229,1340,562,584]" italics="true" pageId="4" pageNumber="279">Pedoniphis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A219578BFFEAB610FAFDFDA6FA386B5F" box="[1346,1436,562,584]" pageId="4" pageNumber="279" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FDDBFA386B88" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FDDBFA386B88" pageId="4" pageNumber="279">
- Dorsal shield with 27 pairs of setae (J1 and S1 absent, s6 outside shield); peritremes reduced in length, with anterior tip not exceeding anterior margin of coxae II; podonotal shield narrowed posteriorly, not completely covering anterior dorsal surface (setae s6 on soft integument); femur II with all setae uniform, needle-like.......................
<taxonomicName id="4C5E4D61FFEAB610FB7DFD1EFA386B88" authority="Karg, 1976" authorityName="Karg" authorityYear="1976" box="[1218,1436,649,671]" class="Arachnida" family="Eviphididae" genus="Scamaphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB7DFD1EFA8E6B88" box="[1218,1322,650,671]" italics="true" pageId="4" pageNumber="279">Scamaphis</emphasis>
<bibRefCitation id="EFCF4B13FFEAB610FA8FFD1EFA386B88" author="Karg" box="[1328,1436,649,671]" pageId="4" pageNumber="279" refString="Karg, W. (1976) Eine neue Milbengattung der Eviphididae Berlese, 1913 (Acarina, Parasitiformes). Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 50, 1 - 6." type="journal article" year="1976">Karg, 1976</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FD33FA3F6BE0" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FD33FA3F6BCD" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FD33FA3F6BCD" pageId="4" pageNumber="279">9. Post-stigmatic section of peritrematal shields expanded and elongated, with tip reaching beyond level of posterior margin of epigynal shield...................................................................................... 10</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FD76FA3F6BE0" blockId="4.[151,1436,151,2021]" box="[151,1435,737,759]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FD76FA3F6BE0" box="[151,1435,737,759]" pageId="4" pageNumber="279">- Post-stigmatic section of peritrematal shields short, reaching at most as far as posterior margin of coxae IV............. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FD6BFA3F6AB0" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FD6BFA386A58" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FD6BFA386A58" pageId="4" pageNumber="279">
10. Coxae II–IV on left side close to those on right side; sternal and epigynal shields anteriorly constricted; strips of soft integument between epigynal, peritrematal and anal shields very narrow; exopodal platelets II and III well developed; metapodal platelets and exopodals IV usually closely adjacent..................................
<taxonomicName id="4C5E4D61FFEAB610FB6BFCAEFA386A58" authority="Karg, 1963" authorityName="Karg" authorityYear="1963" box="[1236,1436,825,847]" class="Arachnida" family="Eviphididae" genus="Evimirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB6BFCAEFA8E6A58" box="[1236,1322,826,847]" italics="true" pageId="4" pageNumber="279">Evimirus</emphasis>
Karg, 1963
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FCC3FA3F6AB0" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FCC3FA3F6AB0" pageId="4" pageNumber="279">- Coxae II–IV on left side well separated from those on right side; sternal and epigynal shields not constricted; epigynal, peritrematal and anal shields separated by wide strips of integument; exopodal platelets II and III absent; metapodals and exopodals IV well separated........................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FC3BFA386D40" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FC3BFA326AE8" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FC3BFA326AE8" pageId="4" pageNumber="279">
11. Idiosoma highly domed, almost hemispherical; vertical setae j1 minute and needle-like; posterior ends of peritrematal shields greatly enlarged; metapodal platelets completely or partly fused with peritrematal shields, occasionally unfused but very narrowly separated from it; trochanter I with five setae, genu III with seven setae..................
<taxonomicName id="4C5E4D61FFEAB610FB7CFC7EFA326AE8" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1219,1430,1001,1023]" class="Arachnida" family="Eviphididae" genus="Eviphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB7CFC7EFAAF6AE8" box="[1219,1291,1002,1023]" italics="true" pageId="4" pageNumber="279">Eviphis</emphasis>
Berlese, 1903
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FB93FA386D40" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FB93FA386D40" pageId="4" pageNumber="279">
- Idiosoma dorsoventrally flattened; vertical setae j1 stout and lance-like; posterior ends of peritrematal shields less enlarged; metapodal platelets widely separated from peritrematal shields by a strip of soft integument; trochanter I with six setae, genu III with eight setae.................................................................
<taxonomicName id="4C5E4D61FFEAB610FB0DFBD6FA386D40" authority="Berlese, 1910" authorityName="Berlese" authorityYear="1910" box="[1202,1436,1089,1111]" class="Arachnida" family="Eviphididae" genus="Copriphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB0DFBD6FAB66D40" box="[1202,1298,1090,1111]" italics="true" pageId="4" pageNumber="279">Copriphis</emphasis>
Berlese, 1910
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FBCBFA386DB8" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FBCBFA386D85" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FBCBFA386D85" pageId="4" pageNumber="279">
12. Coxae I and II with at least one ventral seta modified into flat oval disk-like protuberance; genital setae inserted outside epigynal shield................................................................
<taxonomicName id="4C5E4D61FFEAB610FBC5FBE9FA386D85" authority="Womersley, 1956" authorityName="Womersley" authorityYear="1956" box="[1146,1436,1148,1170]" class="Arachnida" family="Eviphididae" genus="Scarabaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FBC5FBE9FB4B6D85" box="[1146,1263,1149,1170]" italics="true" pageId="4" pageNumber="279">Scarabaspis</emphasis>
Womersley, 1956
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FB0EFA386DB8" blockId="4.[151,1436,151,2021]" box="[151,1436,1177,1199]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FB0EFA386DB8" box="[151,1436,1177,1199]" pageId="4" pageNumber="279">- Coxae I and II with normal needle-like setae; genital setae inserted on epigynal shield............................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FB23FA3F6C48" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FB23FA3F6C10" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FB23FA3F6C10" pageId="4" pageNumber="279">13. Soft integument with granular incrustation, at least on surface adjacent to dorsal shield; highly domed forms with some dorsal shield setae apically rounded; dorsal shield not completely covering dorsum; anterior hyaline margin of epigynal shield subtriangular and mostly pointed, anteriorly reaching beyond posterior margin of sternal shield; presternal platelets absent.... 14</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FA9BFA3F6C48" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FA9BFA3F6C48" pageId="4" pageNumber="279">- Soft integument striated, without additional granular incrustation; dorso-ventrally flattened forms with all dorsal shield setae pointed; dorsal shield almost completely covering dorsum; anterior hyaline margin of epigynal shield straight or rounded; presternal platelets usually present........................................................................ 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28FAF3FA3F6F18" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28FAF3FA386CA0" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FAF3FA386CA0" pageId="4" pageNumber="279">
14. Vertical setae j1 elongated and pointed, lanceolate; idiosoma widely oval, with regularly rounded caudal surface; most dorsal setae thickened, stick- or rod-shaped, with blunt or rounded tip; dorsolateral and opisthogastric integument with 14 pairs of setae; epistome with one point.........................................................
<taxonomicName id="4C5E4D61FFEAB610FB02FA36FA386CA0" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1213,1436,1441,1463]" class="Arachnida" family="Eviphididae" genus="Uroiphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FB02FA36FAB66CA0" box="[1213,1298,1442,1463]" italics="true" pageId="4" pageNumber="279">Uroiphis</emphasis>
Berlese, 1903
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28FA2BFA3F6F18" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28FA2BFA3F6F18" pageId="4" pageNumber="279">- Vertical setae j1 shortened and rounded, columnar; idiosoma lemon-shaped, with a caudal projection bearing a pair of marginal setae; most dorsal setae pointed, needle-like; dorsolateral and opisthogastric integument with ten pairs of setae; epistome with three points......................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AA2E47FFEAB610FF28F983FA386E00" pageId="4" pageNumber="279">
<paragraph id="8BE136E2FFEAB610FF28F983FA386FB5" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28F983FA386FB5" pageId="4" pageNumber="279">
15. Dorsal shield with medial protuberance, suboval, sometimes slightly constricted medially, widest in anterior part, and with punctate-reticulate sculpture; anterior extension of dorsal shield expanded onto ventral surface beyond vertex, fused to anterior parts of peritrematal shields to form an ventral arch-shaped shield structure; setae j1 and z1 displaced ventrally; endopodal platelets II-III fused to sternal shield; each metasternal seta st4 and associated pore inserted together on small metasternal platelet...................................................................
<taxonomicName id="4C5E4D61FFEAB610FBEDF919FA386FB5" authority="Masan &amp; Halliday, 2009" authorityName="Masan &amp; Halliday" authorityYear="2009" box="[1106,1436,1676,1698]" class="Arachnida" family="Eviphididae" genus="Alloseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FBEDF919FB0E6FB5" box="[1106,1194,1677,1698]" italics="true" pageId="4" pageNumber="279">Alloseius</emphasis>
Mašán &amp; Halliday, 2009
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28F93EFA386E00" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28F93EFA386E00" pageId="4" pageNumber="279">
- Dorsal shield flat, rectangular, with parallel lateral margins, widest in posterior part, with coarse rugose-verrucose sculpture; anterior end of dorsal shield not expanded ventrally beyond vertex, anterior sections of peritrematal shields not fused into conspicuous ventral arch-shaped shield structure; setae j1 and z
<quantity id="4CA69B07FFEAB610FC98F970FCEF6FED" box="[807,843,1764,1786]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="279" unit="in" value="1.0">1 in</quantity>
normal dorsal positions; endopodal platelets II-III well separated from sternal shield; metasternal setae st4 and associated pores on soft integument..
<taxonomicName id="4C5E4D61FFEAB610FBFDF896FA386E00" authority="Masan &amp; Halliday, 2009" authorityName="Masan &amp; Halliday" authorityYear="2009" box="[1090,1436,1793,1815]" class="Arachnida" family="Eviphididae" genus="Halolaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FBFDF896FB0E6E00" box="[1090,1194,1794,1815]" italics="true" pageId="4" pageNumber="279">Halolaspis</emphasis>
Mašán &amp; Halliday, 2009
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BE136E2FFEAB610FF28F88BFA386E9A" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28F88BFA386E9A" pageId="4" pageNumber="279">
<keyStep id="30AA2E47FFEAB610FF28F88BFA386E9A" pageId="4" pageNumber="279">
16. Dorsal shield with punctate-reticulate sculpture; anterior extension of dorsal shield overlapping onto ventral surface beyond vertex, fused to anterior parts of peritrematal shields to form a ventral arch-shaped shield structure; setae j1 and z1 displaced ventrally; movable digit of chelicera bidentate; metasternal setae st4 on soft integument; endopodal platelets II-III almost free, fused to sternal shield only at their narrow anterior ends.......................
<taxonomicName id="4C5E4D61FFEAB610FBA3F8E3FA386E9A" authority="Masan &amp; Halliday, 2009" authorityName="Masan &amp; Halliday" authorityYear="2009" box="[1052,1436,1911,1933]" class="Arachnida" family="Eviphididae" genus="Pseudoalliphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEAB610FBA3F8E3FB0E6E9B" box="[1052,1194,1911,1932]" italics="true" pageId="4" pageNumber="279">Pseudoalliphis</emphasis>
Mašán &amp; Halliday, 2009
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEAB610FF28F801FA386EF2" blockId="4.[151,1436,151,2021]" pageId="4" pageNumber="279">
<keyLead id="30AF95D7FFEAB610FF28F801FA386EF2" pageId="4" pageNumber="279">
- Dorsal shield with smooth or finely reticulated surface; anterior end of dorsal shield not expanded ventrally beyond vertex, anterior sections of peritrematal shields not fused into conspicuous ventral arch-shaped shield structure; setae j1 and z
<quantity id="4CA69B07FFEAB610FAF2F826FACB6EDF" box="[1357,1391,1970,1992]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="279" unit="in" value="1.0">1 in</quantity>
normal dorsal position; movable digit of chelicera with one robust subdistal tooth; metasternal setae st4 on small metasternal
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEBB611FF5EFF03FA3F69F0" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="280">
platelets; endopodal platelets II-III and sternal shield completely fused.......................................... 17
<keyStep id="30AA2E47FFEBB611FF28FF20FA3F69F0" pageId="5" pageNumber="280">
<keyLead id="30AF95D7FFEBB611FF28FF20FA3F69F0" pageId="5" pageNumber="280">
17. Dorsal shield setae with extreme variation in length, longest setae about ten times as long as shortest setae, S5 usually several times as long as Z5................................................................
<taxonomicName id="4C5E4D61FFEBB611FB18FF46FA3F69F0" authority="Berlese, 1911" authorityName="Berlese" authorityYear="1911" box="[1191,1435,209,231]" class="Arachnida" family="Eviphididae" genus="Pelethiphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEBB611FB18FF46FAB769F0" box="[1191,1299,210,231]" italics="true" pageId="5" pageNumber="280">Pelethiphis</emphasis>
Berlese, 1911
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BE136E2FFEBB611FF28FF7BFA3F6835" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="280">
<keyLead id="30AF95D7FFEBB611FF28FF7BFA3F6835" pageId="5" pageNumber="280">- Dorsal shield setae without such extreme variation in length, longest setae at most three times as long as shortest setae, Z5 and S5 subequal in length................................................................................. 18</keyLead>
</paragraph>
<keyStep id="30AA2E47FFEBB611FF28FEBDFA3868C5" pageId="5" pageNumber="280">
<paragraph id="8BE136E2FFEBB611FF28FEBDFA3F686D" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="280">
<keyLead id="30AF95D7FFEBB611FF28FEBDFA3F686D" pageId="5" pageNumber="280">
18. Peritrematal shield developed along whole length of peritreme; anterior section of peritrematal shield wide and completely fused to dorsal shield; setae j1 and z1 inserted dorsally; posterior dorsal shield setae subequal or only moderately varying in length; exopodal platelets I-II and II-III present.............................................
<taxonomicName id="4C5E4D61FFEBB611FB78FEF1FA3F686D" authority="Halbert, 1923" authorityName="Halbert" authorityYear="1923" box="[1223,1435,356,378]" class="Arachnida" family="Eviphididae" genus="Alliphis" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEBB611FB78FEF1FAB6686D" box="[1223,1298,357,378]" italics="true" pageId="5" pageNumber="280">Alliphis</emphasis>
Halbert, 1923
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE136E2FFEBB611FF28FE16FA3868C5" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="280">
<keyLead id="30AF95D7FFEBB611FF28FE16FA3868C5" pageId="5" pageNumber="280">
- Peritrematal shield not reaching the anterior tip of peritremes; peritrematal shields or peritremes not connected to dorsal shield; setae j1 and z1 usually displaced ventrally; posterior marginal setae on dorsal shield (S- and Z-setae) conspicuously longer than medial J-setae; exopodal platelets I-II and II-III absent.............................
<taxonomicName id="4C5E4D61FFEBB611FBC4FE29FA3868C5" authority="Masan, 1994" authorityName="Masan" authorityYear="1994" box="[1147,1436,444,466]" class="Arachnida" family="Eviphididae" genus="Scarabacariphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAF0FFEBB611FBC4FE29FABD68C5" box="[1147,1305,445,466]" italics="true" pageId="5" pageNumber="280">Scarabacariphis</emphasis>
<bibRefCitation id="EFCF4B13FFEBB611FA9FFE28FA3868C5" author="Masan" box="[1312,1436,444,466]" pageId="5" pageNumber="280" refString="Masan, P. (1994) The eviphidid mites (Acarina: Mesostigmata; Eviphididae) associated with scarabaeid and carrion beetles (Coleoptera: Scarabaeidae, Silphidae) in central Europe. Acarologia, 35, 3 - 19." type="journal article" year="1994">Mašán, 1994</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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