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<document ID-DOI="http://doi.org/10.5281/zenodo.3789914" ID-GBIF-Dataset="d33a3344-7f9a-4028-8569-2b7643d9b43f" ID-GBIF-Taxon="163794641" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789914" ID-ZooBank="9CACC88D-12A3-4FEC-948E-90365B649BB6" approvalRequired="163" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="158" approvalRequired_for_treatments="2" checkinTime="1588012210414" checkinUser="plazi" docAuthor="van Nieukerken, Erik, Laštůvka, Aleš &amp; Laštůvka, Zdeněk" docDate="2010" docId="03F787B3D459FFD1FF681396FB806D84" docLanguage="en" docName="ZK_article_2128.pdf" docOrigin="ZooKeys 32 (32)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@3c5ded30" docStyleName="zookeys.2008.journal_article" docTitle="Ectoedemia (Ectoedemia) intimella" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFCEFFCBD44CFFC4FFE41275FFCD693A" masterDocTitle="Western Palaearctic Ectoedemia (Zimmermannia) Hering and Ectoedemia Busck s. str. (Lepidoptera: Nepticulidae): five new species and new data on distribution, hostplants and recognition" masterLastPageNumber="82" masterPageNumber="1" pageId="21" pageNumber="22" updateTime="1643558219303" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Western Palaearctic Ectoedemia (Zimmermannia) Hering and Ectoedemia Busck s. str. (Lepidoptera: Nepticulidae): five new species and new data on distribution, hostplants and recognition</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Nieukerken, Erik</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Laštůvka, Aleš</mods:namePart>
<mods:affiliation>Mendel University of Agriculture &amp; Forestry, Brno ,, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Laštůvka, Zdeněk</mods:namePart>
<mods:affiliation>Mendel University of Agriculture &amp; Forestry, Brno ,, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-01-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>32</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>32</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>82</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.32.282</mods:identifier>
<mods:identifier type="GBIF-Dataset">d33a3344-7f9a-4028-8569-2b7643d9b43f</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576594</mods:identifier>
<mods:identifier type="ZooBank">9CACC88D-12A3-4FEC-948E-90365B649BB6</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789914" ID-GBIF-Taxon="163794641" ID-Zenodo-Dep="3789914" LSID="urn:lsid:plazi:treatment:03F787B3D459FFD1FF681396FB806D84" httpUri="http://treatment.plazi.org/id/03F787B3D459FFD1FF681396FB806D84" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection box="[140,649,483,510]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph blockId="21.[140,649,483,510]" box="[140,649,483,510]" pageId="21" pageNumber="22">
<heading box="[140,649,483,510]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="3">
<taxonomicName authority="(Zeller)" baseAuthorityName="Zeller" baseAuthorityYear="1848" box="[140,649,483,510]" class="Insecta" family="Nepticulidae" genus="Ectoedemia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="intimella" subGenus="Ectoedemia">
<emphasis bold="true" box="[140,557,483,510]" italics="true" pageId="21" pageNumber="22">Ectoedemia (Ectoedemia) intimella</emphasis>
(Zeller)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="biology_ecology">
<paragraph blockId="21.[140,1108,553,1214]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[140,237,553,579]" pageId="21" pageNumber="22">
<emphasis box="[140,233,553,579]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Biology</emphasis>
.
</emphasis>
Hostplants:
<taxonomicName box="[380,508,554,580]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="caprea">
<emphasis box="[380,508,554,580]" italics="true" pageId="21" pageNumber="22">Salix caprea</emphasis>
</taxonomicName>
L.,
<taxonomicName box="[551,654,554,580]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cinerea">
<emphasis box="[551,654,554,580]" italics="true" pageId="21" pageNumber="22">S. cinerea</emphasis>
</taxonomicName>
L.,
<taxonomicName box="[697,834,554,580]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="pentandra">
<emphasis box="[697,834,554,580]" italics="true" pageId="21" pageNumber="22">S. pentandra</emphasis>
</taxonomicName>
L.,
<taxonomicName box="[876,975,554,580]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="fragilis">
<emphasis box="[876,975,554,580]" italics="true" pageId="21" pageNumber="22">S. fragilis</emphasis>
</taxonomicName>
L.,
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="phylicifolia">
<emphasis italics="true" pageId="21" pageNumber="22">S. phylicifolia</emphasis>
</taxonomicName>
L.,
<taxonomicName authority="Wimm. (Ellis 2007)" authorityName="Wimm. (Ellis" authorityYear="2007" box="[249,621,589,615]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dasyclados">
<emphasis box="[249,384,589,615]" italics="true" pageId="21" pageNumber="22">S. dasyclados</emphasis>
Wimm. (
<bibRefCitation author="Ellis WN" box="[493,613,589,615]" pageId="21" pageNumber="22" refId="ref39166" refString="Ellis WN (2007) Nederlandse bladmineerders / Dutch leafminers. http: // www. bladmineerders. nl / index. htm [accessed 23 VI 2009]." type="url" year="2007">Ellis 2007</bibRefCitation>
)
</taxonomicName>
. In
<collectingCountry box="[662,749,589,615]" name="Finland" pageId="21" pageNumber="22">Finland</collectingCountry>
a population of
<taxonomicName baseAuthorityName="Zeller" baseAuthorityYear="1848" box="[932,1061,589,615]" class="Insecta" family="Nepticulidae" genus="Ectoedemia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="intimella">
<emphasis box="[932,1061,589,615]" italics="true" pageId="21" pageNumber="22">E. intimella</emphasis>
</taxonomicName>
was discovered feeding on
<taxonomicName authority="L. Th" authorityName="L. Th" box="[393,670,624,651]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="balsamifera">
<emphasis box="[393,607,624,650]" italics="true" pageId="21" pageNumber="22">Populus balsamifera</emphasis>
L. Th
</taxonomicName>
ese mines resemble those of
<taxonomicName baseAuthorityName="Glitz" baseAuthorityYear="1872" class="Insecta" family="Nepticulidae" genus="Ectoedemia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hannoverella">
<emphasis italics="true" pageId="21" pageNumber="22">E. hannoverella</emphasis>
</taxonomicName>
, but the mine usually enters the leaf blade further away from the petiole and the larva doesn’t show the characteristic plates of
<taxonomicName authority="(Gustafsson and Nieukerken 1990)" baseAuthorityName="Gustafsson and Nieukerken" baseAuthorityYear="1990" class="Insecta" family="Nepticulidae" genus="Ectoedemia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hannoverella">
<emphasis box="[650,819,695,721]" italics="true" pageId="21" pageNumber="22">E. hannoverella</emphasis>
(
<bibRefCitation author="Gustafsson B &amp; Nieukerken EJ van" pageId="21" pageNumber="22" pagination="323 - 356" refId="ref39235" refString="Gustafsson B, Nieukerken EJ van (1990) Larvae of Nepticulidae. In: Johansson R, Nielsen ES, Nieukerken EJ van, Gustafsson B (Eds) Th e Nepticulidae and Opostegidae (Lepidoptera) of NW Europe. E. J. Brill, Leiden, 323 - 356." type="book chapter" year="1990">Gustafsson and Nieukerken 1990</bibRefCitation>
)
</taxonomicName>
. Univoltine, larvae in September to November, adults from May to July.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="distribution">
<paragraph blockId="21.[140,1108,553,1214]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[185,344,765,791]" pageId="21" pageNumber="22">
<emphasis box="[185,339,765,791]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Distribution</emphasis>
.
</emphasis>
<collectingCountry box="[357,452,766,792]" name="Norway" pageId="21" pageNumber="22">Norway</collectingCountry>
:
<bibRefCitation author="Aarvik L &amp; Bakke SA &amp; Berg Y &amp; Berggren K &amp; Hansen LO &amp; Myhr K &amp; Svendsen S" box="[469,693,765,792]" pageId="21" pageNumber="22" pagination="55 - 70" refId="ref38028" refString="Aarvik L, Bakke SA, Berg Y, Berggren K, Hansen LO, Myhr K, Svendsen S (1997) Contribution to the knowledge of the Norwegian Lepidoptera 5. Fauna Norvegica Series B 44: 55 - 70." type="journal article" year="1997">Aarvik et al. 1997</bibRefCitation>
;
<collectingCountry box="[712,796,765,791]" name="Ireland" pageId="21" pageNumber="22">Ireland</collectingCountry>
:
<bibRefCitation author="Shackleton J" box="[813,1012,765,791]" pageId="21" pageNumber="22" pagination="98" refId="ref41856" refString="Shackleton J (1977) Ectoedemia intimella (Zeller) and E. subbimaculella (Haworth): two species of Lepidoptera (Nepticulidae) new to Ireland. Irish Naturalist's Journal 19: 98." type="journal article" year="1977">Shackleton 1977</bibRefCitation>
;
<collectingCountry box="[1031,1104,765,791]" name="Latvia" pageId="21" pageNumber="22">Latvia</collectingCountry>
:
<bibRefCitation author="Sulcs A &amp; Sulcs I" box="[140,379,798,828]" pageId="21" pageNumber="22" pagination="67 - 72" refId="ref42034" refString="Sulcs A, Sulcs I (1989) Neue und wenig bekannte Arten der Lepidopteren-Fauna Lettlands. 12. Mitteilung. Notulae Entomologicae 69: 67 - 72." type="journal article" year="1989">Šulcs and Šulcs 1989</bibRefCitation>
;
<collectingCountry box="[391,505,800,826]" name="Lithuania" pageId="21" pageNumber="22">Lithuania</collectingCountry>
:
<bibRefCitation author="Ivinskis P &amp; Pakalniskis S &amp; Puplesis R" box="[515,731,800,827]" pageId="21" pageNumber="22" pagination="1 - 240" refId="ref39437" refString="Ivinskis P, Pakalniskis S, Puplesis R (1985) Augalus Minuojantys Vabzdziai [Leafmining insects]. Mokslas, Vilnius, 1 - 240 pp." type="book chapter" year="1985">Ivinskis et al. 1985</bibRefCitation>
;
<collectingCountry box="[743,840,800,826]" name="Slovakia" pageId="21" pageNumber="22">Slovakia</collectingCountry>
:
<bibRefCitation author="Lastuvka A &amp; Lastuvka Z" pageId="21" pageNumber="22" pagination="185 - 192" refId="ref39848" refString="Lastuvka A, Lastuvka Z (1990 b) Zur Faunistik der Nepticulidae-Arten in der Tschechoslowakei (Lepidoptera). Casopis Moravskeho Muzea Vedy Prirodni 75: 185 - 192." type="journal article" year="1990">Laštůvka and Laštůvka 1990b</bibRefCitation>
;
<collectingCountry box="[228,295,835,861]" name="Spain" pageId="21" pageNumber="22">Spain</collectingCountry>
:
<bibRefCitation author="Lastuvka A &amp; Lastuvka Z" box="[306,632,835,862]" pageId="21" pageNumber="22" pagination="457 - 464" refId="ref39974" refString="Lastuvka A, Lastuvka Z (2008) Seven Nepticulidae new to the Iberian Peninsula and several new province records (Lepidoptera: Nepticulidae). SHILAP Revista de Lepidopterologia 36: 457 - 464." type="journal article" year="2008">Laštůvka and Laštůvka 2008</bibRefCitation>
. For new French records: Van
<bibRefCitation author="Nieukerken EJ van &amp; Lastuvka A &amp; Lastuvka Z" pageId="21" pageNumber="22" pagination="1 - 114" refId="ref40987" refString="Nieukerken EJ van, Lastuvka A, Lastuvka Z (2006) Annotated catalogue of the Nepticulidae and Opostegidae (Lepidoptera: Nepticuloidea) of France and Corsica. Zootaxa 1216: 1 - 114. http: // www. mapress. com / zootaxa / 2006 / zt 01216 p 114. pdf." type="journal article" year="2006">Nieukerken et al. 2006</bibRefCitation>
, here we record it new from southern
<collectingCountry box="[685,764,871,897]" name="France" pageId="21" pageNumber="22">France</collectingCountry>
. Also found in far eastern
<collectingCountry name="Russia" pageId="21" pageNumber="22">Russia</collectingCountry>
:
<collectingRegion box="[184,281,906,932]" country="Russia" name="Sakhalin" pageId="21" pageNumber="22">Sakhalin</collectingRegion>
(
<bibRefCitation author="Puplesis R" box="[296,460,906,932]" pageId="21" pageNumber="22" refId="ref41419" refString="Puplesis R (1994) Th e Nepticulidae of eastern Europe and Asia. Western, central and eastern parts. Backhuys Publishers, Leiden, 290 pp." type="book" year="1994">Puplesis 1994</bibRefCitation>
). Here for the first time recorded from
<collectingCountry box="[907,973,906,932]" name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
,
<collectingRegion box="[986,1103,906,932]" country="Japan" name="Hokkaido" pageId="21" pageNumber="22">Hokkaido</collectingRegion>
, not far from Sakhalin.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="materials_examined">
<paragraph blockId="21.[140,1108,553,1214]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[185,387,976,1002]" pageId="21" pageNumber="22">
<emphasis box="[185,289,976,1002]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Material</emphasis>
.
<collectingCountry box="[300,382,976,1002]" name="France" pageId="21" pageNumber="22">France</collectingCountry>
:
</emphasis>
<specimenCount box="[392,470,976,1002]" pageId="21" pageNumber="22" type="larva">1 larva</specimenCount>
, Alpes Maritimes, Les Mesches,
<quantity box="[828,885,976,1002]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="km" value="6.0">6 km</quantity>
WSW Tende,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="22" unit="m" value="1450.0">1450 m</quantity>
,
<date box="[177,301,1012,1038]" pageId="21" pageNumber="22" value="2008-10-10">10.X.2008</date>
, leafmine on
<taxonomicName box="[453,584,1011,1037]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="caprea">
<emphasis box="[453,584,1011,1037]" italics="true" pageId="21" pageNumber="22">Salix caprea</emphasis>
</taxonomicName>
, EJvN2008224,
<collectionCode box="[778,873,1012,1038]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
.
<accessionNumber box="[876,1007,1011,1038]" httpUri="http://www.ncbi.nlm.nih.gov/protein/INS17603" pageId="21" pageNumber="22">
<collectionCode box="[876,925,1012,1038]" pageId="21" pageNumber="22">INS</collectionCode>
17603
</accessionNumber>
, E.J. van Nieukerken &amp; C. Doorenweerd (
<collectionCode box="[512,610,1047,1073]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
).
<emphasis bold="true" box="[628,728,1046,1072]" pageId="21" pageNumber="22">
<collectingCountry box="[628,724,1046,1072]" name="Finland" pageId="21" pageNumber="22">Finland</collectingCountry>
:
</emphasis>
<specimenCount box="[734,824,1047,1073]" pageId="21" pageNumber="22" type="larva">4 larvae</specimenCount>
, leafmines, OK: Joensuu,
<date box="[140,264,1082,1109]" pageId="21" pageNumber="22" value="2005-10-18">18.X.2005</date>
,
<emphasis box="[277,378,1082,1108]" italics="true" pageId="21" pageNumber="22">
<taxonomicName box="[277,361,1082,1108]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
?
</emphasis>
<taxonomicName authority=", A. Karhu &amp; A. Matero" authorityName="A. Karhu &amp; A. Matero" box="[378,779,1082,1108]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="balsamifera">
<emphasis box="[378,502,1082,1108]" italics="true" pageId="21" pageNumber="22">balsamifera</emphasis>
, A. Karhu &amp; A. Matero
</taxonomicName>
(
<collectionCode box="[795,893,1082,1108]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
).
<emphasis bold="true" box="[913,975,1082,1108]" pageId="21" pageNumber="22">
<collectingCountry box="[913,970,1082,1108]" name="Italy" pageId="21" pageNumber="22">Italy</collectingCountry>
:
</emphasis>
larvae, Cuneo, Palanfré, W.,
<quantity box="[353,520,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.49" metricValueMax="1.54" metricValueMin="1.44" pageId="21" pageNumber="22" unit="m" value="1490.0" valueMax="1540.0" valueMin="1440.0">1440–1540 m</quantity>
,
<date box="[533,657,1117,1143]" pageId="21" pageNumber="22" value="2008-10-14">14.X.2008</date>
, mines on
<taxonomicName box="[786,918,1117,1143]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="caprea">
<emphasis box="[786,918,1117,1143]" italics="true" pageId="21" pageNumber="22">Salix caprea</emphasis>
</taxonomicName>
, EJvN2008300,
<collectionCode box="[140,235,1152,1178]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
.
<accessionNumber box="[238,363,1152,1178]" httpUri="http://www.ncbi.nlm.nih.gov/protein/INS12990" pageId="21" pageNumber="22">
<collectionCode box="[238,287,1152,1178]" pageId="21" pageNumber="22">INS</collectionCode>
12990
</accessionNumber>
, 17642–3, E.J. van Nieukerken &amp; C. Doorenweerd (
<collectionCode box="[958,1056,1152,1178]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
).
<emphasis bold="true" pageId="21" pageNumber="22">
<collectingCountry name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
:
</emphasis>
<specimenCount box="[199,229,1188,1214]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
,
<collectingRegion box="[242,359,1187,1213]" country="Japan" name="Hokkaido" pageId="21" pageNumber="22">Hokkaido</collectingRegion>
, Kitami-shi, Toyochi-oohashi,
<date box="[705,825,1188,1214]" pageId="21" pageNumber="22" value="2004-06-08">
8.
<collectionCode box="[721,755,1188,1214]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="21" pageNumber="22">VI</collectionCode>
.2004
</date>
, S. Kawahara (
<collectionCode box="[990,1088,1188,1214]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22" type="Museum">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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